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Lecture 3: Plant anatomy and
physiology
by
Edgar Moctezuma, Ph.D.
Today…
• Plant Anatomy
– Cells
– Tissues
– Organs
• Plant Physiology
– Water & sugar transport
– Plant hormones
• Plant Classification
From smallest to largest plants
What is plant anatomy?
• ANATOMY: study of the structure of
organisms… looking at cells, tissues
• (Morphology: Study of form)
What is plant physiology?
• PHYSIOLOGY: study of the function of
cells, tissues, organs of living things;
and the physics/chemistry of these functions…
“Structure correlates to function”
Always keep in mind that in plant anatomy,
morphology & physiology…
• How can water
move from
the ground
all the way
to the top
of a 100 m
tall redwood
tree?
Plant Anatomy: Cells
• Plant cells are basic building blocks
• Can specialize in form and function
• By working together, forming tissues, they can
support each other and survive
• Levels of organization
atoms > molecules > cells > tissues > organs > whole plant > pop.
Plant Tissues Types
All plant organs (roots, stems, leaves) are
composed of the same tissue types.
There are three types of tissue:
• 1. Dermal – outermost layer
• 2. Vascular – conducting tissue, transport
• 3. Ground – bulk of inner layers
1. Dermal tissue
• Epidermis is the outermost layer of cells
• Like the “skin” of animals
• In stems and leaves,
epidermis has cuticle,
a waxy layer that prevents
water loss.
• Some have trichomes, hairs.
• Root epidermis has root hairs, for
water and nutrient absorption
2. Vascular tissue
• Transports water and organic materials (sugars)
throughout the plant
• Xylem – transports water and
dissolved ions from the root
to the stem and leaves.
• Phloem – carries dissolved sugars
from leaves to rest of the plant
Xylem
• Transports water and dissolved minerals
• Tracheids: long, thin tube like structures
without perforations at the ends
• Vessel elements: short, wide tubes perforated at
the ends (together form a pipe, called vessel).
• Both cells have pits (thin sections) on the walls
Tracheids Vessel elements
Xylem cells
• Xylem cells are dead!
• They are hollow cells
and consist
only of
cell wall
Phloem
• Cells that transport organic materials (sugars)
• Phloem cells are ALIVE! (unlike xylem)
• However, they lack
nucleus and
organelles
Phloem: transports sugars
• Phloem composed of cells called sieve tube
members (STM)
• Companion cells join sieve tube members, are
related, and help to load materials into STM
• End walls of STM have large pores called
sieve plates
Sieve tube member
Companion cells
Sieve plates
3. Ground tissue
• Makes up the bulk of plant organs.
• Functions: Metabolism, storage and support.
Root Stem Leaf
Plant Organs
Organs: tissues that act together to serve a
specific function
• Roots
• Stems
• Leaves
Dermal
Vascular
Ground
Dermal
Vascular
Ground
Dermal
Vascular
Ground
Functions of plant organs:
• ROOTS: Anchorage, water/nutrient absorption
from soil, storage, water/nutrient transport
• STEMS: Support, water/nutrient transport
• LEAVES: Photosynthesis (food production)
ROOTS
• ROOTS “the hidden half”
• Functions of roots:
• Ancorage
• Absorption of water & dissolved minerals
• Storage (surplus sugars, starch)
• Conduction water/nutrients
Anatomy of a root
epidermis
cortex
vascular
Root Epidermis
• Outermost, single layer of cells that:
– Protects (from diseases)
– Absorbs water and nutrients
• ROOT HAIRS: tubular extensions
of epidermal cells.
• Increase surface area of root,
for better water/nutrient
absorption
Root Hairs: water and mineral
absorption
Root hairs
increase surface
area for better
absorption
Root Cortex
• Stores starch, sugars and other substances
Root Ground tissue
• In roots, ground tissue (a.k.a. cortex)
provides support, and
often stores sugars and starch
(for example: yams, sweet potato, etc.)
Hey!
I yam
what I
yam,
man!
You’re not a
yam, you’re a
sweetpotato!
cortex
Root Cortex: Endodermis
• Endodermis: the innermost layer of the
cortex
Root cortex: Casparian strip
• The Casparian strip is a water-impermeable
strip of waxy material found in the
endodermis (innermost layer of the cortex).
• The Casparian strip helps to control the
uptake of minerals into the xylem: they have
to go through the cytoplasm of the cell!
STEMS
• Above-ground organs (usually)
• Support leaves and fruits
• Conduct water and sugars
throughout plant (xylem and phloem)
Stem Anatomy
• Bud – undeveloped shoot.
• Node – Location of leaf or bud on stem.
• Internode – Space between nodes.
• Pith – Spongy tissue in the center of the
stem.
• Lenticel – Pore in the outer layer of the
stem.
