The document discusses rhizosphere and rhizoplane microflora. It defines the rhizosphere microbiome as the microbial population in the area surrounding plant roots, which is much higher than in bulk soil. It provides examples of common microorganisms found in the rhizosphere like bacteria, fungi, protozoa. Rhizosphere microbes can positively impact plants by increasing nutrient supply, stimulating growth, and preventing pathogens. However, they can also negatively affect plants through competition for resources and by acting as pathogens themselves. The document also discusses actinorhizal bacteria like Frankia that can form symbiotic nitrogen-fixing nodules on roots of certain plants.
The rhizosphere is the region of soil surrounding plant roots that is influenced by root secretions like mucilage, exudates, and lysates. It contains many microorganisms in complex relationships with the plant roots. Root secretions, collectively known as rhizodeposition, enrich the soil environment and stimulate microbial growth in the rhizosphere compared to bulk soil, as measured by the R:S ratio of microorganisms. Rhizodeposition includes a variety of organic compounds that influence soil nutrients and microbes.
Microbial interactions are ubiquitous, diverse, critically important in the function of any biological community.
The most common cooperative interactions seen in microbial systems are mutually beneficial. The interactions between the two populations are classified according to whether both populations and one of them benefit from the associations, or one or both populations are negatively affected.
this presentation show details regarding how the concept of agricultural microbiology came into existance and also the contribution of various scientists
This document provides an overview of mycorrhiza, which is a symbiotic relationship between fungi and plant roots. It defines mycorrhiza and explains that 95% of plant species form these relationships. It then classifies and describes the main types of mycorrhizal associations like ectomycorrhiza, endomycorrhiza, and orchid mycorrhiza. The document outlines the importance and benefits of mycorrhizal relationships for plant growth and health. It also discusses methods for isolating, mass producing, and applying mycorrhizal fungi.
This document summarizes a seminar presentation about plant-microbe interactions given by Manisha Thakur. It discusses how plants constantly encounter biotic and abiotic stresses. Microbes that colonize plants can have pathogenic, symbiotic, or associative relationships. Specific examples provided include mutualistic relationships like rhizobia in root nodules, and types of pathogenic relationships such as necrotrophy and biotrophy. The document also discusses concepts like the rhizosphere and how root exudates influence microbial communities in and around plant roots and leaves.
This document discusses plant growth promoting rhizobacteria (PGPR). It begins by defining PGPR as beneficial bacteria that colonize plant roots and promote plant growth. It then covers the classification, characteristics, and mechanisms of action of PGPR, including direct mechanisms like nitrogen fixation, phosphate solubilization, and phytohormone production as well as indirect mechanisms like siderophore production and induced systemic resistance. The document also discusses the roles, commercialization, and importance of PGPR as biofertilizers for sustainable agriculture.
The rhizosphere is the region of soil surrounding plant roots that is influenced by root secretions like mucilage, exudates, and lysates. It contains many microorganisms in complex relationships with the plant roots. Root secretions, collectively known as rhizodeposition, enrich the soil environment and stimulate microbial growth in the rhizosphere compared to bulk soil, as measured by the R:S ratio of microorganisms. Rhizodeposition includes a variety of organic compounds that influence soil nutrients and microbes.
Microbial interactions are ubiquitous, diverse, critically important in the function of any biological community.
The most common cooperative interactions seen in microbial systems are mutually beneficial. The interactions between the two populations are classified according to whether both populations and one of them benefit from the associations, or one or both populations are negatively affected.
this presentation show details regarding how the concept of agricultural microbiology came into existance and also the contribution of various scientists
This document provides an overview of mycorrhiza, which is a symbiotic relationship between fungi and plant roots. It defines mycorrhiza and explains that 95% of plant species form these relationships. It then classifies and describes the main types of mycorrhizal associations like ectomycorrhiza, endomycorrhiza, and orchid mycorrhiza. The document outlines the importance and benefits of mycorrhizal relationships for plant growth and health. It also discusses methods for isolating, mass producing, and applying mycorrhizal fungi.
This document summarizes a seminar presentation about plant-microbe interactions given by Manisha Thakur. It discusses how plants constantly encounter biotic and abiotic stresses. Microbes that colonize plants can have pathogenic, symbiotic, or associative relationships. Specific examples provided include mutualistic relationships like rhizobia in root nodules, and types of pathogenic relationships such as necrotrophy and biotrophy. The document also discusses concepts like the rhizosphere and how root exudates influence microbial communities in and around plant roots and leaves.
This document discusses plant growth promoting rhizobacteria (PGPR). It begins by defining PGPR as beneficial bacteria that colonize plant roots and promote plant growth. It then covers the classification, characteristics, and mechanisms of action of PGPR, including direct mechanisms like nitrogen fixation, phosphate solubilization, and phytohormone production as well as indirect mechanisms like siderophore production and induced systemic resistance. The document also discusses the roles, commercialization, and importance of PGPR as biofertilizers for sustainable agriculture.
This document discusses plant growth promoting rhizobacteria (PGPR) and their ability to solubilize inorganic phosphate. Some key points:
- PGPR are bacteria that live in the rhizosphere and provide benefits to plants. An important function is solubilizing insoluble phosphate minerals making phosphorus available for plant uptake.
- Common insoluble phosphates include tricalcium phosphate, dicalcium phosphate, and hydroxyapatite. Bacteria secrete organic acids like lactic acid and acetic acid to solubilize these minerals.
- Successful phosphate solubilizing bacteria include species from Bacillus, Pseudomonas, and Rhizobium genera. Screening methods involve checking for clearing zones
The document discusses the rhizosphere and phyllosphere, which are the regions of soil and plant surfaces influenced by microorganisms. The rhizosphere refers to the region of soil directly influenced by root secretions and microbes. It includes the inner and outer rhizosphere zones. Microbes in the rhizosphere play important roles in plant nutrition, growth promotion, and disease suppression. The phyllosphere refers to the interface between leaves and air, and is inhabited by bacteria, yeast and fungi that can benefit plants through nutrient management, disease control and stress tolerance.
he rhizosphere is the narrow region of soil or substrate that is directly influenced by root secretions and associated soil microorganisms known as the root microbiome.
The phyllosphere is a term used in microbiology to refer to the total above-ground portions of plants as habitat for microorganisms.
This document discusses the microbial flora of soil. It defines soil microflora as microorganisms that contribute to the biological properties of soil, including bacteria, fungi, algae, and protozoa. The five major groups interact and form a soil food web, with bacteria and fungi serving important roles in decomposition. Physical and chemical factors like pH and nutrients influence the growth and activity of soil microflora. The microflora are then classified and examples are given of bacteria, fungi, algae, protozoa, and their functions in the soil ecosystem.
