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STEMS
•   Origin
•   Functions
•   External Anatomy
•   Internal Anatomy
•   Specialized
    Stems
Origin
• First stem of a plant develops from part
  of a seed embryo called epicotyl, w/c is a
  continuation of the hypocotyl
The Plant Body: Stems
FUNCTIONS OF STEMS
• Produces & support appendages
  of plant (leaves, flowers, fruits)
• transport water and solutes
  between roots and leaves.
• Stems in some plants are
  photosynthetic.
• Produce & store materials
  necessary for life (e.g., water,
  starch, sugar).
• In some plants, stems have
  become adapted for specialized
  functions.
Stems support a display of
leaves.

Stems orient the leaves
toward the light with minimal
overlap among the leaves.
The stem supports a display of flowers
Opuntia-prickly pear
The stem does photosynthesis…

                                and stores water.
This stem does
 photosynthesis, stores
 water, but also produces a
 defense chemical:
 mescaline…a hallucinogen.




Lophophora williamsii - peyote
Two Types of Aerial Stems
       Herbaceous Stems                 Woody Stems
•   Soft & green                • Tough & not green
•   Little growth in diameter   • Considerable growth in
•   Tissues chiefly primary       diameter
•   Chiefly annual              • Tissues chiefly secondary
•   Covered by epidermis        • Chiefly perennial
•   Buds mostly naked           • Covered by corky bark
                                • Buds chiefly covered by
                                  scales
Herbaceous Stem   woody stem
EXTERNAL ANATOMY
STEM APICAL MERISTEM
PRIMARY & SECONDARY GROWTH
Apical Dominance
       Apical dominance refers
        to the suppression of
        growth by hormones
        produced in the apical
        meristem.

        Lateral branch growth are
         inhibited near the shoot
         apex, but less so farther
         from the tip.
        Apical dominance is
         disrupted in some plants by
         removing the shoot tip,
         causing the plant to become
         bushy.
INTERNAL STEM
   ANATOMY
Monocotyledonous &
Dicotyledonous Flowering Plants
Monocot Stem – cross section
Typical Stem Cross Section (Dicot Stem)

  Epidermis                                Helianthus annuus-
                                           sun flower annual

Cortex

              A ring of vascular bundles




                          Pith
Epidermis
   - window, reduce water loss
Cortex Collenchyma
             - extensible support
Cortex Parenchyma
           - photosynthesis, etc.
Fibers- rigid support

Functional Phloem
      - conduct sugars etc. away
         from leaf to rest of plant
Vascular Cambium
 - adds 2° xylem and 2° phloem
Xylem
    -conduct water and minerals
                      up from soil
Pith
         -water storage, defense?
VIP Stem: Provide both name and function labels:
                   Epidermis: reduce evaporation, gas exchange
                     Cortex: photosynthesis, collenchyma support
                       Vascular Bundles: conduction
                       Pith: water storage? defense? disintegrate?




                                  Vascular Bundle:
to center            outside
                                    Phloem Fibers: support
                                    Functional Phloem:
                                         conduct CH2O away from leaf
                                     Vascular Cambium:
                                          add 2° Xylem and 2° Phloem
                                     Xylem:
                                          conduct minerals up from soil
outside             to center
Vitis vinifera - grape
Notice how the vascular cambia
 of adjacent vascular bundles
 line up side by side.


 Notice that cambium tissue
 differentiates between the
 bundles, connecting the cambia
 together.

 Remnants of the procambium:
      Intrafasicular cambium
      Interfasicular cambium


Vitis vinifera - grape
The vascular cambium makes 2° tissues:
                                         Vitis vinifera - grape




                                    2° xyl
                                      2°
                                      ph em
                                         loe
                                             m
Basswood – 1 & 2 years old
Three years of Secondary Growth
                                              Tilia - basswood


                                   Secondary
                 c ambium
                                         Phloem

                                  Secondary
                                  Xylem
A cork cambium differentiates and produces a periderm.
                                    Epidermis
    cutin



                          suberin

                                     Cork Cells

                                     Cork Cambium
                                     Phelloderm
Over time, the epidermis dies.
The cork cells build up to for
a thick layer for the bark of a
tree. We use this to make
stoppers for wine bottles and
so on.



