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Naveed Akhtar
Assistant Professor of Zoology
Introduction
 Animal behavior includes all the ways animals interact
with other organisms and the physical environment.
 Behavior can also be defined as a change in the activity of
an organism in response to a stimulus, an external or
internal cue or combo of cues
 Behavior refers to the varied activities that an animal
performs during its lifetime. Internal physiological
conditions, environmental stimuli, and social situations
influence specific behavioral responses

 Proximate factors that influence the behavior of an
animal include genetics, developmental experiences,
and the current environment, including photoperiod,
season, and temperature
 The ultimate factor that influences the behavior of an
animal is natural selection. Animals that possess certain
traits are more successful at surviving and reproducing.
 Some behaviors are innate, or genetically hardwired,
while others are learned, or developed through
experience. In many cases, behaviors have both an innate
component and a learned component.
 Individual Behaviour: Individual behaviour
includes those related to movement or physiological
regulation
 Social Behaviour: Social behaviour is related to
interactions with other organisms. This includes
behaviours related to courtship, reproduction,
parenting, feeding, defence strategies as well as
communication within and between groups of animals
Types of Behaviour
 The following the three categories of behavioral
pattern of animals. The categories are:
 1. Instinctive Behaviors or Fixed Action Pattern
(FAP)
 2. Learning Behaviour
 3. Complex Behaviour
Instinctive Behaviors or Fixed
Action Pattern (FAP)
 Innate behaviour is generally any behaviour that is not
learned.
 Instinct is the innate ability to complete a specific
behaviour pattern with no conscious intention.
 A releaser is a specific signal called a sign stimulus that
results in an innate behavioral response
 Instinctive Behaviours are genetically inherited
characteristics that impel animals to behave in a certain
fixed way.
 It is also referred to as Fixed Action Pattern (FAP) or
Innate Behaviour or Inborn Behaviour or Inherent
Behaviour.
 Instinct is generally described as patterns of inherited
pre-set behavioural responses which develop along
with the developing nervous system.
 In the animal kingdom, there are a number of
behavioural patterns which are depicted in the genes.
To name a few — the courtship display and mating in
most animals, feeding patterns, nest building, parental
care, singing, wings cleaning, territoriality and
aggression, construction of web by spiders
construction of nest by birds etc..
 Examples
 1. During the breeding season a peacock starts dancing
as soon as it sees a peahen. The dancing act is not
learnt by watching.
 2. A weaver bird never learns to construct a nest. It
simply knows it.
 3. A tailor bird brings two leaves together and stitches
them with a long flexible grass and then cushions the
bottom with soft grasses, where it lays her eggs. This
vital information were passed on to them through the
genes of their parents.
Tailor bird nest building
 4. Bees do not learn to build their hive, this property is
acquired genetically.
 Another very interesting example of instinctive
behaviour or FAP is provided by the Mason wasp
(Manobia sp). A female wasp, after emerging from her
cell, has a brief moment of interaction with another
wasp when she mates with a male.
 Later, totally on her own, she selects a hollow stem and
builds at its inner end, a partition of mud mixed with
her saliva
 Close to this partition she lays an egg.
 Thereafter, she hunts for caterpillars which she lightly
paralyses with her sting.
 She carries 5-8 of them to the cell which would act as
future food supply for the larva when it hatches.
 After being satisfied, she builds another partition
sealing off the egg with its food supply.
 nstinct is understood as an innate behaviour
mechanism that impel animals to behave in a certain
fixed way. The idea of Fixed Action Pattern was
formulated by Konrad Lorenz (1958).
Sign Stimulus or Releaser
 FAP is a specific and stereotyped sequence of activities
that are triggered by particular stimulus called sign
stimulus or releaser.
 Definite sign stimulus or releaser works and initiates
instinctive acts.
Examples
 1.The red belly in the males of three-spined stickleback
fish is one specific sign stimulus, responsible for
releasing territorial defence and other reproductive
activities.
 2. The red spot present on the bill of herring gull is
pecked by its chicks which acts as a sign stimulus. This
induces the parent to regurgitate food called crop
milk.
 3. When the parent of song birds alights on the nest
with food in their beak, it creates a jerk. This acts as a
sign stimuli for the begging behaviour of the chicks.
Characteristics of Instincts
 1. Instincts are behaviours that are basically
determined and transferred through genes.
 2. Instincts are similar in all individuals of a species.
 3. Instincts involve complex and highly rigid pattern of
behaviour involving numerous muscles, muscle
groups, organs and systems that function in an entirely
coordinated manner.