This document discusses plant growth-promoting microbes (PGPM) which are soil microorganisms that can positively influence plant growth. PGPM are divided into two main groups - plant growth-promoting rhizobacteria (PGPR) and plant growth-promoting fungi (PGPF). PGPR and PGPF help plants grow through mechanisms like nitrogen fixation, phosphate solubilization, siderophore production, phytohormone synthesis, and protecting plants from pathogens. They play important roles in agricultural sustainability by improving soil fertility, supplying nutrients, suppressing diseases, and assisting in environmental remediation.
Soil microbiology is the study of microorganisms in soil such as bacteria, actinomycetes, fungi, algae and protozoa. These microorganisms are important because they affect soil structure and fertility through organic matter decomposition, nutrient transformations, and symbiotic relationships with plants. The four major groups of microbes found in soil are bacteria, actinomycetes, fungi, and algae, each playing an important role in soil health and plant growth.
This document provides an overview of plant growth promoting rhizobacteria (PGPR). It discusses that PGPR are a group of soil bacteria that colonize plant roots and enhance plant growth directly or indirectly. Direct mechanisms include biological nitrogen fixation, phosphate solubilization, phytohormone production, siderophore production, and antibiotic production. Indirect mechanisms include inducing systemic resistance in plants, production of lytic enzymes, and stress tolerance effects. The document reviews the specific mechanisms of several important PGPR functions and commercially available PGPR products.
soil characteristics influencing growth and activity of microfloraIGKV
This document discusses soil characteristics that influence the growth and activity of microflora in soil. It outlines several key factors: soil moisture and temperature are essential for microbial life and activity; most bacteria, algae and protozoa prefer neutral-slightly alkaline pH between 4.5-8.0; aerobic microbes require oxygen while anaerobic microbes do not; cultural practices like tilling and crop rotation alter microbial populations; root exudates and soil organic matter provide nutrients and food for microbes; microbial interactions can be symbiotic or antagonistic; and the physical, chemical and nutrient properties of different soil types impact microbial communities.
Soil organic matter has long been recognized as one of the most important components in maintaining soil fertility, soil quality, and agricultural sustainability. The soil zone strongly influenced by plant roots, the rhizosphere, plays an important role in regulating soil organic matter decomposition and nutrient cycling. Processes that are largely controlled or directly influenced by roots are often referred to as rhizosphere processes. These processes may include exudation of soluble compounds, water uptake, nutrient mobilization by roots and microorganisms, rhizosphere-mediated soil organic matter decomposition, and the subsequent release of CO2 through respiration. Rhizosphere processes are major gateways for nutrients and water. At the global scale, rhizosphere processes utilize approximately 50% of the energy fixed by photosynthesis in terrestrial ecosystems, contribute roughly 50% of the total CO2 emitted from terrestrial ecosystems, and mediate virtually all aspects of nutrient cycling. Therefore, plant roots and their rhizosphere interactions are at the center of many ecosystem processes. However, the linkage between rhizosphere processes and soil organic matter decomposition is not well understood. Because of the lack of appropriate methods, rates of soil organic matter decomposition are commonly assessed by incubating soil samples in the absence of vegetation and live roots with an implicit assumption that rhizosphere processes have little impact on the results. Our recent studies have overwhelmingly proved that this implicit assumption is often invalid, because the rate of soil organic matter decomposition can be accelerated by as much as 380% or inhibited by as much as 50% by the presence of live roots. The rhizosphere effect on soil organic matter decomposition is often large in magnitude and significant in mediating plant-soil interactions.
Mycorrhiza Biofertilizer is also known as VAM (Myco = Fungal + rrhiza = roots) adheres to plants rhizoids leading to development of hyphae. Hyphae boost development and spreading of white root in to soil leading to significant increase in rhizosphere. These hyphae further penetrate and form arbuscules within the root cortical. VAM fungi form a special symbiotic relationship with roots of plant that can enhance growth and survivability of colonized plants. Mycorrhiza Biofertilizer is very useful in organic farming as well as normal commercial farming
This document discusses Azotobacter, a genus of nitrogen-fixing bacteria that can be used as a biofertilizer. It describes the key species of Azotobacter, their identifying characteristics, and their benefits to agriculture. Azotobacter promotes plant growth by fixing atmospheric nitrogen and producing plant hormones. It also functions as a biocontrol agent by suppressing plant pathogens. The document outlines Azotobacter's mode of action in plants and provides examples of increased crop yields and quality from its use as an inoculant. It also discusses the maintenance, selection, and mass production methods for Azotobacter cultures.
The document discusses various types of interactions between microorganisms including mutualism, commensalism, parasitism, predation, competition, and synergism. Specific examples are provided for each type of interaction such as lichens exhibiting mutualism between fungi and cyanobacteria. Both beneficial and harmful relationships between microbes and other organisms like plants, animals, and humans are explored.
The document discusses different types of biofertilizers and their production. It describes biofertilizers as microbial inoculants that establish symbiotic relationships with plants to enrich soil nutrients and promote crop growth. Major biofertilizers include rhizobia, azotobacter, algae, and phosphate-solubilizing bacteria. Rhizobium and cyanobacteria (blue-green algae) are discussed in detail, outlining their role in nitrogen fixation and methods for mass production, including trough, pit and field methods.
Microbiology is the study of microscopic organisms. There are several branches of microbiology including bacteriology, mycology, and virology. Microbes are found in diverse habitats and have relationships with other organisms. Important bacterial genera include Escherichia, Lactobacillus, Streptococcus, and Clostridium. Viruses can cause diseases like hepatitis, smallpox, and the common cold. Fungi include yeasts and molds and are found worldwide in various environments. Yeasts are used to produce alcoholic beverages and leaven baked goods.
This document summarizes bacterial blight, a common rice disease caused by the bacterium Xanthomonas oryzae. It first appears as water-soaked leaf spots and streaks, which can coalesce and spread to grains. The disease spreads via seeds, infected plant debris, and rain splash. Management strategies include seed treatment, multiple fungicide applications, chlorinated irrigation water, burning debris, isolation of nurseries, and growing resistant varieties.
DEFINITION
RHIZOSPHERE EFFECT
MICROORGANISMS FOUND IN RHIZOSPHERE
FACTORS INFLUENCING THEIR GROWTH AND ACTIVITIES
POSITIVE EFFECT OF RHIZOSPHERIC MICROORGANISMS ON PLANTS
NEGATIVE EFFECT OF RHIZOSPHERIC MICROORGANISMS ON PLANTS
Halophiles (Introduction, Adaptations, Applications)Jamil Ahmad
Introduction
Halophiles are organisms that thrive in high salt concentrations.
They are a type of extremophile organisms. The name comes from the Greek word for "salt-loving".
