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http://photography.nationalgeographic.com/staticfiles/NGS/Shared/StaticFiles/Photography/Images/Content/sea-star-pair-758303-sw.jpg
Echinoderm Classes
Asteroidea – Sea Stars:
 Multiple arms radiating from
central disk
 Tube feet on bottom
Riley Galton
Asteroidea – Sea Stars:
 Multiple arms radiating from
central disk
 Tube feet on bottom
Ophiuroidea – Brittle Stars:
 Distinct central disk with snake like arms
 Tube feet lack suckers
http://photography.nationalgeographic.com/staticfiles/
http://environment.nationalgeographic.com/staticfiles/
Echinoderm Classes
Echinoidea – Sea Urchins and Sand Dollars:
 No arms, but 5 rows of tube feet
 Have muscles that can move
 spines
Riley Galton
Echinoidea – Sea Urchins and Sand Dollars:
 No arms, but 5 rows of tube feet
 Have muscles that can move
 spines
http://environment.nationalgeographic.com/staticfiles/
www.emiliogrossi.com/images/Photo
graphyiles/
Crinoidea – Feather Stars
and Sea Lilies
 Flexible arms used in
suspension feeding
 Haven’t changed in 500 million
years
http://lh6.ggpht.com http://image44.webshots.com
Echinoderm Classes
Holothuroidea – Sea Cucumbers
 Reduced endoskeleton
 Five rows of tube feet
Riley Galton
Holothuroidea – Sea Cucumbers
 Reduced endoskeleton
 Five rows of tube feet
Concentricycloidea – Sea Daisies
 Only two known species
 Disk shaped, less than 1cm in diameter
http://blog.case.edu/case-news/2008/03/06/seacucumber.jpg
http://www.sfu.ca/~fankbone/v/xyloplax.jpg
Echinoderm Sensory
Structures
 Eyespots on the ends of ambulacra are
sensitive to light
 Pedicellariae (jaw like skeletal structures used to keep
organisms from settling on skin), tube feet, and spines
are sensitive to touch
http://www.ucmp.berkeley.edu/echinodermata/echinomm.html
Riley Galton
 Eyespots on the ends of ambulacra are
sensitive to light
 Pedicellariae (jaw like skeletal structures used to keep
organisms from settling on skin), tube feet, and spines
are sensitive to touch
http://www.ucmp.berkeley.edu/echinodermata/echinomm.html
Echinoderm Reproduction
Sexual:
• Eggs and sperm are released into the
water where they are fertilized
• Larvae are planktonic
• Internal fertilization has been observed
in certain species of sea stars, brittle
stars, and sea cucumbers
Asexual:
• Echinoderms can regenerate body
parts and organs
• Asexual reproduction can occur
(though not by will) if the echinoderm is
torn with enough of the body on each
part
http://www.oceaninn.com/guides/echino.htm
Riley Galton
http://www.biolbull.org/content/vol204/issue3/images/medium/cov
erfig.gif
Sexual:
• Eggs and sperm are released into the
water where they are fertilized
• Larvae are planktonic
• Internal fertilization has been observed
in certain species of sea stars, brittle
stars, and sea cucumbers
Asexual:
• Echinoderms can regenerate body
parts and organs
• Asexual reproduction can occur
(though not by will) if the echinoderm is
torn with enough of the body on each
part
http://www.oceaninn.com/guides/echino.htm
http://www.starfish.ch/Fotos/echinoderms
Body Cavity (Coelom)
 PRESENT!
 Coelom = A fluid-filled cavity between body wall and gut;
lined by mesoderm
 Two parts:
Perivisceral coelom – large, fluid-filled cavity where major
organs, digestive tube and sex organs, are suspended
Water Vascular System - network of hydraulic canals unique
to echinoderms that branches into extensions (tube feet)
which function in locomotion, feeding and gas exchange
Julia Heunis
 PRESENT!