 4. Instinctive actions are specific and stereotyped
sequence of activities that are triggered by complex
stimuli called sign stimuli.
Learned Behaviors
 What is learning?
 The modification of behavior through experience
 In general, a learned behavior is one that an organism
develops as a result of experience.
 Learned behaviors contrast with innate behaviors, which
are genetically hardwired and can be performed without
any prior experience or training.
 Some behaviors have both learned and innate elements.
Example# 1 , Zebra finches are genetically
preprogrammed to learn a song, but the song they sing
depends on what they hear from their fathers.
 Example# 1
 Young bobcats raised in isolation without the chance
to catch live prey did not attack a white rat placed with
them, unless the rat tried to escape.
 At first, their attacks were not efficient, but after some
experience, they were seizing prey by the neck and
rapidly killing them.
 Apparently, learning improve inherited components of
this behavior
 Example # 3
 Another example involving instinctive and learned
components to behavior is the nut-cracking
behavior of squirrels.
 Squirrels gnaw and pry to open a nut. Inexperienced
squirrels are not efficient; they gnaw and pry at
random on the nut.
 Experienced squirrels, however, gnaw a furrow on the
broad side, then wedge their lower incisors into the
furrow and crack the nut open
Examples of Instinctive and
Learned Combination
Types of Learned Behaviour
 Simple Learned Behaviour
 Conditioned behaviors
 1. Habituation
 Habituationi s the simplest and perhaps most common type of
behavior in many different animals
 Habituation involves a wanin or decrease in response to repeated
or continuous stimulation
 This is a form of non-associative learning, meaning that the
stimulus is not linked with any punishment or reward.
 An animal learns not to respond to stimuli in its
environment that are constant and probably relatively
unimportant
Conti….
 By habituating to unimportant stimuli, an animal
conserves energy and time that are better spent on other
important function
 Example:
 Prairie dogs typically sound an alarm call when threatened
by a predator.
 At first, they will give this alarm call in response to hearing
human steps, which indicate the presence of a large and
potentially hungry animal
 However, the prairie dogs gradually become habituated to
the sound of human footsteps, as they repeatedly
experience the sound without anything bad happening
Prairie dogs Habituation
2. Imprinting
 Imprinting is a simple and highly specific type of
learning that occurs at a particular age or life stage
during the development of certain animals, such as
ducks and geese
 The attachment usually forms only during a specific
critical period soon after hatching or birth and is not
reversible
 Imprinting is a rapid learning process that apparently
occurs without reinforcement.
 Example
 Konrad Lorenz (1903–1989) an Austrian zoologist,
ethologist, and ornithologist conducted experiments
with geese in which he allowed the geese to imprint on
him.
 The goslings followed him as though he was their mother.
In nature, many species of birds in which the young follow
the parent soon after birth use imprinting so that the
young can identify with or recognize their parent
 They can then be led successfully to the nest or to water.
Both visual and auditory cues are important in imprinting
systems.
Conditioned Behaviors
 Conditioned behaviors are types of associative
learning where a stimulus becomes associated
with a consequence
 Conditioned behaviors are the result of associative
learning, which takes two forms;
 Classical conditioning
 Operant conditioning
Classical Conditioning
 A type of learning in which a neutral stimulus comes
to bring about a response after it is paired with a
stimulus that naturally brings that response
 Classical conditioning is a type of learning
documented by Russian physiologist, Ivan Pavlov
(1849–1936).
 The “Pavlov’s Dog” Experiment
 In his classic experiment on the salivary reflex in dogs,
Pavlov presented food right after the sound of a bell
 After a number of such presentations, the dogs were
conditioned—they associated the sound of the
bell with food. It was then possible to elicit the dog’s
usual response to food—salivation—with just the
sound of the bell
 The food was a positive reinforcement for salivating
behavior, but responses could also be conditioned
using negative reinforcement
The “Pavlov’s Dog” Experiment
INSTRUMENTAL CONDITIONING
 In instrumental conditioning (also known as trial-
and-error learning), the animal learns while carrying
out certain searching actions, such as walking and
moving about
 Example
 If the animal find food during these activities, the food
reinforces the behavior, and the animal associates the
reward with the behavior.