While most halophiles are classified into the Archaea domain, there are also bacterial halophiles and some eukaryota, such as the alga Dunaliella salina or fungus Wallemia ichthyophaga
Satellite viruses are sub-viral agents that depend on a helper virus for replication. The first reported satellite virus was Tobacco necrosis satellite virus. Satellite viruses contain nucleic acids enclosed in a protein coat and lack genes for replication. Satellite genomes can be single-stranded RNA, DNA, or circular RNA.
Satellite RNAs are small, linear or circular RNA strands found in certain multicomponent virus particles. They do not encode their own coat protein and depend on a helper virus for replication and encapsidation.
Viroids were discovered in 1971 and are small, circular, naked RNA molecules that replicate independently using host polymerases. Well-studied viroids include potato spindle tuber viroid and av
This document discusses several fungal plant diseases. It begins by covering wheat rust, describing the three types (leaf rust, stem rust, and stripe rust), their symptoms, and control methods. It also discusses cedar apple rust in apples, coffee leaf rust, and various root rots caused by fungi in different plants. Verticillium wilt and oak wilt are described. Finally, it discusses loose smut of barley and maize smut, detailing their symptoms, life cycles, and prevention strategies.
Interactions between microorganisms and plantsMSC yusur alani
Microorganisms interact with plants in various ways. Some interactions are mutualistic, such as nitrogen-fixing bacteria that form symbiotic relationships with legumes and cyanobacteria that provide nutrients to plants like Azolla. Mycorrhizal fungi also have mutualistic relationships with plant roots, helping uptake nutrients and providing other benefits. However, some microbes harm plants as pathogens, through production of diseases. The document also discusses how plant growth-promoting bacteria can stimulate plant growth through hormone production or nutrient transformations in the rhizosphere. Microorganisms additionally play roles in biocontrol of agricultural pests and pathogens.
This document discusses plant growth promoting rhizobacteria (PGPR) and their ability to solubilize inorganic phosphate. Some key points:
- PGPR are bacteria that live in the rhizosphere and provide benefits to plants. An important function is solubilizing insoluble phosphate minerals making phosphorus available for plant uptake.
- Common insoluble phosphates include tricalcium phosphate, dicalcium phosphate, and hydroxyapatite. Bacteria secrete organic acids like lactic acid and acetic acid to solubilize these minerals.
- Successful phosphate solubilizing bacteria include species from Bacillus, Pseudomonas, and Rhizobium genera. Screening methods involve checking for clearing zones
The document discusses the rhizosphere and phyllosphere, which are the regions of soil and plant surfaces influenced by microorganisms. The rhizosphere refers to the region of soil directly influenced by root secretions and microbes. It includes the inner and outer rhizosphere zones. Microbes in the rhizosphere play important roles in plant nutrition, growth promotion, and disease suppression. The phyllosphere refers to the interface between leaves and air, and is inhabited by bacteria, yeast and fungi that can benefit plants through nutrient management, disease control and stress tolerance.
he rhizosphere is the narrow region of soil or substrate that is directly influenced by root secretions and associated soil microorganisms known as the root microbiome.
The phyllosphere is a term used in microbiology to refer to the total above-ground portions of plants as habitat for microorganisms.
This document discusses the microbial flora of soil. It defines soil microflora as microorganisms that contribute to the biological properties of soil, including bacteria, fungi, algae, and protozoa. The five major groups interact and form a soil food web, with bacteria and fungi serving important roles in decomposition. Physical and chemical factors like pH and nutrients influence the growth and activity of soil microflora. The microflora are then classified and examples are given of bacteria, fungi, algae, protozoa, and their functions in the soil ecosystem.
This document discusses plant growth-promoting microbes (PGPM) which are soil microorganisms that can positively influence plant growth. PGPM are divided into two main groups - plant growth-promoting rhizobacteria (PGPR) and plant growth-promoting fungi (PGPF). PGPR and PGPF help plants grow through mechanisms like nitrogen fixation, phosphate solubilization, siderophore production, phytohormone synthesis, and protecting plants from pathogens. They play important roles in agricultural sustainability by improving soil fertility, supplying nutrients, suppressing diseases, and assisting in environmental remediation.
Soil microbiology is the study of microorganisms in soil such as bacteria, actinomycetes, fungi, algae and protozoa. These microorganisms are important because they affect soil structure and fertility through organic matter decomposition, nutrient transformations, and symbiotic relationships with plants. The four major groups of microbes found in soil are bacteria, actinomycetes, fungi, and algae, each playing an important role in soil health and plant growth.
This document provides an overview of plant growth promoting rhizobacteria (PGPR). It discusses that PGPR are a group of soil bacteria that colonize plant roots and enhance plant growth directly or indirectly. Direct mechanisms include biological nitrogen fixation, phosphate solubilization, phytohormone production, siderophore production, and antibiotic production. Indirect mechanisms include inducing systemic resistance in plants, production of lytic enzymes, and stress tolerance effects. The document reviews the specific mechanisms of several important PGPR functions and commercially available PGPR products.
soil characteristics influencing growth and activity of microfloraIGKV
This document discusses soil characteristics that influence the growth and activity of microflora in soil. It outlines several key factors: soil moisture and temperature are essential for microbial life and activity; most bacteria, algae and protozoa prefer neutral-slightly alkaline pH between 4.5-8.0; aerobic microbes require oxygen while anaerobic microbes do not; cultural practices like tilling and crop rotation alter microbial populations; root exudates and soil organic matter provide nutrients and food for microbes; microbial interactions can be symbiotic or antagonistic; and the physical, chemical and nutrient properties of different soil types impact microbial communities.
Soil organic matter has long been recognized as one of the most important components in maintaining soil fertility, soil quality, and agricultural sustainability. The soil zone strongly influenced by plant roots, the rhizosphere, plays an important role in regulating soil organic matter decomposition and nutrient cycling. Processes that are largely controlled or directly influenced by roots are often referred to as rhizosphere processes. These processes may include exudation of soluble compounds, water uptake, nutrient mobilization by roots and microorganisms, rhizosphere-mediated soil organic matter decomposition, and the subsequent release of CO2 through respiration. Rhizosphere processes are major gateways for nutrients and water. At the global scale, rhizosphere processes utilize approximately 50% of the energy fixed by photosynthesis in terrestrial ecosystems, contribute roughly 50% of the total CO2 emitted from terrestrial ecosystems, and mediate virtually all aspects of nutrient cycling. Therefore, plant roots and their rhizosphere interactions are at the center of many ecosystem processes. However, the linkage between rhizosphere processes and soil organic matter decomposition is not well understood. Because of the lack of appropriate methods, rates of soil organic matter decomposition are commonly assessed by incubating soil samples in the absence of vegetation and live roots with an implicit assumption that rhizosphere processes have little impact on the results. Our recent studies have overwhelmingly proved that this implicit assumption is often invalid, because the rate of soil organic matter decomposition can be accelerated by as much as 380% or inhibited by as much as 50% by the presence of live roots. The rhizosphere effect on soil organic matter decomposition is often large in magnitude and significant in mediating plant-soil interactions.
