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Dr.	A.S.	Naik	
Head,	Dept.	of	Botany,	
S.B.E.S.	College	of	Science.	
MORPHOLOGY	OF	ANGIOSPERMS-4	
FLOWER
Reproduc+ve	structure-flower
Dissected	flower
TERMS	USED	TO	DESCRIBE	FLOWERS	
																																Pedicillate/Sessile
Complete/Incomplete	flowers	or	Perfect/
Imperfect	flowers
Symmetrical/Asymmetrical	flowers
Bisexual/Unisexual	flowers	
e.g	Pterygota	alata	
												Female	flowers																																																									Male	flowers	
With	nonfunc+onal	stamens																														with	monoadephous	stamens	
						(Pis+late	flowers)																																																			(staminate	flowers)
A.	Staminate,	B.	Bisexual	and	C.	pis+late	flowers	of		
Carica	papaya
Bracteate/Ebracteate	flowers
Floral	bracts:	Sepaloid/petaloid	
												Sunflower																																											Banana																															Bougainvillea	
				Anthurium																																												Poinse5a
Inser+on	of	floral	whorls	on	thalamus/receptacle
Hypogynae:	Bloodroot,	Hibiscus																																Epigynae	in	Cucumber	
																				Perigynae	in	Prunus
Extension	of	receptacle	axixs	
Androphore,	Gynophore	and	Gynandrophore	
•  GYNOPHORE	IN	Capparis
Gynandrophore	
Passion	flower
Non-essen+al/Accessory	whorls:	
1.	Calyx-Lowermost,	protec+ve	whorl	
		Free	sepals-Polysepalous																														Fused	sepals-	Gamosepalous	
											e.g.	Mustard																																																												e.g.	Datura	
•  	Members	called	as	sepals
2.	Corolla	
Meant	for	aVrac+on	
Polypetalous-	Free	petals																																										Gamopetalous-	Fused	petals	
											e.g.	Hibiscus																																																																				e.g.	Datura	
																																	Members	called	as	petals
Types	of	Corolla:	
INFUNDIBULIFORM	COROLLA	
1.Infundibuliform-Funnel	shaped	
	
•  The	corolla	resembles	an	inverted	
cone	like	a	funnel	as	seen	in	
Datura	or	many	plants	of	the	
family	Convolvulaceae	like	
Ipomoea	pulchella	.	
	
e.g.Datura
CRUCIFORM	COROLLA	
2.	Cruciform:	
Four	free	clawed	petals	are	arranged	
	in	the	form	of	a	cross	as	in	the	
	Cruciferae	family,		
	
	
e.g.,	mustard		
	
			e.g.	Mustard	(Brassica)
BILABIATE	COROLLA	
3.Bilabiate:	Two	lipped	
•  Limb	is	divided	into	an	upper	
posterior	part	(the	union	of	two	
petal	lobes)	and	an	unequal	lower	
anterior	part	(	the	union	of	three	
petal	lobes)	with	the	mouth	
gaping	wide	open.	
•  This	is	characteris+c	of	the	family	
Labiatae	:	Basil,	Salvia	
								e.g.	Salvia	sp
Papilionaceous	/	Vexillary	Corolla	
							e.g.	Clitoria	ternatea	
4.Papilionaceous	corolla		
	
•  Five	free	petals	resemble	a	buVerfly	
(papilion=buVerfly).	The	posterior	
superior	petal	is	larger	than	the	
others	and	is	termed	the	vexillum	or	
standard.	
	
•  two	anterior	inferior	ones	are	
usually	more	or	less	united	forming	
a	boat-shaped	struc-ture	and	are	
called	the	carina	or	keel.	
	
•  Two	lateral	petals	called	alae	or	
wings	overlap	the	carina	and	are	
themselves	overlapped	by	the	
vexillum.
AESTIVATION	
•  Aes:va:on	or	es:va:o
n	refers	to	the	
posi+onal	arrangement	
of	the	parts	of	a	flower	
within	a	flower	bud	
before	it	has	opened.
Types	of	Aes+va+on	
1.valvate	–	margins	of	
adjacent	petals	or	sepals	touch	
each	other	without	
overlapping.	
	
2.	Twisted:	margins	of	
adjacent	petals	or	sepals	
overlapping.
Types	of	Aes+va+on	
3.	Imbricate:	One	petal		
innermost,	one	petal	
outermost	and	
Rest	of	the	petals	twisted.	
	