When suberin is fully
developed, the cortex cells
will eventually be in the
dark. So these chloroplasts
will lose their function!
Bark =
   epidermis +
    periderm +
      cortex +
     phloem +
      vascular
     cambium

Wood =
     secondary
   xylem only!

Pith =
        a small
 percentage of
 tree diameter
    at maturity
Anatomy of a Woody Stem
The trees pictured below have long lost their
epidermis on the woody portion of the stem




         Sequoia sempervirens - giant sequoia
The study of the growth rings in wood: Dendrochronology
Each year the cambium
produces a layer of secondary
xylem and a layer of secondary
phloem.

This photo shows secondary
xylem from parts of three years
in Pinus strobus (white pine).




   spring of the next year
  winter of that year
   fall of that year


   mid-summer of one year
Stems
Stems—the axes of plants—consist of nodes (where
leaves and axillary buds are produced) separated by
internodes.
Node - region of the stem where the leaf and bud are
borne.

Internode - the part of the stem between two adjacent
nodes.

Herbaceous - not woody; dying down at the end of the
growing season.

Woody - hard in texture, containing secondary xylem, and
persisting more than one growing season.

Acaulescent - having an inconspicuous stem.

Caulescent - having a distinct stem.
Modified & Specialized Stems
Bulbs
• Bulbs - large buds
  with a small stem at
  the lower end
  surrounded by
  numerous fleshy
  leaves that store
  nutrients; adventitious
  roots at base
• Eg. onion, tulip, hyacinth,
  daffodil and lily
Corms
• Corms - resemble
  bulbs but composed
  entirely of stem tissue
  surrounded by a few
  papery scale like
  leaves, food storage
  organs with
  adventitious roots at
  the base of corms
• Eg. crocus and
  gladiolus.
Rhizomes
• Rhizomes - horizontal
  stems that grow
  below the ground with
  adventitious roots

• Eg. irises, ferns, and
  grasses.
Cladophylls
•Cladophylls or
cladodes - leaf-like
stems modified for
photosynthesis

•Eg. butcher's broom,
asparagus, orchids (eg.
Epidendrum)
Succulent Stems
• Succulent stems -
  stout fleshy stems
  that are modified for
  water and food
  storage
 Eg. cacti
Thorns- for protection from grazing animals
• Bougainvillea
• Honey locust (modified
  stem)
• Black Locust (modified
  leaf stipules)
Food Storage Stems
Prickly Pear
Cactus




Bamboo Shoots     Kohlrabi
Food Storage Stems - Sugarcane
Tubers
• Tubers – swollen
  regions of stems that
  store food for
  subsequent growth
• The "eyes" of a potato
  (irish potatoes Solanum
  tuberosum) are the nodes
  of a starch-ladened stem
Stolons
• Stolons or runners -            Bermuda grass (Cynodon dactylon).
  horizontal stem that
  grow above the
  ground with long
  internodes
Eg. Bermuda grass
      (Cynodon dactylon)
    Spider plant (Chlorophytum)
    Fern (Nephrolepis)
Rosette
• Rosette - stem with short internodes and leaves
  attached at nodes
Wild Radish – Rosette & Bolt
       A FLOWERING ANNUAL




     YEAR ONE          YEAR ONE
Common Mullen – Rosette & Bolt
  A FLOWERING BIENNIAL




       YEAR ONE


                         YEAR TWO
Buds
Buds are short embryonic stems. In
angiosperms they are found at the nodes,
in the leaf axil (the angle formed by the
stem and the petiole of the leaf).

Axillary bud - a bud located in the leaf
axil.
Terminal bud - a bud at the apex of a
stem.
PHYSIOLOGY OF STEMS
• Conduction of Materials by Xylem
  1) Root pressure – powered by transpiration of water from
  the leaves
  2) Transpiration pull and water cohesion –
  water is pulled up from the roots due to adhesion of water to the
  xylem walls & tension generated by the water-potential gradient bet.
  leaves & xylem
  Other contributing factors:
  3) Atmospheric pressure
  4) Action of Living cells
  5) Imbibition in cell walls of xylem
  6) Capillary attraction
PHYSIOLOGY OF STEMS
• Conduction of Materials by Phloem
  - nutrient-rich fluid in the phloem moves from
   areas of high solute concentration & water
   pressure to areas of low solute concentration &
   water pressure

