SlideShare a Scribd company logo
CHAPTER 16
Sensory, Motor, and
Integrative Systems
Principles of
Anatomy and
Physiology
14th Edition
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Sensation is the conscious or
subconscious awareness of changes in
the external or internal environment.
 Perception is the conscious interpretation
of sensations performed mainly by the
cerebral cortex.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Each type of sensation is called a
sensory modality. This includes: touch,
pain, vision and hearing.
 Sensory modalities are grouped into either
general senses or special senses.
 General senses: somatic-(tactile,
thermal, proprioceptive); visceral-
pressure, chemicals, stretch, nausea,
hunger, temperature.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Special senses: smell, taste, vision,
hearing and equilibrium.
 Different types of sensory receptors are
involved in the different senses.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Free nerve endings are used to detect
pain, temperature, tickle, itch and some
touch.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Encapsulated nerve endings are used to
detect pressure, vibration and some touch.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Sensory receptors for some special senses
include gustatory receptor cells in taste
buds, photoreceptors in the retina of the
eye and hair cells in the inner ear for
hearing.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Anatomy Overview
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 The Nervous System
You must be connected to the Internet and in Slideshow Mode to
run this animation.
 Receptors may also be grouped based on
location of the receptors and the origin of
the stimuli that activate them.
 Exteroreceptors include: hearing,
vision, smell, taste, touch, pressure,
vibration and pain.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Interoreceptors monitor the body’s
internal environment.
 Proprioceptors provide information about
body position, muscle length and
tension and the position and movement
of joints.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Sensation
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Somatic sensations include tactile,
thermal, pain and proprioceptive.
 Tactile sensations: touch, pressure,
vibration, itch and tickle.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 There are two types of pain: fast and
slow. Fast pain (acute, sharp or pricking)
perceived within 0.1 second.
 Slow pain (chronic, burning, aching or
throbbing) is perceived a second or more
after the stimulus.
 Superficial somatic pain: arising from
skin receptors.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Deep somatic pain: skeletal muscles,
joints, tendons and fascia.
 Stimulation of pain sensors in visceral
organs is visceral pain. This type of pain
usually presents in or just deep to the skin
that overlies the stimulated organ.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Proprioception: recognizing position of
body parts.
 Proprioceptors: in muscles and tendons.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Two types of proprioceptors are muscle spindles and tendon
organs. Muscle spindles: in skeletal muscles monitor their length
and are involved in stretch reflexes.
 Tendon organs: at the junction of a tendon and a muscle protect
muscles and tendons from damage due to overstretching.
 Joint kinesthetic receptors exist within and around the joint
capsule of synovial joints. They respond to pressure and
acceleration and deceleration during movement. Joint ligaments
contain receptors to protect against excessive strain.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensations
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic sensory pathways carry
information from somatic sensory receptors
to the primary somatosensory area in the
cerebral cortex and to the cerebellum. The
pathways to the cortex consist of thousands
of sets of three neurons classified as first,
second and third-order neurons.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 First-order neurons: impulses from
somatic receptors to the brain stem or
spinal cord.
 Second-order neurons: impulses from
the brain stem and spinal cord to the
thalamus.
 Third-order neurons: impulses from the
thalamus to the primary somatosensory
area of the cortex on the same side.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Somatic sensory impulses ascend to the
cerebral cortex along three general
pathways:
 The posterior column-medial lemniscus
pathway (impulses from the limbs, trunk,
neck and posterior head).
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensory
Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
The anterolateral (spinothalamic) pathway
(impulses for pain, temperature, itch, tickle
and posterior head).
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensory
Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 The trigeminothalamic pathway
(impulses for most somatic sensations—
tactile, thermal and pain—from the face,
nasal cavity, oral cavity and teeth).
 Somatic sensory impulses reach the
cerebellum via the spinocerebellar
tracts.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic
Sensory
Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Postcentral gyri located on both parietal
lobes of the brain are the sites for the
primary somatosensory area.
 Each region in this area receives sensory
input from a different part of the body on
the opposite side.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
The posterior spinocerebellar tract and
the anterior spinocerebellar tract carry
proprioceptive impulses to the cerebellum.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Sensory Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Nerves that extend out of the brain stem and
spinal cord are called lower motor neurons
(LMNs). These nerves innervate skeletal
muscles of the face and head through
cranial nerves, and skeletal muscles of the
limb and trunk through spinal nerves.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic motor pathways provide the input into
lower motor neurons and are divided into four
distinct circuits.
1. Local circuit neurons are located close to LMNs
in the brain stem and spinal cord.
2. Upper motor neurons (UMNs): input to both
lower circuit neurons and LMNs.
3. Basal nuclei neurons: assist movement by
providing input to UMNs.
4. Cerebellar neurons: assist movement via
control of activity of UMNs.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 UMNs extend to LMNs via two types of
pathways:
 Direct motor pathways deliver signals to
LMNs from the cerebral cortex.
 Indirect motor pathways deliver signals
to LMNs from motor centers in the basal
nuclei, cerebellum and cerebral cortex.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
There are two direct motor pathways used
for voluntary movement. These are the two
corticospinal pathways (the lateral
corticospinal tract and the anterior
corticospinal tract) and the corticobulbar
pathway.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Motor
Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Motor
Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic
Motor
Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Indirect motor pathways (extrapyramidal
pathways): rubrospinal, tectospinal,
vestibulospinal, lateral reticulospinal and
medial reticulospinal tracts.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
The cerebellum performs 4 activities:
1. Monitoring intentions for movement
2. Monitoring actual movement
3. Comparing command signals with
sensory information
4. Sending out corrective feedback
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Somatic Motor Pathways
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
 Wakefulness and sleep: relies on the
reticular activating system (RAS).
 Learning and memory: includes
immediate, short-term and long-term
memory.
Integrative Functions of the Cerebrum
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Copyright 2014 John Wiley & Sons, Inc.
All rights reserved. Reproduction or translation of this work beyond
that permitted in section 117 of the 1976 United States Copyright Act
without express permission of the copyright owner is unlawful.
Request for further information should be addressed to the
Permission Department, John Wiley & Sons, Inc. The purchaser may
make back-up copies for his/her own use only and not for distribution
or resale. The Publisher assumes no responsibility for errors,
omissions, or damages caused by the use of these programs or from
the use of the information herein.
End of Chapter 16
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.

