Biology is the study of life. It began with early Greek philosophers who began classifying living things. Important figures like Aristotle, Galen, and Andreas Vesalius made advances in anatomy and classification. William Harvey discovered blood circulation. The development of the microscope allowed Anton van Leeuwenhoek to discover microorganisms. Charles Darwin developed the theory of evolution by natural selection. The major divisions of biology are botany, which is the study of plants, and zoology, which is the study of animals. Key areas of biology include anatomy, physiology, ecology, taxonomy, embryology, genetics, and microbiology.