 Coelom = A fluid-filled cavity between body wall and gut;
lined by mesoderm
 Two parts:
Perivisceral coelom – large, fluid-filled cavity where major
organs, digestive tube and sex organs, are suspended
Water Vascular System - network of hydraulic canals unique
to echinoderms that branches into extensions (tube feet)
which function in locomotion, feeding and gas exchange
http://www.colin.edu/science/awashington/syllabi/General%20Zoology%20Terminology.htm
http://biology.kenyon.edu/courses/biol112/Biol112WebPage/Syllabus/Topics/Week%207/Resources/coelom.GIF
http://www.britannica.com/EBchecked/topic/452108/perivisceral-coelom
http://www.und.nodak.edu/instruct/jvaughan/b151/VOCAB_vertebrates.html
Body Symmetry
 evolved from animals with bilateral
symmetry
 adult echinoderms possess radial
symmetry
 All echinoderms exhibit fivefold radial
symmetry in portions of their body at some
stage of life (5 parts around central axis),
even if they have secondary bilateral
symmetry
 For the most part…
LARVAE ~ bilateral
ADULTS ~ radial
Julia Heunis
http://tbn2.google.com/images?q=tbn:KrJSbsj0l4ImqM:http://
photos.tobinhosting.com/d/117553-2/sea%2Burchins.jpg
 evolved from animals with bilateral
symmetry
 adult echinoderms possess radial
symmetry
 All echinoderms exhibit fivefold radial
symmetry in portions of their body at some
stage of life (5 parts around central axis),
even if they have secondary bilateral
symmetry
 For the most part…
LARVAE ~ bilateral
ADULTS ~ radial
http://en.wikipedia.org/
wiki/Echinoderm
Excretory System
 NO true excretory
system!
 Main opening of a
sponge used only to
EXPEL WASTE
 Anus leads directly from
stomach/digestive tract
 Anus = top
mouth = bottom
Julia Heunis
 NO true excretory
system!
 Main opening of a
sponge used only to
EXPEL WASTE
 Anus leads directly from
stomach/digestive tract
 Anus = top
mouth = bottom
universe-review.ca
Olivia Heunis
http://universe-review.ca/I10-82-starfish.jpg
Skeletal Type
Echinodermata: “Spiny Skinned”
 DO NOT possess an exoskeleton
 a thin outermost skin covers a mesodermal
endoskeleton made of tiny calcified plates and
spines - forms rigid support contained within
tissues of the organism
 Skeleton composed of skeletal plates called
ossicles
 Ossicles = small bones
 In some species, such as the sea urchin, plates
of the skeleton are locked together to form a
rigid structure.
 most sea stars and brittle stars can flex their
‘arms’ = skeleton has gaps, flexible plate
junctures
Julia Heunis
 DO NOT possess an exoskeleton
 a thin outermost skin covers a mesodermal
endoskeleton made of tiny calcified plates and
spines - forms rigid support contained within
tissues of the organism
 Skeleton composed of skeletal plates called
ossicles
 Ossicles = small bones
 In some species, such as the sea urchin, plates
of the skeleton are locked together to form a
rigid structure.
 most sea stars and brittle stars can flex their
‘arms’ = skeleton has gaps, flexible plate
junctures
http://ebiomedia.com/prod/BOechinoderms.html
http://en.wikipedia.org/wiki/File:Sanc0521.jpg
Other Unique Features
 Echinoderms can REGENERATE
ex: sea cucumbers can eject a portion of
gut in response to predators and
regenerate when safe
 Water vascular system (described earlier)
is unique to echinoderms; varies water
pressure to control tube feet
 Crinoids and some brittle stars = passive
filter-feeders, absorbing suspended
particles from passing water
sea urchins = grazers
sea cucumbers = deposit feeders
starfish = active hunters
Julia Heunis
 Echinoderms can REGENERATE
ex: sea cucumbers can eject a portion of
gut in response to predators and
regenerate when safe
 Water vascular system (described earlier)
is unique to echinoderms; varies water
pressure to control tube feet
 Crinoids and some brittle stars = passive
filter-feeders, absorbing suspended
particles from passing water
sea urchins = grazers
sea cucumbers = deposit feeders
starfish = active hunters
http://www.starfish.ch/reef/echinoderms.html
More Unique Features
 Autonomy = The spontaneous
self amputation of an
appendage when the organism
is injured or under attack. The
autotomized part is usually
regenerated.

have more because of
regeneration
 mutable collagenous tissue ~
connect ossicles; can maintain
different positions without
much effort
Julia Heunis
 Autonomy = The spontaneous
self amputation of an
appendage when the organism
is injured or under attack. The
autotomized part is usually
regenerated.