Stem
Stem Modifications
• Tuber – underground stem with nodes
• Rhizome – underground stem with buds
• Stolon – aboveground stem with shoot
buds
• Bulb – underground stem with fleshy
leaves
• Corm – underground stem with papery
leaves
Modified Stem
Thorn
Tendril
Onion set Rhizome
Modified Stem II
Asparagus Stolon
Stem anatomy
• Dermal, ground and vascular tissues…
pith
cortex
epidermis
Vascular
bundles
Types of Stems
Monocot stem Dicot stem Root
Types of stems
• Herbaceous vs. Woody stems
Tissues of stems
• Epidermis (Dermal tissue type)
• Provides protection
• Has cuticle (wax) prevents water loss
• Trichomes (hairs) for protection, to release
scents, oils, etc.
Stem Vascular tissue
• Vascular bundles – composed of both
xylem and phloem
• Xylem
– Conducts water
– Support
• Phloem
– Conducts food
– Support
Vascular
cambium
Vascular cambium
• Occurs in woody stems
• Vascular cambium located in the middle of
the vascular bundle, between xylem and
phloem
Vascular tissue: Trees
• Vascular tissue is located on the outer layers
of the tree.
wood
phloem
xylem
bark
Vascular
cambium
Girdling: cutting around a tree
• Damages the phloem and xylem, eventually
killing the tree!
Vascular tissue forms rings in trees
• Annual rings: xylem formed by the
vascular cambium during one growing
season
• One ring = one year
History of the tree: annual rings
1489: Tree is planted
by Native American
1492: Columbus lands in
the Americas
1620: Pilgrims land
in Plymouth, Mass.
1776: Declaration
of US independence
1861: Start of
Civil War
1969: Man
lands on Moon
1917 & 1945: Tree
Survives two World
Wars
1971: Birth Year
of the IDIOT
who cut down
this tree!!!
Dendrochronology : tree time-keeping
Ground tissue: Cortex & pith
• Stores food (e.g. potato)
• Site of Photosynthesis (when green)
• Support cells
pith
cortex
LEAVES:
• ‘Photosynthetic factories’ of the plant…
• Function: Photosynthesis – food
production for the whole plant
• Blade: Flat expanded area
• Petiole: stalk that connects
leaf blade to stem, and
transports materials
BLADE
Leaf Anatomy
• Leaf anatomy is correlated to photosynthesis:
Carbon dioxide + Water  sugars + oxygen
dermal
ground
vascular
dermal
Leaf epidermis
• Is transparent – so that sun light can go through.
• Waxy cuticle protects against drying out
• Lower epidermis: stomata with guard cells –
for gas exchange (CO2, H2O in; O2 out)
Leaf epidermis
• Trichomes (give fuzzy texture)
(“Panda plant”)
Leaf vascular tissue
• VEINS  vascular tissue of leaves.
• Veins are composed of xylem (water transport)
phloem (food transport)
and bundle sheaths,
cells surrounding the
xylem/phloem for
strength & support
Leaf Mesophyll
• Middle of the leaf (meso-phyll)
• Composed of photosynthetic ground cells:
• Palisade parenchyma
(long columns below epidermis;
have lots chloroplasts for
photosynthesis)
Spongy parenchyma
(spherical cells)
with air spaces around,
(for gas exchange)
Plant Cells
• Formed at meristems.
– Mitosis in meristem produces new cells.
• Two types of meristems.
– Apical – produces primary growth, ex. tip of
root or shoot.
– Lateral – produces secondary growth, ex.
cambium.
Apical Meristem
Flowers
• Flower Parts -- Male
– Stamen: Male part of the flower.
– Filament: Stalk like in the stamen that holds up
the anther
– Anther: Sack-like structure that contains
pollen.
Flowers
• Flower Parts -- Male
– Pollen grains are released from the anther that
contains sperm.
– Staminate: Flowers that have only male parts.
Flowers
• Flower Parts – Female
– Pistil: Female part of the flower
– Stigma: Sticky part of the pistil that is
receptive to pollen.
– Style: Rod shaped middle part that has a
swollen base (ovary) containing eggs
Flowers
• Flower Parts – Neither male or female
– Petals: colorful leaf-like structures which
attract animals and insects.
– Corolla: When all of the petals are fused
together.
– Sepals: Green leaves that protect the flower
before it opens.
Flowers
• Flower Parts – Niether male or female
– Calyx: When all of the sepals are fused
together.
Flowers
• Flower Types:
– Perfect Flower: Has both male and female
parts.
– Imperfect Flower: A flower that is missing
either male or female parts.
– Complete Flower: Flowers that have sepals,
petals, pistils, and stamens.
Flowers
• Flower Types:
– Incomplete Flowers: When a flower is missing
sepals, petals, pistils, or stamen.
– Imperfect Flowers are always incomplete.
Incomplete flowers may or may not be
imperfect
From ovary to fruit
• The ovary of the flower contains the
ovules.
• As fertilized ovules develop into seeds,
the ovary wall develops into the fruit.
• In science, the term “fruit” refers to a
mature ovary that contains seeds.
