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Sponges and Placozoans
Chapter 12
Phylum Porifera (sponges)
Diversity
- 18,644 species (Van Soest et al.
2012)
- 8,600 species (Hickman et al.
2017)
- 4 classes - Demospongiae,
Hexactinellida, Calcera,
Homoscleromorpha
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Where found?
- All aquatic, mostly marine, a few freshwater.
General characteristics
- No germ layers
- All sessile (attached to substrate and do not move).
- All but one are filter feeders (feed on particles suspended in water)
- Cellular level of organization (no tissues, organs, or organ systems)
Phylum Porifera (sponges)
General characteristics (cont.)
Skeleton in 2 parts
Rigid skeleton = spicules
Fibrous skeleton = spongin
Homoscleromorpha
Calcera
Demospongiae
Hexactinellida
Phylum Porifera (sponges)
Body Plan
Body a series of interconnected pores and canals.
- Movement of the flagella on the choanocyte draws water into the sponge.
- Water enters through pores known as dermal ostia.
- The collar of the choanocyte captures food particles.
- Water exits through the osculum (oscula = pl.)
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Body Plan (cont.)
- Cell types - 3 broad classes recognized (many subclass though)
- Choanocytes - generate water currents and capture food.
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Body Plan (cont.)
Cell types - 3 broad classes recognized (many subclass though)
- Archaeocytes - amoeboid cells; receive food from choanocytes
- Capable of differentiating into more specialized cells
- Sclerocytes - secrete spicules
- Spongocytes - secrete spongin
- Collencytes - secrete collagen
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Body Plan (cont.)
- Cell types - 3 broad classes recognized (many subclass though)
- Pinacocytes - form the outer covering of sponges.
- Some, myocytes, are capable of contracting; closes ostia.
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Body Plan (cont.)
- Extracellular materials
- Mesohyl - functions as connective tissue (but not a true tissue).
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Arrangement of canals and choanocytes ranges from simple to complex.
- Greater complexity allows for more food to be gathered and thus larger size.
Asconoid - simplest arrangements and therefore the smallest size.
Syconoid - intermediate arrangement and therefore capable of intermediate size.
Leuconoid - most complex arrangement and therefore capable of largest size.
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Phylum Porifera (sponges)
Homeostatic functions and structures
- Osmotic balance - none
- Excretion - diffusion
- Respiration - diffusion
- Temperature regulation - none
- Circulation - none (all processes by diffusion)
Phylum Porifera (sponges)
Nervous system
- Lacking along with sensory structures .
- Can respond to localized stimuli.
Phylum Porifera (sponges)
Regenerative and reproductive processes
- Great ability to regenerate and replace lost parts.
- Undergo somatic embryogenesis when fragmented.
Phylum Porifera (sponges)
Regenerative and reproductive processes
- Capable of asexual and sexual reproduction.
- Asexual - budding (external buds), gemmulation (internal buds)
- Gemmulation - common in freshwater sponges where freezing and
drought are problematic.
Phylum Porifera (sponges)
Regenerative and reproductive processes
- Capable of asexual and sexual reproduction.
- Sexual - most sponges are monecious; gametes from choanocytes.
- Most are viviparous and produce a ciliated larvae.
Phylum Placozoa (placozoans)
Diversity - 1 species (Trichoplax adhaerens)
Where found - marine
General characteristics
- Plate-like body; asymmetrical; diploblastic without tissues or organs
- Dorsal and ventral sides, but no head
- Dorsal has flagellated cells with bright spheres
- Ventral has columnar ciliated cells with glands.
Phylum Placozoa (placozoans)
General characteristics
- Fibrous contractile cells between dorsal and ventral
Feeding
- Position above food item; secrete enzymes; absorb
remains.
Reproduction
- Asexual known; sexual not known
Phylum Placozoa (placozoans)

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