Mycorrhiza Biofertilizer is also known as VAM (Myco = Fungal + rrhiza = roots) adheres to plants rhizoids leading to development of hyphae. Hyphae boost development and spreading of white root in to soil leading to significant increase in rhizosphere. These hyphae further penetrate and form arbuscules within the root cortical. VAM fungi form a special symbiotic relationship with roots of plant that can enhance growth and survivability of colonized plants. Mycorrhiza Biofertilizer is very useful in organic farming as well as normal commercial farming
This document discusses Azotobacter, a genus of nitrogen-fixing bacteria that can be used as a biofertilizer. It describes the key species of Azotobacter, their identifying characteristics, and their benefits to agriculture. Azotobacter promotes plant growth by fixing atmospheric nitrogen and producing plant hormones. It also functions as a biocontrol agent by suppressing plant pathogens. The document outlines Azotobacter's mode of action in plants and provides examples of increased crop yields and quality from its use as an inoculant. It also discusses the maintenance, selection, and mass production methods for Azotobacter cultures.
The document discusses various types of interactions between microorganisms including mutualism, commensalism, parasitism, predation, competition, and synergism. Specific examples are provided for each type of interaction such as lichens exhibiting mutualism between fungi and cyanobacteria. Both beneficial and harmful relationships between microbes and other organisms like plants, animals, and humans are explored.
The document discusses different types of biofertilizers and their production. It describes biofertilizers as microbial inoculants that establish symbiotic relationships with plants to enrich soil nutrients and promote crop growth. Major biofertilizers include rhizobia, azotobacter, algae, and phosphate-solubilizing bacteria. Rhizobium and cyanobacteria (blue-green algae) are discussed in detail, outlining their role in nitrogen fixation and methods for mass production, including trough, pit and field methods.
Microbiology is the study of microscopic organisms. There are several branches of microbiology including bacteriology, mycology, and virology. Microbes are found in diverse habitats and have relationships with other organisms. Important bacterial genera include Escherichia, Lactobacillus, Streptococcus, and Clostridium. Viruses can cause diseases like hepatitis, smallpox, and the common cold. Fungi include yeasts and molds and are found worldwide in various environments. Yeasts are used to produce alcoholic beverages and leaven baked goods.
This document summarizes bacterial blight, a common rice disease caused by the bacterium Xanthomonas oryzae. It first appears as water-soaked leaf spots and streaks, which can coalesce and spread to grains. The disease spreads via seeds, infected plant debris, and rain splash. Management strategies include seed treatment, multiple fungicide applications, chlorinated irrigation water, burning debris, isolation of nurseries, and growing resistant varieties.
DEFINITION
RHIZOSPHERE EFFECT
MICROORGANISMS FOUND IN RHIZOSPHERE
FACTORS INFLUENCING THEIR GROWTH AND ACTIVITIES
POSITIVE EFFECT OF RHIZOSPHERIC MICROORGANISMS ON PLANTS
NEGATIVE EFFECT OF RHIZOSPHERIC MICROORGANISMS ON PLANTS
Halophiles (Introduction, Adaptations, Applications)Jamil Ahmad
Introduction
Halophiles are organisms that thrive in high salt concentrations.
They are a type of extremophile organisms. The name comes from the Greek word for "salt-loving".
While most halophiles are classified into the Archaea domain, there are also bacterial halophiles and some eukaryota, such as the alga Dunaliella salina or fungus Wallemia ichthyophaga
Satellite viruses are sub-viral agents that depend on a helper virus for replication. The first reported satellite virus was Tobacco necrosis satellite virus. Satellite viruses contain nucleic acids enclosed in a protein coat and lack genes for replication. Satellite genomes can be single-stranded RNA, DNA, or circular RNA.
Satellite RNAs are small, linear or circular RNA strands found in certain multicomponent virus particles. They do not encode their own coat protein and depend on a helper virus for replication and encapsidation.
Viroids were discovered in 1971 and are small, circular, naked RNA molecules that replicate independently using host polymerases. Well-studied viroids include potato spindle tuber viroid and av
This document discusses several fungal plant diseases. It begins by covering wheat rust, describing the three types (leaf rust, stem rust, and stripe rust), their symptoms, and control methods. It also discusses cedar apple rust in apples, coffee leaf rust, and various root rots caused by fungi in different plants. Verticillium wilt and oak wilt are described. Finally, it discusses loose smut of barley and maize smut, detailing their symptoms, life cycles, and prevention strategies.
Interactions between microorganisms and plantsMSC yusur alani
Microorganisms interact with plants in various ways. Some interactions are mutualistic, such as nitrogen-fixing bacteria that form symbiotic relationships with legumes and cyanobacteria that provide nutrients to plants like Azolla. Mycorrhizal fungi also have mutualistic relationships with plant roots, helping uptake nutrients and providing other benefits. However, some microbes harm plants as pathogens, through production of diseases. The document also discusses how plant growth-promoting bacteria can stimulate plant growth through hormone production or nutrient transformations in the rhizosphere. Microorganisms additionally play roles in biocontrol of agricultural pests and pathogens.
Biofertilizers contain living microorganisms that colonize plant roots and soil to promote plant growth. They do this through natural processes like nitrogen fixation, phosphorus solubilization, and stimulating plant growth with substances. There are several types of biofertilizers including bacterial (Rhizobium, Azotobacter), fungal (VAM), and algal biofertilizers. They help increase soil fertility and nutrient supply to plants, reducing the need for chemical fertilizers. Mass production of biofertilizers involves growing the microorganisms in large quantities and mixing them with a carrier for storage and application to soil and plants.
Roots release chemicals called rhizodeposits into the surrounding soil, known as the rhizosphere. This biologically active area is populated by microorganisms that interact with and benefit from root exudates. Rhizodeposits include mucilage, root exudates, and sloughed-off root cells and layers. They support soil microbial life and their composition varies by plant type, climate, soil properties, and nutrient availability. Root exudates communicate with both microbes and other plant roots, selectively fostering beneficial microbes while inhibiting competing plants or pathogens through allelopathy. The rhizosphere is crucial for nutrient acquisition and microbial symbioses that are important for plant growth and health.
This document discusses soil microorganisms and their classification. It notes that soil microorganisms play important roles in soil nitrogen utilization, organic matter decomposition, nutrient transformations, and soil formation. The microorganisms are classified as bacteria, actinomycetes, fungi, and algae. Bacteria are the most abundant and play key roles such as organic matter decomposition and enzymatic reactions. Actinomycetes are filamentous bacteria that also decompose organic matter. Fungi possess filamentous mycelium and are important for organic matter decomposition. Algae are photoautotrophs that fix carbon and are present in soils.