have more because of
regeneration
 mutable collagenous tissue ~
connect ossicles; can maintain
different positions without
much effort
http://www.starfish.ch/reef/echinoderms.html
Nervous System
 Decentralized
 central nerve ring surrounds gut, connect
radial nerves.
 Radial nerves run under each arm, coordinate
movement, etc.
 Do not have “brains,” but some have ganglia
along radial nerves
David Lin
 Decentralized
 central nerve ring surrounds gut, connect
radial nerves.
 Radial nerves run under each arm, coordinate
movement, etc.
 Do not have “brains,” but some have ganglia
along radial nerves
http://www.tolweb.org/Echinodermata
Digestive System
 Complete digestive system
(tubular gut), mouth to anus.
(bottom to top)
 Crinoidia: filter-feed, use cilia
 Asteroidea: Cardiac (evert),
pyloric stomach, digestive glands
 Urchins: “Aristotle’s lantern”
(masticatory apparatus)
 Holothurians: suspension/deposit
feeders, use tentacles.
http://www.experiencefestival.com/a/Sea_star_-
_Internal_Anatomy/id/2099728
David Lin
 Complete digestive system
(tubular gut), mouth to anus.
(bottom to top)
 Crinoidia: filter-feed, use cilia
 Asteroidea: Cardiac (evert),
pyloric stomach, digestive glands
 Urchins: “Aristotle’s lantern”
(masticatory apparatus)
 Holothurians: suspension/deposit
feeders, use tentacles.
http://www.experiencefestival.com/a/Sea_star_-
_Internal_Anatomy/id/2099728
http://userwww.sfsu.edu/~biol240/labs/lab_17hydrostatics
k/pages/echinoderm.html
Locomotion/Musculature
 Utilize water vascular
system and tube feet
 Water in from
madreporite, pumped into
ampulla by radial canal
 Ampulla contracts, water
to podia
 Podia contracts, water to
ampulla
– Podia bend, shorten
allows movement
(vacuum and suction)
David Lin
 Utilize water vascular
system and tube feet
 Water in from
madreporite, pumped into
ampulla by radial canal
 Ampulla contracts, water
to podia
 Podia contracts, water to
ampulla
– Podia bend, shorten
allows movement
(vacuum and suction)
http://animaldiversity.ummz.umich.edu/site/resources/biodidac/basicechino.jpg/mediu
m.jpg
http://userwww.sfsu.edu/~biol240/labs/lab_17hydrostaticsk/pages/echinoderm.html
Gas Exchange
 Asteroids, crinoids: dermal
gills (thin epidermis)
 Urchins: infolds of body wall
(bursae) opening to outside
 Sand dollars: modified podia
on top (petaloids) which are
thin-walled and flaplike.
 Holothurians: highly branched
hindgut called "respiratory
tree"
David Lin
 Asteroids, crinoids: dermal
gills (thin epidermis)
 Urchins: infolds of body wall
(bursae) opening to outside
 Sand dollars: modified podia
on top (petaloids) which are
thin-walled and flaplike.
 Holothurians: highly branched
hindgut called "respiratory
tree"
http://faculty.vassar.edu/mehaffey/academic/animalstructure/outlines/echinode
rmata.html
Circulatory System
 Mostly in perivisceral
coelom, enhanced by
water vascular system
and hemal system.
– Hemal system: series
of canals and spaces
mostly in coelomic
channels. Fluid
movement by cilia.
Used to distribute
nutrients. No
respiratory system.
David Lin
 Mostly in perivisceral
coelom, enhanced by
water vascular system
and hemal system.
– Hemal system: series
of canals and spaces
mostly in coelomic
channels. Fluid
movement by cilia.
Used to distribute
nutrients. No
respiratory system.
http://faculty.vassar.edu/mehaffey/academic/animalstructure/outlines/
echinodermata.html
http://www.dailygalaxy.com/photos/uncategorized/urchins_1.jpg
Quiz
1. Echinodermata have ________ symmetry as
larvae, but display ________ symmetry as adults.
2. What allows Echinoderms to function in locomotion,
feeding and gas exchange?
3. Sea stars can grow new arms and sea cucumbers
can replace their gut after ejecting them. What
feature of echinoderms does this demonstrate?
4. Name the stomachs of a sea star. Functions?
1. Echinodermata have ________ symmetry as
larvae, but display ________ symmetry as adults.
2. What allows Echinoderms to function in locomotion,
feeding and gas exchange?