 This association is repeated several times, the
animal learns that the behavior leads to
reinforcement
 Example:
 A classic example of instrumental conditioning is that
of a rat in a “Skinner box,” developed by B. F. Skinner
(1904–1990), a prominent psychologist
 When placed in the box, the rat begins to explore. It
moves all about the box and, by accident, eventually
presses a lever and is rewarded with a food pellet.
 Because food rewards are provided each time the rat
presses the lever, the rat associates the reward with the
behavior.
 Through repetition, the rat learns to press the lever
right away to receive the reward. In this type of
learning, the animal is instrumental in providing its
own reinforcement.
 In instrumental conditioning, providing the
reinforcement (food) whenever the animal comes close
to the lever and continuing to supply reinforcement
when the animal touches the lever “shapes” the
behavior. Finally, the animal learns to press the lever to
obtain food.
 Young animals’ attempts to learn new motor patterns
often involve instrumental conditioning. A young bird
learning to fly or a young mammal at play may improve
coordination of certain movements or behavior
patterns by practice during these activities.
LATENT LEARNING
 Latent learning, sometimes called exploratory
learning, involves making associations without
immediate reinforcement or reward. The reward is not
obvious.
 An animal is apparently motivated, however, to learn
about its surroundings
 Example:
 If a rat is placed in a maze that has no food or reward,
it explores the maze, although rather slowly.
 If food or another reward is provided, the rat quickly
runs the maze.
 Apparently, previous learning of the maze had
occurred but remained latent, or hidden, until an
obvious reinforcement was provided.
 Latent learning allows an animal to learn about its
surroundings as it explores. Knowledge about an
animal’s home area may be important for its survival,
perhaps enabling it to escape from a predator or
capture prey.
Example: 2
INSIGHT LEARNING
 What is insight learning?
 In insight learning or intellectual learning, the
animal uses cognitive or mental processes to associate
experiences and solve problems
 Example:
 The classic example is the work of Wolfgang
Kohler (1887–1967) on chimpanzees that were
trained to use tools to obtain food rewards
Stages of Insight Learning
 One chimpanzee was given some bamboo poles that
could be joined to make a longer pole, and some
bananas were hung from the ceiling.
 Once the chimp formed the longer pole, it used the
pole to knock the bananas to the cage floor. Kohler
believed that the animal used insight learning to get
the bananas
 In addition, Jane van Lawick-Goodall (1934– ) has
observed chimpanzees in the wild using tools to
accomplish various tasks. For example, they use
crumpled leaves as a sponge for drinking water
END OF CHAPTER

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Kinds of Animal Behavior.pdf

  • 2. Introduction  Animal behavior includes all the ways animals interact with other organisms and the physical environment.  Behavior can also be defined as a change in the activity of an organism in response to a stimulus, an external or internal cue or combo of cues  Behavior refers to the varied activities that an animal performs during its lifetime. Internal physiological conditions, environmental stimuli, and social situations influence specific behavioral responses 
  • 3.  Proximate factors that influence the behavior of an animal include genetics, developmental experiences, and the current environment, including photoperiod, season, and temperature  The ultimate factor that influences the behavior of an animal is natural selection. Animals that possess certain traits are more successful at surviving and reproducing.  Some behaviors are innate, or genetically hardwired, while others are learned, or developed through experience. In many cases, behaviors have both an innate component and a learned component.
  • 4.  Individual Behaviour: Individual behaviour includes those related to movement or physiological regulation  Social Behaviour: Social behaviour is related to interactions with other organisms. This includes behaviours related to courtship, reproduction, parenting, feeding, defence strategies as well as communication within and between groups of animals
  • 5. Types of Behaviour  The following the three categories of behavioral pattern of animals. The categories are:  1. Instinctive Behaviors or Fixed Action Pattern (FAP)  2. Learning Behaviour  3. Complex Behaviour
  • 6.
  • 7. Instinctive Behaviors or Fixed Action Pattern (FAP)  Innate behaviour is generally any behaviour that is not learned.  Instinct is the innate ability to complete a specific behaviour pattern with no conscious intention.  A releaser is a specific signal called a sign stimulus that results in an innate behavioral response  Instinctive Behaviours are genetically inherited characteristics that impel animals to behave in a certain fixed way.  It is also referred to as Fixed Action Pattern (FAP) or Innate Behaviour or Inborn Behaviour or Inherent Behaviour.
  • 8.  Instinct is generally described as patterns of inherited pre-set behavioural responses which develop along with the developing nervous system.  In the animal kingdom, there are a number of behavioural patterns which are depicted in the genes. To name a few — the courtship display and mating in most animals, feeding patterns, nest building, parental care, singing, wings cleaning, territoriality and aggression, construction of web by spiders construction of nest by birds etc..