Mechanism of disease control by endophytesPooja Bhatt
The document discusses alternative methods for pest management to address problems with chemical pesticides such as development of resistance and environmental contamination. It suggests that biological control using endophytic microorganisms is a promising alternative as endophytes have antagonistic properties against plant pathogens. Endophytes can inhibit pathogens through direct mechanisms such as hyperparasitism, competition, antibiosis, and lytic enzyme production or indirect induction of host plant resistance. Case studies provide examples of endophytes inhibiting fungal plant pathogens through siderophore production, parasitic growth, and antibiotic compounds.
This document discusses mycorrhizae, which are symbiotic associations between fungi and plant roots. It describes the different types of mycorrhizae including endomycorrhizae, ectomycorrhizae, and ectendomycorrhizae. Applications of mycorrhizae include increasing nutrient uptake, plant diversity, and resistance to diseases and drought. The document also discusses methods for isolating and mass producing mycorrhizal fungi for use as biofertilizers to improve soil health and crop yields.
The document discusses mycorrhizae and their potential as biocontrol agents. It describes the evolution and discovery of mycorrhizal fungi. There are different types of mycorrhizal associations based on the relationship with host plants. The document discusses the distribution of mycorrhizal fungi among host and non-host plants. It explains how vesicular arbuscular mycorrhizae (VAM) can act as biocontrol agents by suppressing soil-borne pathogens through various mechanisms like physiological alterations in host plants, competition for space and nutrients, and changes in root exudates. The mycorrhizal intensity was found to be higher in healthy plants compared to diseased plants in fields studied
Lichens=(Algae+Fungi) Symbiotic Association (Phycobiont+ Mycobiont), Idealistic marriage, Pioneers species of Xerosere succession Shows Dual Nature, Trinity=(One Algae+Two Fungi), Natural farmers, it melt stone convert stone to soil particles
Plants require essential nutrients for their growth and development that are mainly acquired from soil by their roots. Nutrient stress is an environmental condition that can seriously affect the production and quality of crop produce. Biofertilizers are the organisms (Bacteria, fungi, cyanobacteria, etc.) that enrich the nutrient quality of soil. Plants have a number of beneficial relationship with such organisms. Among these AM-Fungi are ubiquitous and form a mutuality relationship with roots of most plant species.
1. Microbes play a central role in nitrogen availability and life on Earth. Certain bacteria can fix nitrogen from the atmosphere in forms usable by plants through free-living or symbiotic relationships.
2. The document discusses nitrogen fixation by bacteria, which make nitrogen available to support plant growth. It also describes symbiotic nitrogen fixation between rhizobia bacteria and legume plants in root nodules.
3. Organic farming aims to feed the soil by promoting nutrient cycling without synthetic pesticides or fertilizers. It uses techniques like crop rotation, cover crops, composting, and biological pest control.
Fungi play several important ecological roles. As decomposers, they break down dead organic matter and release nutrients back into the soil, recycling them for other organisms. As mycorrhizal partners with plants, fungi help plants absorb water and minerals in exchange for carbohydrates. Lichens, which consist of fungi and algae, are pioneers in plant succession and weather rocks into soil. Some fungi also help bioremediate polluted environments and produce antibiotics. However, some fungi can also cause diseases or destroy plants, wood, food, and other materials.
B.sc. agri sem ii agricultural microbiology unit 2 soil microorganismsRai University
The document discusses the various microorganisms found in soil, including bacteria, actinomycetes, fungi, algae, protozoa, and higher animal forms. It describes the different groups of bacteria and microorganisms, their roles in soil processes like decomposition and nutrient cycling, and how they vary depending on environmental conditions. Methods for enumerating and classifying soil microorganisms are also summarized.
1. Mycorrhiza plays an important role to establish forest in unfavourable location, barren land, waste lands etc.
2. Trees with facultative endomycorrhiza act as first invader in waste lands as pioneer in plant succession.
3. The application of mycorrhizal fungi in forest bed enhances the formation of mycorrhizal association that prevents the entry of fungal root pathogens. This method is very much effective in the root of Pinus clausa against Phytophthora cinnamoni infection.
4. Mycorrhiza mixed nitrogenous compounds such as nitrate; ammonia etc. is available to the plants. Thus it helps in plant growth, especially in acid soil.
Mycorrhizal fungi form mutualistic relationships with the roots of most plant species. They help plants absorb water and mineral nutrients from soil in exchange for carbohydrates. There are two main types of mycorrhizal associations - ectomycorrhizas, which cover tree roots with a sheath and branching structures, and endomycorrhizas, whose thread-like structures penetrate root cells. Mycorrhizal fungi play an important ecological role in nutrient cycling, plant community development, and soil health.
(i) Biofertilizers help crop plants uptake nutrients through interactions in the rhizosphere. They contain living microorganisms that promote nutrient availability in a plant-accessible form.
(ii) Organic agriculture promotes agroecosystem health through practices preferring natural inputs over synthetic ones. It emphasizes biodiversity, biological cycles and soil activity.
(iii) Overuse of chemical fertilizers pollutes soil and water, destroys microorganisms, and reduces soil fertility. Biofertilizers are more sustainable, efficient and affordable options for farmers.
This document discusses different forms of plant-microbe interactions. It provides examples of mutualistic relationships between plants and microbes, including decomposition, mycorrhizal associations, and nitrogen fixation. Decomposition and nitrogen fixation are carried out by various bacteria and fungi. Mycorrhizal associations involve fungi colonizing plant roots and increasing nutrient and water uptake for the plant. The rhizosphere, or area of soil surrounding plant roots, contains many microbes due to root exudates and supports various interactions between plants and beneficial, neutral, or pathogenic microbes.
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1. Agnihotra is an ancient Vedic practice of performing small fire sacrifices at sunrise and sunset using dried rice grains, ghee, and mantras to purify the atmosphere.
2. It was discovered by Rishi Kanva and mentioned in Vedic texts as a way to remove pollution and contamination through the transforming power of fire.
3. Regular Agnihotra practice is believed to positively impact microorganisms, plants, humans, animals and the environment by healing and harmonizing the atmosphere.
This document discusses bioaerosols and droplet nuclei. It defines bioaerosols as small airborne biological material, either attached to particulate matter or not, including microorganisms, their byproducts, and fragments. Bioaerosols can be viable, such as living bacteria and viruses, or non-viable, like toxins and proteins. Droplet nuclei are small aerosols formed from the evaporation of respiratory droplets smaller than 5 micrometers that can transmit diseases through airborne transmission. Common sources of bioaerosols include outdoor environmental sources, indoor materials and occupants, and human activities like coughing or talking. Exposure to some bioaerosols is associated with respiratory illnesses, but others may benefit immune
Soil is a habitat for many microorganisms that are involved in important biogeochemical cycles like carbon, nitrogen, and sulphur. Microbes interact with each other in the soil environment and carry out processes in these elemental cycles that are essential for life.