3. Sea stars can grow new arms and sea cucumbers
can replace their gut after ejecting them. What
feature of echinoderms does this demonstrate?
4. Name the stomachs of a sea star. Functions?
Answers
 Bilateral, radial
 Water vascular
system
 Regeneration
 Cardiac, pyloric;
evert and enzymes,
further digestion
 Bilateral, radial
 Water vascular
system
 Regeneration
 Cardiac, pyloric;
evert and enzymes,
further digestion

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Echinodermata presentation

  • 2. Echinoderm Classes Asteroidea – Sea Stars:  Multiple arms radiating from central disk  Tube feet on bottom Riley Galton Asteroidea – Sea Stars:  Multiple arms radiating from central disk  Tube feet on bottom Ophiuroidea – Brittle Stars:  Distinct central disk with snake like arms  Tube feet lack suckers http://photography.nationalgeographic.com/staticfiles/ http://environment.nationalgeographic.com/staticfiles/
  • 3. Echinoderm Classes Echinoidea – Sea Urchins and Sand Dollars:  No arms, but 5 rows of tube feet  Have muscles that can move  spines Riley Galton Echinoidea – Sea Urchins and Sand Dollars:  No arms, but 5 rows of tube feet  Have muscles that can move  spines http://environment.nationalgeographic.com/staticfiles/ www.emiliogrossi.com/images/Photo graphyiles/ Crinoidea – Feather Stars and Sea Lilies  Flexible arms used in suspension feeding  Haven’t changed in 500 million years http://lh6.ggpht.com http://image44.webshots.com
  • 4. Echinoderm Classes Holothuroidea – Sea Cucumbers  Reduced endoskeleton  Five rows of tube feet Riley Galton Holothuroidea – Sea Cucumbers  Reduced endoskeleton  Five rows of tube feet Concentricycloidea – Sea Daisies  Only two known species  Disk shaped, less than 1cm in diameter http://blog.case.edu/case-news/2008/03/06/seacucumber.jpg http://www.sfu.ca/~fankbone/v/xyloplax.jpg
  • 5. Echinoderm Sensory Structures  Eyespots on the ends of ambulacra are sensitive to light  Pedicellariae (jaw like skeletal structures used to keep organisms from settling on skin), tube feet, and spines are sensitive to touch http://www.ucmp.berkeley.edu/echinodermata/echinomm.html Riley Galton  Eyespots on the ends of ambulacra are sensitive to light  Pedicellariae (jaw like skeletal structures used to keep organisms from settling on skin), tube feet, and spines are sensitive to touch http://www.ucmp.berkeley.edu/echinodermata/echinomm.html
  • 6. Echinoderm Reproduction Sexual: • Eggs and sperm are released into the water where they are fertilized • Larvae are planktonic • Internal fertilization has been observed in certain species of sea stars, brittle stars, and sea cucumbers Asexual: • Echinoderms can regenerate body parts and organs • Asexual reproduction can occur (though not by will) if the echinoderm is torn with enough of the body on each part http://www.oceaninn.com/guides/echino.htm Riley Galton http://www.biolbull.org/content/vol204/issue3/images/medium/cov erfig.gif Sexual: • Eggs and sperm are released into the water where they are fertilized • Larvae are planktonic • Internal fertilization has been observed in certain species of sea stars, brittle stars, and sea cucumbers Asexual: • Echinoderms can regenerate body parts and organs • Asexual reproduction can occur (though not by will) if the echinoderm is torn with enough of the body on each part http://www.oceaninn.com/guides/echino.htm http://www.starfish.ch/Fotos/echinoderms
  • 7. Body Cavity (Coelom)  PRESENT!  Coelom = A fluid-filled cavity between body wall and gut; lined by mesoderm  Two parts: Perivisceral coelom – large, fluid-filled cavity where major organs, digestive tube and sex organs, are suspended Water Vascular System - network of hydraulic canals unique to echinoderms that branches into extensions (tube feet) which function in locomotion, feeding and gas exchange Julia Heunis  PRESENT!  Coelom = A fluid-filled cavity between body wall and gut; lined by mesoderm  Two parts: Perivisceral coelom – large, fluid-filled cavity where major organs, digestive tube and sex organs, are suspended Water Vascular System - network of hydraulic canals unique to echinoderms that branches into extensions (tube feet) which function in locomotion, feeding and gas exchange http://www.colin.edu/science/awashington/syllabi/General%20Zoology%20Terminology.htm http://biology.kenyon.edu/courses/biol112/Biol112WebPage/Syllabus/Topics/Week%207/Resources/coelom.GIF http://www.britannica.com/EBchecked/topic/452108/perivisceral-coelom http://www.und.nodak.edu/instruct/jvaughan/b151/VOCAB_vertebrates.html
