The pneumatic tire was developed in 1888 by John Boyd Dunlop for bicycles but was quickly adapted for motor vehicles. Tires form the first part of the vehicle's suspension by cushioning it with air to absorb shocks from the road. Early tires used inner tubes but most modern tires are tubeless, which allows them to better seal air and self-seal when punctured. Tires must support the vehicle, provide adhesion, minimize rolling resistance, transmit drive, and absorb shock. The main tire types are radial and cross-ply tires, with radials having plies at 90 degrees to the bead for better road handling.