Soil microbiology is the study of microorganisms in soil. The document provides information about soil microbiology from the Department of Microbiology at MUSKAN. It thanks the reader for their time.
Aeromicrobiology is the study of microbes suspended in air, known as bioaerosols. These microbes can travel long distances via wind and precipitation, potentially spreading disease widely. Bioaerosols originate from terrestrial and aquatic environments and are launched into the air by turbulence, transported by winds, and deposited through mechanisms like gravity or rain. Microbes in air must adapt to stresses like desiccation and UV exposure.
BIOMOLECULE PRESENTATION BY MUSKAN.pptxMuskan Ashi
Enzymes are biological macromolecules that act as catalysts to speed up biochemical reactions without undergoing any changes themselves. They greatly increase both the rate and specificity of metabolic reactions. Enzymes are usually proteins that catalyze reactions under mild conditions like normal body temperature and pH. They are highly specific to their substrates and can have their activity regulated by substrate or allosteric effector concentrations. Factors like temperature, pH, enzyme and substrate concentrations affect the rate of enzymatic reactions. The mechanisms of enzyme action include the lock-and-key and induced fit models where the flexible active site binds and may change shape to accommodate different substrates.
This document discusses viruses, including their structure, classification, and discovery. It notes that viruses consist of nucleic acid and a protein coat, and are able to multiply only within host cells. It describes some of the early discoveries of viruses in the late 19th century. It also summarizes different classification systems for viruses, including those based on nucleic acid type, structure, and genome, such as the Baltimore classification system. The document provides an overview of viruses with relevant details on their composition, life cycles, and taxonomic organization.
A review of the growth of the Israel Genealogy Research Association Database Collection for the last 12 months. Our collection is now passed the 3 million mark and still growing. See which archives have contributed the most. See the different types of records we have, and which years have had records added. You can also see what we have for the future.
How to Manage Your Lost Opportunities in Odoo 17 CRMCeline George
Odoo 17 CRM allows us to track why we lose sales opportunities with "Lost Reasons." This helps analyze our sales process and identify areas for improvement. Here's how to configure lost reasons in Odoo 17 CRM
A Strategic Approach: GenAI in EducationPeter Windle
Artificial Intelligence (AI) technologies such as Generative AI, Image Generators and Large Language Models have had a dramatic impact on teaching, learning and assessment over the past 18 months. The most immediate threat AI posed was to Academic Integrity with Higher Education Institutes (HEIs) focusing their efforts on combating the use of GenAI in assessment. Guidelines were developed for staff and students, policies put in place too. Innovative educators have forged paths in the use of Generative AI for teaching, learning and assessments leading to pockets of transformation springing up across HEIs, often with little or no top-down guidance, support or direction.
This Gasta posits a strategic approach to integrating AI into HEIs to prepare staff, students and the curriculum for an evolving world and workplace. We will highlight the advantages of working with these technologies beyond the realm of teaching, learning and assessment by considering prompt engineering skills, industry impact, curriculum changes, and the need for staff upskilling. In contrast, not engaging strategically with Generative AI poses risks, including falling behind peers, missed opportunities and failing to ensure our graduates remain employable. The rapid evolution of AI technologies necessitates a proactive and strategic approach if we are to remain relevant.
Strategies for Effective Upskilling is a presentation by Chinwendu Peace in a Your Skill Boost Masterclass organisation by the Excellence Foundation for South Sudan on 08th and 09th June 2024 from 1 PM to 3 PM on each day.
ISO/IEC 27001, ISO/IEC 42001, and GDPR: Best Practices for Implementation and...PECB
Denis is a dynamic and results-driven Chief Information Officer (CIO) with a distinguished career spanning information systems analysis and technical project management. With a proven track record of spearheading the design and delivery of cutting-edge Information Management solutions, he has consistently elevated business operations, streamlined reporting functions, and maximized process efficiency.
Certified as an ISO/IEC 27001: Information Security Management Systems (ISMS) Lead Implementer, Data Protection Officer, and Cyber Risks Analyst, Denis brings a heightened focus on data security, privacy, and cyber resilience to every endeavor.
His expertise extends across a diverse spectrum of reporting, database, and web development applications, underpinned by an exceptional grasp of data storage and virtualization technologies. His proficiency in application testing, database administration, and data cleansing ensures seamless execution of complex projects.
What sets Denis apart is his comprehensive understanding of Business and Systems Analysis technologies, honed through involvement in all phases of the Software Development Lifecycle (SDLC). From meticulous requirements gathering to precise analysis, innovative design, rigorous development, thorough testing, and successful implementation, he has consistently delivered exceptional results.
Throughout his career, he has taken on multifaceted roles, from leading technical project management teams to owning solutions that drive operational excellence. His conscientious and proactive approach is unwavering, whether he is working independently or collaboratively within a team. His ability to connect with colleagues on a personal level underscores his commitment to fostering a harmonious and productive workplace environment.
Date: May 29, 2024
Tags: Information Security, ISO/IEC 27001, ISO/IEC 42001, Artificial Intelligence, GDPR
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How to Add Chatter in the odoo 17 ERP ModuleCeline George
In Odoo, the chatter is like a chat tool that helps you work together on records. You can leave notes and track things, making it easier to talk with your team and partners. Inside chatter, all communication history, activity, and changes will be displayed.
বাংলাদেশের অর্থনৈতিক সমীক্ষা ২০২৪ [Bangladesh Economic Review 2024 Bangla.pdf] কম্পিউটার , ট্যাব ও স্মার্ট ফোন ভার্সন সহ সম্পূর্ণ বাংলা ই-বুক বা pdf বই " সুচিপত্র ...বুকমার্ক মেনু 🔖 ও হাইপার লিংক মেনু 📝👆 যুক্ত ..
আমাদের সবার জন্য খুব খুব গুরুত্বপূর্ণ একটি বই ..বিসিএস, ব্যাংক, ইউনিভার্সিটি ভর্তি ও যে কোন প্রতিযোগিতা মূলক পরীক্ষার জন্য এর খুব ইম্পরট্যান্ট একটি বিষয় ...তাছাড়া বাংলাদেশের সাম্প্রতিক যে কোন ডাটা বা তথ্য এই বইতে পাবেন ...