	
	
	
4.	Vexillary:		5	petals-one	
largest	outermost	standard/
vexillum/banner	encloses	two	
middle	smaller	petals-wings,	
which	in	turn	enclose	two	
smallest	and	innermost,	folded	
petals-keel
Essen+al	whorls:	1.	Androecium	
Male	reproduc+ve	whorl
Types	of	anthers		
Filament	to	anther	aVachment	
•  Dorsifixed:	The	aVachment	is	
dorsifixed	when	the	filament	
is	firmly	fixed	to	some	
posi+on	on	the	back	of	the	
an-ther	as	in	passion-flower,	
Sesbania,	etc.		
•  Basifixed:	The	filament	ends	
just	at	the	base	of	the	anther,	
the	laVer	being	firmly	fixed	
on	the	top	of	the	former.	This	
condi+on	is	called	basifixed	.	
•  Versa+le:	where	the	filament,	
is	aVached	merely	at	a	point	
about	the	middle	of	the	
connec+ve	so	that	the	anther	
can	swing	on	it	freely.
Anther	lobe	types
Stamen	arrangement	&	grouping	
•  Didynamous:	
4	stamens,	2	short,	2	long	
e.g.	Mint	
	
	
•  Tetradynamous:	
6	stamens,	2	outer	short	and	4	inner	
Long.	
e.g.	Cabbage
Stamen	cohesion	and	adelphy	
•  Cohesion	usually	
involves	either	
only	the	filaments	
(adelphy)	or	only	
the	anthers	
(syngeny).		
•  In	adelphy,	all	the	
stamens	may	unite	
by	their	filaments	
forming	one	
bundle	of	stamens	
with	all	the	
anthers	free.
Stamen	cohesion	types
Monoadelphy,	diadelphyphy	and	polyadelphy	
•  monadelphy:		In	the	family	
Malvaceae	and	in	many	other	flowers	
the	united	filamen’s	form	a	staminal	
lube	through	which	the	long	style	of	
the	pis+l	passes.	
•  Diadelphy:		(two	bundles)	is	very	
commonly	seen	in	Papilionaceous	
flowers	where	rine	stamens	form	one	
bundle	and	the	tenth	remains	free	as	
the	second	bundle		
•  Polyadelphy:		In	the	silk-coVon	tree	
(Salmalia	or	Bombax	ceiba)	the	
stamens	form	several	separate	
groups	with	the	filaments	uni+ng	to	
form	several	bundles	to	form	the	
polyadelphous	condi+on
Syngeny	and	synandry	
•  Syngeny:	In	the	family	
Compositae,		the	
syngenesious	anthers	
form	a	tube	enclosing	the	
style	and	the	s+gma.		
	
	
•  Synandrous:	In	Family	
cucurbitaceae	complete	
union	of	the	filaments	as	
well	as	the	sinuous	
anthers	.	This	is	called	the	
synandrous	condi+on.
Cohesion	of	stamens
Adhesion	of	stamens	
								Epipetalous	stamens:	e.g.	Brinjal-Fusion	of	androecium	and		
																																																									corolla	
	
									Epiphyllous:	e.g.	Parnassia-Fusin	of	androecium	and		
																																													gynoecium
Gynoestagium	of	Calotropis	
•  The	s+gmas		and		
androeciums	are	fused	
to	form	a	gynostegium	.	
•  The	pollen	are	enclosed	
in	pollinia	(a	coherent	
mass	of	pollen	grains)		
•  When	the	bee	visits		
flower,	the	flower	is	
pollinated	by	the	
adhering	of	pollinium	on	
the	legs	of	bee.
Gynoecium	
						Female	reproduc+ve	essen+al	whorl
Types	of	gynoecium	
1.	Apocarpous-	free	carpels
2.	Syncarpous-	fused	carpels
Chambers/locules	of	ovary
Placenta+on:	AVachment	of	ovules	to	Ovary
Axile	placenta+on	e.g.	Hibiscus	
•  The	placenta	is	axial	
and	the	ovules	are	
aVached	to	the	bi-or	
mul+locular	superior	
ovary.	Example:	China	
rose,	lemon	and	orange
Marginal	Placenta+on	
	
	
The	placenta	forms	a	
ridge	along	the	ventral	
suture	of	the	ovary	and	
the	ovules	are	borne	on	
the	ridge.	Example:	Peas
Basal	Placenta+on	
•  In	this	type	of	
placenta+on,	the	
placenta	is	situated	at	
the	base	of	the	ovary	
and	a	single	ovule	is	
aVached	to	it.	Example:	
Sunflower
Parietal	Placenta+on	
•  In	this	type	of	
placenta+on,	the	ovules	
develop	on	the	inner	
wall	of	the	ovary.	Ovary	
has	a	single	chamber	
but	appears	to	be	
double-chambered	
because	of	the	
forma+on	of	a	false	
septum.	Example:	
Mustard	and	argemone.
Seed	dispersal
Seed	dispersal-Hydrochory
2.	Anemochory-Wind	dispersal
Hydrochory
Video	autochory

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