  • 8. Body Symmetry  evolved from animals with bilateral symmetry  adult echinoderms possess radial symmetry  All echinoderms exhibit fivefold radial symmetry in portions of their body at some stage of life (5 parts around central axis), even if they have secondary bilateral symmetry  For the most part… LARVAE ~ bilateral ADULTS ~ radial Julia Heunis http://tbn2.google.com/images?q=tbn:KrJSbsj0l4ImqM:http:// photos.tobinhosting.com/d/117553-2/sea%2Burchins.jpg  evolved from animals with bilateral symmetry  adult echinoderms possess radial symmetry  All echinoderms exhibit fivefold radial symmetry in portions of their body at some stage of life (5 parts around central axis), even if they have secondary bilateral symmetry  For the most part… LARVAE ~ bilateral ADULTS ~ radial http://en.wikipedia.org/ wiki/Echinoderm
  • 9. Excretory System  NO true excretory system!  Main opening of a sponge used only to EXPEL WASTE  Anus leads directly from stomach/digestive tract  Anus = top mouth = bottom Julia Heunis  NO true excretory system!  Main opening of a sponge used only to EXPEL WASTE  Anus leads directly from stomach/digestive tract  Anus = top mouth = bottom universe-review.ca
  • 11. Skeletal Type Echinodermata: “Spiny Skinned”  DO NOT possess an exoskeleton  a thin outermost skin covers a mesodermal endoskeleton made of tiny calcified plates and spines - forms rigid support contained within tissues of the organism  Skeleton composed of skeletal plates called ossicles  Ossicles = small bones  In some species, such as the sea urchin, plates of the skeleton are locked together to form a rigid structure.  most sea stars and brittle stars can flex their ‘arms’ = skeleton has gaps, flexible plate junctures Julia Heunis  DO NOT possess an exoskeleton  a thin outermost skin covers a mesodermal endoskeleton made of tiny calcified plates and spines - forms rigid support contained within tissues of the organism  Skeleton composed of skeletal plates called ossicles  Ossicles = small bones  In some species, such as the sea urchin, plates of the skeleton are locked together to form a rigid structure.  most sea stars and brittle stars can flex their ‘arms’ = skeleton has gaps, flexible plate junctures http://ebiomedia.com/prod/BOechinoderms.html http://en.wikipedia.org/wiki/File:Sanc0521.jpg
  • 12. Other Unique Features  Echinoderms can REGENERATE ex: sea cucumbers can eject a portion of gut in response to predators and regenerate when safe  Water vascular system (described earlier) is unique to echinoderms; varies water pressure to control tube feet  Crinoids and some brittle stars = passive filter-feeders, absorbing suspended particles from passing water sea urchins = grazers sea cucumbers = deposit feeders starfish = active hunters Julia Heunis  Echinoderms can REGENERATE ex: sea cucumbers can eject a portion of gut in response to predators and regenerate when safe  Water vascular system (described earlier) is unique to echinoderms; varies water pressure to control tube feet  Crinoids and some brittle stars = passive filter-feeders, absorbing suspended particles from passing water sea urchins = grazers sea cucumbers = deposit feeders starfish = active hunters http://www.starfish.ch/reef/echinoderms.html
  • 13. More Unique Features  Autonomy = The spontaneous self amputation of an appendage when the organism is injured or under attack. The autotomized part is usually regenerated.  have more because of regeneration  mutable collagenous tissue ~ connect ossicles; can maintain different positions without much effort Julia Heunis  Autonomy = The spontaneous self amputation of an appendage when the organism is injured or under attack. The autotomized part is usually regenerated.  have more because of regeneration  mutable collagenous tissue ~ connect ossicles; can maintain different positions without much effort http://www.starfish.ch/reef/echinoderms.html