• Hypotheses of phloem function are:
  1) cytoplasmic streaming
  2) movement through interface
  3) pressure flow or mass flow
Transpiration-Cohesion Hypothesis
       for Water Movement
Sugar Loading of Phloem and Bulk Flow
Sugar Loading of Phloem and Bulk Flow

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Stems 100926175806-phpapp02

  • 1. STEMS • Origin • Functions • External Anatomy • Internal Anatomy • Specialized Stems
  • 2. Origin • First stem of a plant develops from part of a seed embryo called epicotyl, w/c is a continuation of the hypocotyl
  • 3. The Plant Body: Stems FUNCTIONS OF STEMS • Produces & support appendages of plant (leaves, flowers, fruits) • transport water and solutes between roots and leaves. • Stems in some plants are photosynthetic. • Produce & store materials necessary for life (e.g., water, starch, sugar). • In some plants, stems have become adapted for specialized functions.
  • 4. Stems support a display of leaves. Stems orient the leaves toward the light with minimal overlap among the leaves.
  • 5. The stem supports a display of flowers
  • 6. Opuntia-prickly pear The stem does photosynthesis… and stores water.
  • 7. This stem does photosynthesis, stores water, but also produces a defense chemical: mescaline…a hallucinogen. Lophophora williamsii - peyote
  • 8. Two Types of Aerial Stems Herbaceous Stems Woody Stems • Soft & green • Tough & not green • Little growth in diameter • Considerable growth in • Tissues chiefly primary diameter • Chiefly annual • Tissues chiefly secondary • Covered by epidermis • Chiefly perennial • Buds mostly naked • Covered by corky bark • Buds chiefly covered by scales
  • 9. Herbaceous Stem woody stem
  • 11.
  • 13.
  • 14.
  • 15.
  • 17. Apical Dominance  Apical dominance refers to the suppression of growth by hormones produced in the apical meristem. Lateral branch growth are inhibited near the shoot apex, but less so farther from the tip. Apical dominance is disrupted in some plants by removing the shoot tip, causing the plant to become bushy.
  • 18. INTERNAL STEM ANATOMY
  • 20. Monocot Stem – cross section
  • 21. Typical Stem Cross Section (Dicot Stem) Epidermis Helianthus annuus- sun flower annual Cortex A ring of vascular bundles Pith
  • 22. Epidermis - window, reduce water loss Cortex Collenchyma - extensible support Cortex Parenchyma - photosynthesis, etc. Fibers- rigid support Functional Phloem - conduct sugars etc. away from leaf to rest of plant Vascular Cambium - adds 2° xylem and 2° phloem Xylem -conduct water and minerals up from soil Pith -water storage, defense?
  • 23. VIP Stem: Provide both name and function labels: Epidermis: reduce evaporation, gas exchange Cortex: photosynthesis, collenchyma support Vascular Bundles: conduction Pith: water storage? defense? disintegrate? Vascular Bundle: to center outside Phloem Fibers: support Functional Phloem: conduct CH2O away from leaf Vascular Cambium: add 2° Xylem and 2° Phloem Xylem: conduct minerals up from soil outside to center
  • 25. Notice how the vascular cambia of adjacent vascular bundles line up side by side. Notice that cambium tissue differentiates between the bundles, connecting the cambia together. Remnants of the procambium: Intrafasicular cambium Interfasicular cambium Vitis vinifera - grape
  • 26. The vascular cambium makes 2° tissues: Vitis vinifera - grape 2° xyl 2° ph em loe m
  • 27. Basswood – 1 & 2 years old
  • 28. Three years of Secondary Growth Tilia - basswood Secondary c ambium Phloem Secondary Xylem
  • 29. A cork cambium differentiates and produces a periderm. Epidermis cutin suberin Cork Cells Cork Cambium Phelloderm
  • 30. Over time, the epidermis dies. The cork cells build up to for a thick layer for the bark of a tree. We use this to make stoppers for wine bottles and so on. When suberin is fully developed, the cortex cells will eventually be in the dark. So these chloroplasts will lose their function!
