Bryophytes are amphibious primitive land plants that were the first to colonize land. They have a life cycle that alternates between a haploid gametophyte generation that produces gametes and a diploid sporophyte generation that produces spores. The plant body is the gametophyte, which is either a thallus or an erect foliose structure. Bryophytes reproduce sexually through antheridia and archegonia that produce sperm and eggs, requiring water for fertilization. The sporophyte develops from the fertilized egg and produces spores in capsules, which germinate to form new gametophytes.