Skip to main content
NODAL ANATOMY
NODALANATOMY
 Shoots consists of stems bearing
leaves.
 The point where the leaf attached to
the stem called Node. ( site of
attachment of leaves to the stem )
 The distance between two nodes on
the shoot is called Vegetative
Internodes.
LEAF TRACES
 Leaves must have access to a source of water and
essential minerals.
 This transport of substances takes place in primary
vascular connections between the stem vascular system
and the base of leaves called Leaf traces.
 Leaf traces are composed of protoxylem, metaxylem,
protophloem and metaphloem.
 Typically contains transfer cells in both primary xylem
and primary phloem.
LEAF GAP
 The continuity of the cylinder of primary vascular tissue
is broken immediately above the position of outward
divergence of leaf trace, leaving a discontinuity is called
Leaf Gap.
 Since, these leaf gaps doesn’t affect vascular system.
 Lateral connectivity between tissues on top and bottom of
the leaf gap is formed, since vascular system connectivity
is breaked due to leaf gaps.
PHYLLOTAXY
PHYLLOTAXY
 “Phyllotaxis” or “Phyllotaxy” is the arrangement of
leaves on a stem.
 The word phyllotaxy is derived from ancient Greek word.
( Phyllo means leaf and taxis means arrangements )
 Phyllotaxy in mature plants is directly related to the
position & size of leaf primordia produced on apical
meristems of varying size and shape.
TYPES OF PHYLLOTAXY
 Based on the number of leaves and arrangement of leaves
from the nodes, the phyllotaxy has various types.
I. Alternate
II. Opposite
III. Whorled
Alternate Phyllotaxy
 When only one leaf develops at each node. Eg: Brassica
campestris
 Based on the position of leaves at after particular number
of leaves, alternate phyllotaxy is further subdivided into
two.
a) Distichous
b) Tristichous
A) Distichous:
 When 3rd leaf comes over the first one. Eg; Grasses.
B) Tristichous:
 When 4th leaf comes over the first one. Eg; Cypres.
Opposite Phyllotaxy
 When a pair of leaves are present just opposite to each
other at each node, eg; Calotropis.
 There are two types in opposite phyllotaxy.
A. Opposite Deccusate
B. Opposite Superposed
A) Oppposite Decussate:
 When two successive pairs of leaves occur at right angle
to each other, eg; Psidium gujava.
B) Opposite Superposed:
 When all pairs of leaves occur in same plane. Eg;
Combretum.
Whorled Phyllotaxy
 When more than two leaves are arranged in the form of
whorl at each node, the phyllotaxy is called Whorled. Eg;
Hydrilla verticillata, Nerium etc.,
Nodal anatomy, Leaf development & Phyllotaxy