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This document summarizes theories of root apex organization. It defines the root apex as the region where primary tissues are derived and composed of apical meristem cells. Two historical theories are described: 1) The Apical Cell Theory proposed a single apical cell governs growth but was later replaced. 2) The Korper-Kappe Theory proposed cells divide in two planes, with one zone resembling an inverted T and the other a straight T, similar to the tunica-corpus model of shoot apices. However, this theory also fails to fully explain differences between species.












