The phylum Arthropoda contains animals with bilateral symmetry, segmented bodies, and jointed appendages. They have an exoskeleton made of protein and chitin that protects their bodies and allows growth through molting. Arthropods live on land, in water, and everywhere in between, and eat a wide variety of foods. Their circulatory systems use hemolymph instead of blood, and respiratory organs vary between gills, book lungs, and tracheae depending on the environment. Reproduction is usually sexual, with sperm transferred to females in packets. Major subgroups include insects, chelicerates, crustaceans, and myriapods.