  • 14. Nervous System  Decentralized  central nerve ring surrounds gut, connect radial nerves.  Radial nerves run under each arm, coordinate movement, etc.  Do not have “brains,” but some have ganglia along radial nerves David Lin  Decentralized  central nerve ring surrounds gut, connect radial nerves.  Radial nerves run under each arm, coordinate movement, etc.  Do not have “brains,” but some have ganglia along radial nerves http://www.tolweb.org/Echinodermata
  • 15. Digestive System  Complete digestive system (tubular gut), mouth to anus. (bottom to top)  Crinoidia: filter-feed, use cilia  Asteroidea: Cardiac (evert), pyloric stomach, digestive glands  Urchins: “Aristotle’s lantern” (masticatory apparatus)  Holothurians: suspension/deposit feeders, use tentacles. http://www.experiencefestival.com/a/Sea_star_- _Internal_Anatomy/id/2099728 David Lin  Complete digestive system (tubular gut), mouth to anus. (bottom to top)  Crinoidia: filter-feed, use cilia  Asteroidea: Cardiac (evert), pyloric stomach, digestive glands  Urchins: “Aristotle’s lantern” (masticatory apparatus)  Holothurians: suspension/deposit feeders, use tentacles. http://www.experiencefestival.com/a/Sea_star_- _Internal_Anatomy/id/2099728 http://userwww.sfsu.edu/~biol240/labs/lab_17hydrostatics k/pages/echinoderm.html
  • 16. Locomotion/Musculature  Utilize water vascular system and tube feet  Water in from madreporite, pumped into ampulla by radial canal  Ampulla contracts, water to podia  Podia contracts, water to ampulla – Podia bend, shorten allows movement (vacuum and suction) David Lin  Utilize water vascular system and tube feet  Water in from madreporite, pumped into ampulla by radial canal  Ampulla contracts, water to podia  Podia contracts, water to ampulla – Podia bend, shorten allows movement (vacuum and suction) http://animaldiversity.ummz.umich.edu/site/resources/biodidac/basicechino.jpg/mediu m.jpg http://userwww.sfsu.edu/~biol240/labs/lab_17hydrostaticsk/pages/echinoderm.html
  • 17. Gas Exchange  Asteroids, crinoids: dermal gills (thin epidermis)  Urchins: infolds of body wall (bursae) opening to outside  Sand dollars: modified podia on top (petaloids) which are thin-walled and flaplike.  Holothurians: highly branched hindgut called "respiratory tree" David Lin  Asteroids, crinoids: dermal gills (thin epidermis)  Urchins: infolds of body wall (bursae) opening to outside  Sand dollars: modified podia on top (petaloids) which are thin-walled and flaplike.  Holothurians: highly branched hindgut called "respiratory tree" http://faculty.vassar.edu/mehaffey/academic/animalstructure/outlines/echinode rmata.html
  • 18. Circulatory System  Mostly in perivisceral coelom, enhanced by water vascular system and hemal system. – Hemal system: series of canals and spaces mostly in coelomic channels. Fluid movement by cilia. Used to distribute nutrients. No respiratory system. David Lin  Mostly in perivisceral coelom, enhanced by water vascular system and hemal system. – Hemal system: series of canals and spaces mostly in coelomic channels. Fluid movement by cilia. Used to distribute nutrients. No respiratory system. http://faculty.vassar.edu/mehaffey/academic/animalstructure/outlines/ echinodermata.html http://www.dailygalaxy.com/photos/uncategorized/urchins_1.jpg
  • 19. Quiz 1. Echinodermata have ________ symmetry as larvae, but display ________ symmetry as adults. 2. What allows Echinoderms to function in locomotion, feeding and gas exchange? 3. Sea stars can grow new arms and sea cucumbers can replace their gut after ejecting them. What feature of echinoderms does this demonstrate? 4. Name the stomachs of a sea star. Functions? 1. Echinodermata have ________ symmetry as larvae, but display ________ symmetry as adults. 2. What allows Echinoderms to function in locomotion, feeding and gas exchange? 3. Sea stars can grow new arms and sea cucumbers can replace their gut after ejecting them. What feature of echinoderms does this demonstrate? 4. Name the stomachs of a sea star. Functions?
  • 20. Answers  Bilateral, radial  Water vascular system  Regeneration  Cardiac, pyloric; evert and enzymes, further digestion  Bilateral, radial  Water vascular system  Regeneration  Cardiac, pyloric; evert and enzymes, further digestion