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  • 10.  Examples  1. During the breeding season a peacock starts dancing as soon as it sees a peahen. The dancing act is not learnt by watching.  2. A weaver bird never learns to construct a nest. It simply knows it.  3. A tailor bird brings two leaves together and stitches them with a long flexible grass and then cushions the bottom with soft grasses, where it lays her eggs. This vital information were passed on to them through the genes of their parents.
  • 11. Tailor bird nest building
  • 12.  4. Bees do not learn to build their hive, this property is acquired genetically.  Another very interesting example of instinctive behaviour or FAP is provided by the Mason wasp (Manobia sp). A female wasp, after emerging from her cell, has a brief moment of interaction with another wasp when she mates with a male.  Later, totally on her own, she selects a hollow stem and builds at its inner end, a partition of mud mixed with her saliva
  • 13.  Close to this partition she lays an egg.  Thereafter, she hunts for caterpillars which she lightly paralyses with her sting.  She carries 5-8 of them to the cell which would act as future food supply for the larva when it hatches.  After being satisfied, she builds another partition sealing off the egg with its food supply.  nstinct is understood as an innate behaviour mechanism that impel animals to behave in a certain fixed way. The idea of Fixed Action Pattern was formulated by Konrad Lorenz (1958).
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  • 15. Sign Stimulus or Releaser  FAP is a specific and stereotyped sequence of activities that are triggered by particular stimulus called sign stimulus or releaser.  Definite sign stimulus or releaser works and initiates instinctive acts.
  • 16. Examples  1.The red belly in the males of three-spined stickleback fish is one specific sign stimulus, responsible for releasing territorial defence and other reproductive activities.  2. The red spot present on the bill of herring gull is pecked by its chicks which acts as a sign stimulus. This induces the parent to regurgitate food called crop milk.  3. When the parent of song birds alights on the nest with food in their beak, it creates a jerk. This acts as a sign stimuli for the begging behaviour of the chicks.
  • 17. Characteristics of Instincts  1. Instincts are behaviours that are basically determined and transferred through genes.  2. Instincts are similar in all individuals of a species.  3. Instincts involve complex and highly rigid pattern of behaviour involving numerous muscles, muscle groups, organs and systems that function in an entirely coordinated manner.  4. Instinctive actions are specific and stereotyped sequence of activities that are triggered by complex stimuli called sign stimuli.
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  • 19. Learned Behaviors  What is learning?  The modification of behavior through experience  In general, a learned behavior is one that an organism develops as a result of experience.  Learned behaviors contrast with innate behaviors, which are genetically hardwired and can be performed without any prior experience or training.  Some behaviors have both learned and innate elements. Example# 1 , Zebra finches are genetically preprogrammed to learn a song, but the song they sing depends on what they hear from their fathers.
  • 20.  Example# 1  Young bobcats raised in isolation without the chance to catch live prey did not attack a white rat placed with them, unless the rat tried to escape.  At first, their attacks were not efficient, but after some experience, they were seizing prey by the neck and rapidly killing them.  Apparently, learning improve inherited components of this behavior
  • 21.  Example # 3  Another example involving instinctive and learned components to behavior is the nut-cracking behavior of squirrels.  Squirrels gnaw and pry to open a nut. Inexperienced squirrels are not efficient; they gnaw and pry at random on the nut.  Experienced squirrels, however, gnaw a furrow on the broad side, then wedge their lower incisors into the furrow and crack the nut open
  • 22. Examples of Instinctive and Learned Combination
  • 23. Types of Learned Behaviour  Simple Learned Behaviour  Conditioned behaviors  1. Habituation  Habituationi s the simplest and perhaps most common type of behavior in many different animals  Habituation involves a wanin or decrease in response to repeated or continuous stimulation  This is a form of non-associative learning, meaning that the stimulus is not linked with any punishment or reward.  An animal learns not to respond to stimuli in its environment that are constant and probably relatively unimportant
  • 24. Conti….  By habituating to unimportant stimuli, an animal conserves energy and time that are better spent on other important function  Example:  Prairie dogs typically sound an alarm call when threatened by a predator.  At first, they will give this alarm call in response to hearing human steps, which indicate the presence of a large and potentially hungry animal  However, the prairie dogs gradually become habituated to the sound of human footsteps, as they repeatedly experience the sound without anything bad happening
  • 26. 2. Imprinting  Imprinting is a simple and highly specific type of learning that occurs at a particular age or life stage during the development of certain animals, such as ducks and geese  The attachment usually forms only during a specific critical period soon after hatching or birth and is not reversible  Imprinting is a rapid learning process that apparently occurs without reinforcement.