তাই একজন নাগরিক হিসাবে এই তথ্য গুলো আপনার জানা প্রয়োজন ...।
বিসিএস ও ব্যাংক এর লিখিত পরীক্ষা ...+এছাড়া মাধ্যমিক ও উচ্চমাধ্যমিকের স্টুডেন্টদের জন্য অনেক কাজে আসবে ...
Assessment and Planning in Educational technology.pptxKavitha Krishnan
In an education system, it is understood that assessment is only for the students, but on the other hand, the Assessment of teachers is also an important aspect of the education system that ensures teachers are providing high-quality instruction to students. The assessment process can be used to provide feedback and support for professional development, to inform decisions about teacher retention or promotion, or to evaluate teacher effectiveness for accountability purposes.
Thinking of getting a dog? Be aware that breeds like Pit Bulls, Rottweilers, and German Shepherds can be loyal and dangerous. Proper training and socialization are crucial to preventing aggressive behaviors. Ensure safety by understanding their needs and always supervising interactions. Stay safe, and enjoy your furry friends!
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Macroeconomics- Movie Location
This will be used as part of your Personal Professional Portfolio once graded.
Objective:
Prepare a presentation or a paper using research, basic comparative analysis, data organization and application of economic information. You will make an informed assessment of an economic climate outside of the United States to accomplish an entertainment industry objective.
Physiology and chemistry of skin and pigmentation, hairs, scalp, lips and nail, Cleansing cream, Lotions, Face powders, Face packs, Lipsticks, Bath products, soaps and baby product,
Preparation and standardization of the following : Tonic, Bleaches, Dentifrices and Mouth washes & Tooth Pastes, Cosmetics for Nails.
10. Microorganisms found in Rhizosphere (Rhizosphere microbiome)
The microbial population in the rhizosphere consists of different groups of microorganisms like bacteria, fungi, parasites,
viruses, and algae.
The microbial population in the rhizosphere is known as the rhizosphere microbiome and the microbial population in such an
area much higher than the bulk soil.
In the rhizosphere, there is a microbial population distinct from the rest of the soil.
Bacteria in the rhizosphere are larger and have higher proportions of Gram-negative and denitrifying bacteria than those in the
bulk soil.
Rhizosphere fungal populations, abundant in both pathogenic and mycorrhizal species, can be 10 to 20 times higher than those
in the non-rhizosphere.
Protozoa and other microfauna also thrive in the rhizosphere because that is where food is most plentiful.
The type and population of microorganisms in the rhizosphere are highly influenced by the type of plant grown on the soil.
Microbes in the bulk soil often experience long periods of nutrient deprivation; they have different survival strategies in
dealing with starvation and stress.
The rhizosphere bacterial community is recruited from the main reservoir of microorganisms present in the soil.
Next to the recruitment of specific soil microbes into the rhizosphere microbiome, plant roots also influence specific functions
of the microbiome.
Some of the examples of microorganisms found in the rhizosphere region include Bacillus, Arthrobacter, Pseudomonas,
Agrobacterium, Alcaligenes, Clostridium, Flavobacterium, Corynebacterium, Micrococcus, Xanthomonas, Amanita,
Tricholoma, Torrendia, Descomyces, Thelephora, Verticillium, Phytophthora, Rhizoctonia, Micromonospora,
Thermoactinomycetes, Amycolaptosis, Actinomadura, etc.
11. Positive effect of Rhizospheric microorganisms on Plants
Rhizospheric microorganisms play an important role in the ecological fitness of the plant and the
soil.
Important microbial processes like plant protection, growth promotion, production of antibiotics,
geochemical cycling, and plant colonization take place in the rhizosphere.
Rhizospheric microorganisms increase the supply of mineral nutrients from the soil to the plant.
Another group of microorganisms in the rhizosphere stimulate plant growth indirectly by
preventing the growth or activity of plant pathogens.
These microorganisms are responsible for direct growth promotion by the production of
phytohormones.
Plant growth-promoting rhizobacteria act as biofertilizers by enhancing phytochrome production,
phosphate solubilization, and siderophore production.
The capacity of rhizospheric organisms to synthesize anti-fungal metabolites such as antibiotics,
fungal cell wall-lysing enzymes, or hydrogen cyanide suppresses the growth of fungal pathogens.
12. Negative effect of Rhizospheric microorganisms on Plants
One of the most important negative effects of rhizosphere microorganisms is a competition where
the microorganisms compete with the plants for water, nutrient, and space.
Some of the members of the rhizosphere microbiome might act as plant pathogens, resulting in
different forms of plant diseases.
Competition between the microorganisms in the microbial community in the rhizosphere can even
result in the loss of beneficial microorganisms.
13.
14. Estimation
Microorganisms are utilized in agriculture for various purposes; as important components of
organic amendments and composts, as inoculants for biological nitrogen fixation, phosphorous
solubilization and indole acetic acid (IAA), to improve crop quality and yields.
One of the most common strategies to increase agricultural production is through the
improvement of soil fertility. Nitrogen (N) and phosphorus (P) are the two most limiting nutrients
in soil. Indole acetic acid is an essential natural growth promoter that extensively affects plant
growth and development.
Reports have indicated the ability of many bacterial microorganisms to produce phytohormones
that can enhance the plant root contact surface with soil and subsequently the increase of nutrient
uptake via root elongation. Due to this ability, microorganism inoculants can be used as a
substitute for chemical fertilizers in partially fertile soils and/or at least as a supplement for
chemical fertilizers in infertile soils.
Therefore, the estimation was undertaken to screen the rhizosphere, rhizoplane and phyllosphere
bacteria and fungi isolated from rice growing regions of Kenya for their physiological
characteristics, including P-solubilization, N-fixation and IAA production.
15. .
Most of the bacterial isolates from the rhizosphere, rhizoplane as well as the phyllosphere had
ARA though at low levels. Bacterial isolates from the rhizosphere and rhizoplane were found to
be efficient in P solubilization whereas the fungal isolates were mostly non-solubilizers. Although
the percentage of IAA production by the tested isolates was not high, the bacterial isolates
performed better than the fungal isolates.
The results therefore suggest that these microorganisms have the potential to be utilized as
microbial inoculants to replace chemical fertilizers for sustainable rice cultivation in the Kenyan
rice growing regions. Thus the future of biofertilizers based on nitrogen fixing, phosphate
solubilizing and IAA producing bacteria and to some extent fungi seems very promising.
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34. Introduction
An actinomycete (filamentous, branching, gram-positive bacteria) that forms nodules and fixes
dinitrogen in a fashion analogous to that used by rhizobia.
The dominant actinorhizal genus is Frankia, occurring on roots of 8 plant families, encompassing
24 genera and some 230 species of dicotyledons.
Prominent actinorhizal plant families and genera are: Betulaceae, on 47 Alnus species;
Casuarinaceae, on 16 spp. of Casuarina, and 54 spp. of Allocasuarina; Myricaceae, 28 spp. of
Myrica, and Rhamnaceae, 31 spp. of Ceanothus.
