Birds have evolved various modifications to their feet depending on their environment and locomotion. Some key foot modifications include: 1. Running or cursorial feet with reduced toes for fast running in ostriches, bustards, and emus. 2. Perching feet with three toes directed forward and a rear-facing hallux for gripping branches in birds like sparrows and robins. 3. Webbed feet with lobes or fully united toes for efficient swimming in aquatic birds such as ducks, grebes, and pelicans.