  • 27.  Example  Konrad Lorenz (1903–1989) an Austrian zoologist, ethologist, and ornithologist conducted experiments with geese in which he allowed the geese to imprint on him.  The goslings followed him as though he was their mother. In nature, many species of birds in which the young follow the parent soon after birth use imprinting so that the young can identify with or recognize their parent  They can then be led successfully to the nest or to water. Both visual and auditory cues are important in imprinting systems.
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  • 29. Conditioned Behaviors  Conditioned behaviors are types of associative learning where a stimulus becomes associated with a consequence  Conditioned behaviors are the result of associative learning, which takes two forms;  Classical conditioning  Operant conditioning
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  • 31. Classical Conditioning  A type of learning in which a neutral stimulus comes to bring about a response after it is paired with a stimulus that naturally brings that response  Classical conditioning is a type of learning documented by Russian physiologist, Ivan Pavlov (1849–1936).  The “Pavlov’s Dog” Experiment  In his classic experiment on the salivary reflex in dogs, Pavlov presented food right after the sound of a bell
  • 32.  After a number of such presentations, the dogs were conditioned—they associated the sound of the bell with food. It was then possible to elicit the dog’s usual response to food—salivation—with just the sound of the bell  The food was a positive reinforcement for salivating behavior, but responses could also be conditioned using negative reinforcement
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  • 35. INSTRUMENTAL CONDITIONING  In instrumental conditioning (also known as trial- and-error learning), the animal learns while carrying out certain searching actions, such as walking and moving about  Example  If the animal find food during these activities, the food reinforces the behavior, and the animal associates the reward with the behavior.  This association is repeated several times, the animal learns that the behavior leads to reinforcement
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  • 37.  Example:  A classic example of instrumental conditioning is that of a rat in a “Skinner box,” developed by B. F. Skinner (1904–1990), a prominent psychologist  When placed in the box, the rat begins to explore. It moves all about the box and, by accident, eventually presses a lever and is rewarded with a food pellet.  Because food rewards are provided each time the rat presses the lever, the rat associates the reward with the behavior.
  • 38.  Through repetition, the rat learns to press the lever right away to receive the reward. In this type of learning, the animal is instrumental in providing its own reinforcement.  In instrumental conditioning, providing the reinforcement (food) whenever the animal comes close to the lever and continuing to supply reinforcement when the animal touches the lever “shapes” the behavior. Finally, the animal learns to press the lever to obtain food.
  • 39.  Young animals’ attempts to learn new motor patterns often involve instrumental conditioning. A young bird learning to fly or a young mammal at play may improve coordination of certain movements or behavior patterns by practice during these activities.
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  • 42. LATENT LEARNING  Latent learning, sometimes called exploratory learning, involves making associations without immediate reinforcement or reward. The reward is not obvious.  An animal is apparently motivated, however, to learn about its surroundings  Example:  If a rat is placed in a maze that has no food or reward, it explores the maze, although rather slowly.
  • 43.  If food or another reward is provided, the rat quickly runs the maze.  Apparently, previous learning of the maze had occurred but remained latent, or hidden, until an obvious reinforcement was provided.  Latent learning allows an animal to learn about its surroundings as it explores. Knowledge about an animal’s home area may be important for its survival, perhaps enabling it to escape from a predator or capture prey.
  • 45. INSIGHT LEARNING  What is insight learning?  In insight learning or intellectual learning, the animal uses cognitive or mental processes to associate experiences and solve problems  Example:  The classic example is the work of Wolfgang Kohler (1887–1967) on chimpanzees that were trained to use tools to obtain food rewards
  • 46. Stages of Insight Learning
  • 47.  One chimpanzee was given some bamboo poles that could be joined to make a longer pole, and some bananas were hung from the ceiling.  Once the chimp formed the longer pole, it used the pole to knock the bananas to the cage floor. Kohler believed that the animal used insight learning to get the bananas
  • 48.  In addition, Jane van Lawick-Goodall (1934– ) has observed chimpanzees in the wild using tools to accomplish various tasks. For example, they use crumpled leaves as a sponge for drinking water
  • 49.