Frankia is a genus of soil actinomycetes famous for its ability to form N2-fixing root nodule
symbioses with actinorhizal plants.
Although Frankia strains display a high diversity in terms of ecological niches in soil, current
knowledge about Frankia is dominated by its life as an endophyte in root nodules. Increased use
of molecular methods has refined and expanded insights into endophyte-host specificities and
Frankia phylogeny.
Frankia is a genus of soil actinomycetes in the family Frankiaceae occurring also in symbiosis
with certain angiosperms. The actinomycete Frankia is defined as the N2-fixing microsymbiont of
actinorhizal plants.
35. .
Actinorhizal symbioses as examplified from Patagonia,
Argentina. a Discaria chacaye shrubs growing along a
river in northwestern Patagonia; b Discaria trinervis
shoot with mature fruits; c Discaria trinervis multilobed
nodule; d Discaria trinervis mature nodule in
longitudinal section showing characteristic central
vascular tissue (vt), apical meristem (m) and infected
cells full of vesicle clusters stained blue ; e liquid
culture of Frankia strain DcI45 isolated from Discaria
chacaye root nodules; f Frankia strain DaI1 isolated
form Dicsaria articulata showing characteristic hyphae,
multilocular sporangia, spores and a spherical vesicle
(inset)
36. Overview
Actinorhizal plants are classified into four subclasses, eight families, and 25 genera comprising
more than220 species.
There are two main types of symbioses between nitrogen-fixing bacteria and vascular plants: one
between Rhizobium and leguminous plants, and the other between Frankia and actinorhizal
plants.
Actinorhizal plants comprise more than 220 species symbiotically associated with the
filamentous actinomycete Frankia
Gunnera, which establishes a symbiosis with cyanobacteria in a specialized stem structure,
represents a third type of nitrogen-fixing symbiosis. A fourth type is that which occurs between
cyanobacteria and cycads.
Frankia-actinorhizal plant and Rhizobium- legume symbioses have been known for many years to
benefit soil fertility.
A fourth type is that which occurs between cyanobacteria and cycads. Other diverse diazotrophs,
such as Azospirillum, Herbaspirillum, and Acetobacter, have been isolated and identified from the
rhizosphere or from roots of many other plants, generally grasses, but are not symbiotically
associated in root nodules.
37.
38. The bacteria
The microsymbiont of actinorhizal plants was first referred to as Frankia in 1888 by Brunchorst and was
later classified as an actinomycete after studies by Krebber in 1932 (Quispel 1990).
The genusnFrankia is comprised of gram-positive and gram- variable actinomycetes (Lechevalier and
Lechevalier 1990). The first cultured Frankia, isolated from Alnus root nodules was reported by Pommer
(1959), but unfortunately the culture was lost. In 1978, the first successful isolation of Frankia was reported
from Comptonia peregrina root nodules (Callaham and others 1978), beginning a new era in actinorhizal
symbiosis research (Quispel 1990).
Filaments, vesicles, and sporangia have the potential for being infective particles although they must
germinate and grow as new filaments to infect the root. Spores are probably a major means of Frankia
propagation in nature. It has been shown that Frankia cells are distributed through air by birds and also
accumulate in river and lake sediments. All three cell types can be found in the symbiotic state, although
there are some exceptions.
Cultivated Frankia cells behave as heterotrophic aerobic bacteria with doubling times of 15 h, com- pared
with 3 h for rhizobia.
Frankia comprises not only symbiotic bacteria but also free-living actinomycetes in the soil. Frankia cell
wall and cell envelope composition is distinct from that of other bacteria, in particular, because of the
presence of hopanoids in the multi- layer envelope of the vesicle (Harriot and others 1991). This lipid
envelope acts as a gas diffusion barrier to prevent high oxygen tension within vesicles, thereby permitting
nitrogenase expression and activity, both in culture and in symbiotic state.
39. .
It is worth noting that Frankia is not the only microorganism that can be isolated from actinorhizal
root nodules. For instance, a previously unrecognized actinomycete was isolated from root
nodules of Casuarina trees growing in Mexico. This microorganism appears to fix nitrogen, on the
basis of acetylene reduction assays, but does not develop vesicles or sporangia in culture, and
moreover, it is unable to reinfect its original host.
40. THE ACTINORHIZAL PLANT
All the actinorhizal plants are trees or shrubs, except for the genus Datisca, which is herbaceous.
Some species are very well adapted to flooded lands, warm arid and semiarid regions, and areas
of devastation (for example, rock slides).
Actinorhizal plants have numerous uses: soil restoration, fuel wood, production of wood and
derivatives, agroforestry, coastal restoration, and the prevention of desertification.
This tripartite symbiosis gives a high degree of autotrophy to these plant-microorganism
associations. Thus, actinorhizal plants are natural pioneers in succession on land, and they are
frequently the first species colonizing disturbed areas.
Nitrogen fixation by actinorhizal plants in nature seems to be of similar magnitude as that of the
legumes showing diurnal and seasonal variation with an estimated annual rate of 240—350 kg
ha−1 y−1.
Actinorhizal plants are perennial, so their contribution to N cycle through litter fall and soil
decomposition is ecologically relevant.
41. THE ROOT NODULE
Actinorhizal nodules resemble modified lateral roots, having a central vascular bundle. Because of its
indeterminate structure, the development of the symbiotic association is recapitulated longitudinally in a
mature nodule.
At the nodule tip is the uninfected apical meristem from which nodule parenchyma develops. Adjoining the
meristem in a basipetal direction is a region of uninfected cells followed by a region of recently infected
cells without vesicle differentiation, known as the infection zone.
The central nodule tissue, or fixation zone, contains two types of cells: mature infected cells with
differentiated vesicles, where nitrogen fixation takes place, and uninfected cells, which are probably
involved in assimilation of the fixed N and exchange of C.
The distribution of infected and uninfected cells in the fixation zone differ depending on the actinorhizal
plant genus. The different arrangements are attributed to differences in oxygen protection mechanisms. At
the nodule base, the senescent zone is present. Because actinorhizal nodules are perennial, they show
seasonal variations in the proportion of these zones.
42. Applications
Actinorhizal plants could be useful tools to develop a sustainable economy. If there is interest in
extending the ability to establish a nitrogen-fixing symbiosis to a plant species with economic
potential, efforts should concentrate first on close relatives of well known actinorhizal plants.
Beneficial role of the symbiotic state there may be greater pathogenic resistance induced after
Frankia nodulation.
More basic studies on the complex interactions between Frankia and the actinorhizal plants will
help us achieve a better understanding not only of symbiosis but also of plant growth regulation.