Fruit anatomy
Types of dry fruits
Legume
(Bean pod)
Capsule
(Poppy)
Achene
(Sunflower)
Silique
(Money Plant)
Follicle
(Columbine)
Nut
(Hazelnut)
Types of fleshy fruits
Drupe
(Peach)
Pome
(Apple)
Pepo
(Cucumber)
Aggregate
(Strawberry)
Multiple
(Pineapple)
Berry
(Tomato)
Fruit dispersal
• The form of the fruit gives clues about its
dispersal.
• Small, dry fruits with “wings” or
“parachutes” may be wind-dispersed.
Fleshy fruits are often animal dispersed.
Explosive fruits can fling seeds away.
Floating fruits may be water dispersed.
How are these fruits dispersed?
Dandelion Coconut
Maple
Cocklebur Jewelweed
Thinking question:
• Why must fruits be dispersed away from
the parent plant?
Seeds
Ovule to seed
Mature Seed
Seed anatomy
Seed dormancy
• Seeds can remain dormant in the soil for
long periods of time. Dormancy helps
ensure that seeds only germinate when
conditions are right.
• When we weed or cultivate a bare patch of
soil, the weeds that sprout up immediately
usually come from the “seed bank”
already in the soil.
Breaking dormancy
• Seeds require moisture and the right
temperature to germinate.
• In addition, some seeds germinate only
after certain environmental signals:
– Drying
– Temperature (period of cold or heat)
– Disruption of the seed coat
Thinking question
• What could be the advantage of waiting
for each of these signals to germinate?
– Long period of cold
– High heat of a forest fire
– Drying out
– Disruption of the seed coat.
Germination: monocot
Germination: dicot
• Is a seed alive? Is a fruit alive? Answer as
completely as you can on your own paper.
(Hang on to your paper until the end of
class.)
W
O
R
K
T
O
G
E
T
H
E
R
Plant Life Cycle
Asexual Reproduction
• Asexual reproduction is natural “cloning.”
Parts of the plant, such as leaves or stems,
produce roots and become an independent
plant.
• List some benefits and some drawbacks to
asexual reproduction.
Sexual Reproduction
• Sexual reproduction requires fusion of
male cells in the pollen grain with female
cells in the ovule.
• List some advantages and drawbacks to
sexual reproduction.
Terms to know:
• Haploid: having a single set of
chromosomes in each cell.
• Diploid: having two sets of
chromosomes in each cell.
• Mitosis: cell division, which produces
two genetically identical cells.
• Meiosis: reduction division, which
produces four haploid reproductive
cells.
Animals vs. Plants
Plant Reproduction
Animal
Reproduction
Life cycle
Alternation of
generations
No alternation of
generations
Gametes Haploid gametes Haploid gametes
Spores Haploid spores No spores
Gametes made
by
Haploid gametophyte,
by mitosis
Diploid organism, by
meiosis
Spores made
by
Diploid sporophyte, by
meiosis
No spores
Alternation of Generations
• Plants have a double life cycle with two
distinct forms:
– Sporophyte: diploid, produce haploid spores
by meiosis.
– Gametophyte: haploid, produce gametes by
mitosis.
Non-flowering plants
• Mosses, ferns, and related plants have
motile, swimming sperm.
• What kind of environmental conditions
would be required for reproduction in
these plants?
• What kinds of limits does external
reproduction impose on these plants?
Moss Life Cycle
Fern Life Cycle
Conifers
• Conifers (also non-flowering plants) have
reduced gametophytes.
– Male gametophyte is contained in a dry pollen
grain.
– Female gametophyte is a few cells inside of
the structures that become the seed.
Conifer life cycle
Conifer pollination
• Conifers are wind-pollinated plants.
• Chance allows some pollen to land on the
scales of female cones.
• Pollen germinates, grows a pollen tube
into the egg to allow sperm to fertilize the
egg.
• What are some advantages and
disadvantages to wind pollination?
Pollen go-betweens
• Showy flowers are the result of selection
for more efficient pollination strategies.
• Flower parts are modified leaves. Those
that were brightly colored attracted insects
in search of pollen.
• Why would insects search for pollen?
What other rewards do flowers offer?
• What are advantages and disadvantages to
relying on insects as pollinators?
Flowers
Flower Parts
Imperfect flowers
Flowers
• Sexual Reproduction in Plants: Two parents
(meiosis)
– #1 The stamen releases pollen.
– #2 Pollen is carried by wind, gravity, animals, or
insects to the stigma of another flower. (This is when
pollination occurs)
– #3 The pollen moves from the stigma down through in
a pollen tube the style depositing sperm in the ovary.
Flowers
• Sexual Reproduction in Plants:
– #4 When the sperm has been deposited in the
ovary fertilization has occurred.
– #5 When the eggs have been fertilized, the
ovary and surrounding tissue start to enlarge to
become a fruit and the fertilized eggs become
seeds.
Gametogenesis: Male
Gametogenesis: Female
Double Fertilization
Flower to Fruit
Ovule to Seed
Seed Anatomy
Growth Cycle
• Annual – single season
• Biennial – two seasons
• Perennial – multiple seasons
• Evergreen – leaves persist > 2 seasons
• Deciduous – leaves die in cold or dry