  • 31. Bark = epidermis + periderm + cortex + phloem + vascular cambium Wood = secondary xylem only! Pith = a small percentage of tree diameter at maturity
  • 32. Anatomy of a Woody Stem
  • 33. The trees pictured below have long lost their epidermis on the woody portion of the stem Sequoia sempervirens - giant sequoia
  • 34. The study of the growth rings in wood: Dendrochronology
  • 35. Each year the cambium produces a layer of secondary xylem and a layer of secondary phloem. This photo shows secondary xylem from parts of three years in Pinus strobus (white pine). spring of the next year winter of that year fall of that year mid-summer of one year
  • 36. Stems Stems—the axes of plants—consist of nodes (where leaves and axillary buds are produced) separated by internodes.
  • 37. Node - region of the stem where the leaf and bud are borne. Internode - the part of the stem between two adjacent nodes. Herbaceous - not woody; dying down at the end of the growing season. Woody - hard in texture, containing secondary xylem, and persisting more than one growing season. Acaulescent - having an inconspicuous stem. Caulescent - having a distinct stem.
  • 39. Bulbs • Bulbs - large buds with a small stem at the lower end surrounded by numerous fleshy leaves that store nutrients; adventitious roots at base • Eg. onion, tulip, hyacinth, daffodil and lily
  • 40. Corms • Corms - resemble bulbs but composed entirely of stem tissue surrounded by a few papery scale like leaves, food storage organs with adventitious roots at the base of corms • Eg. crocus and gladiolus.
  • 41. Rhizomes • Rhizomes - horizontal stems that grow below the ground with adventitious roots • Eg. irises, ferns, and grasses.
  • 42. Cladophylls •Cladophylls or cladodes - leaf-like stems modified for photosynthesis •Eg. butcher's broom, asparagus, orchids (eg. Epidendrum)
  • 43. Succulent Stems • Succulent stems - stout fleshy stems that are modified for water and food storage Eg. cacti
  • 44.
  • 45. Thorns- for protection from grazing animals • Bougainvillea • Honey locust (modified stem) • Black Locust (modified leaf stipules)
  • 46. Food Storage Stems Prickly Pear Cactus Bamboo Shoots Kohlrabi
  • 47. Food Storage Stems - Sugarcane
  • 48. Tubers • Tubers – swollen regions of stems that store food for subsequent growth • The "eyes" of a potato (irish potatoes Solanum tuberosum) are the nodes of a starch-ladened stem
  • 49. Stolons • Stolons or runners - Bermuda grass (Cynodon dactylon). horizontal stem that grow above the ground with long internodes Eg. Bermuda grass (Cynodon dactylon) Spider plant (Chlorophytum) Fern (Nephrolepis)
  • 50. Rosette • Rosette - stem with short internodes and leaves attached at nodes
  • 51. Wild Radish – Rosette & Bolt A FLOWERING ANNUAL YEAR ONE YEAR ONE
  • 52. Common Mullen – Rosette & Bolt A FLOWERING BIENNIAL YEAR ONE YEAR TWO
  • 53. Buds Buds are short embryonic stems. In angiosperms they are found at the nodes, in the leaf axil (the angle formed by the stem and the petiole of the leaf). Axillary bud - a bud located in the leaf axil. Terminal bud - a bud at the apex of a stem.
  • 54. PHYSIOLOGY OF STEMS • Conduction of Materials by Xylem 1) Root pressure – powered by transpiration of water from the leaves 2) Transpiration pull and water cohesion – water is pulled up from the roots due to adhesion of water to the xylem walls & tension generated by the water-potential gradient bet. leaves & xylem Other contributing factors: 3) Atmospheric pressure 4) Action of Living cells 5) Imbibition in cell walls of xylem 6) Capillary attraction
  • 55. PHYSIOLOGY OF STEMS • Conduction of Materials by Phloem - nutrient-rich fluid in the phloem moves from areas of high solute concentration & water pressure to areas of low solute concentration & water pressure • Hypotheses of phloem function are: 1) cytoplasmic streaming 2) movement through interface 3) pressure flow or mass flow
  • 56. Transpiration-Cohesion Hypothesis for Water Movement
  • 57. Sugar Loading of Phloem and Bulk Flow
  • 58. Sugar Loading of Phloem and Bulk Flow