More Related Content

What's hot

09 [chapter 9 joints]
09 [chapter 9 joints]09 [chapter 9 joints]
09 [chapter 9 joints]
Sompoch Thanachaikan
 
20 [chapter 20 the cardiovascular system the heart]
20 [chapter 20 the cardiovascular system   the heart]20 [chapter 20 the cardiovascular system   the heart]
20 [chapter 20 the cardiovascular system the heart]
Sompoch Thanachaikan
 
10 [chapter 10 muscular tissue]
10 [chapter 10 muscular tissue]10 [chapter 10 muscular tissue]
10 [chapter 10 muscular tissue]
Sompoch Thanachaikan
 
23 [chapter 23 the respiratory system]
23 [chapter 23 the respiratory system]23 [chapter 23 the respiratory system]
23 [chapter 23 the respiratory system]
Sompoch Thanachaikan
 
06 [chapter 6 the skeletal system bone tissue]
06 [chapter 6 the skeletal system   bone tissue]06 [chapter 6 the skeletal system   bone tissue]
06 [chapter 6 the skeletal system bone tissue]
Sompoch Thanachaikan
 
19 [chapter 19 the cardiovascular system the blood]
19 [chapter 19 the cardiovascular system   the blood]19 [chapter 19 the cardiovascular system   the blood]
19 [chapter 19 the cardiovascular system the blood]
Sompoch Thanachaikan
 
MOTOR SYSTEM IV REGULATION OF POSTURE
MOTOR SYSTEM IV REGULATION OF POSTUREMOTOR SYSTEM IV REGULATION OF POSTURE
MOTOR SYSTEM IV REGULATION OF POSTURE
Dr Nilesh Kate
 
Physiology of memory and learning
Physiology of memory and learning  Physiology of memory and learning
Physiology of memory and learning
Ali Faris
 
Reflex action
Reflex actionReflex action
Reflex action
Pavithra Pavi
 
Physiology of movement
Physiology of movementPhysiology of movement
Physiology of movement
aakankshabajpai2
 
12.postural reflexes kjg
12.postural reflexes kjg12.postural reflexes kjg
12.postural reflexes kjg
physiology mgmcri
 
Action Potential
Action Potential Action Potential
Action Potential
rashidrmc
 
Organisation Of Cns
Organisation Of CnsOrganisation Of Cns
Organisation Of Cnsraj kumar
 
Physiology of Sensory system
Physiology of Sensory systemPhysiology of Sensory system
Physiology of Sensory system
Eneutron
 
Muscle physiology
Muscle physiologyMuscle physiology
Muscle physiology
Medical Students
 
03 [chapter 3 the cellular level of organization]
03 [chapter 3 the cellular level of organization]03 [chapter 3 the cellular level of organization]
03 [chapter 3 the cellular level of organization]
Sompoch Thanachaikan
 
08 [chapter 8 the skeletal system appendicular skeleton]
08 [chapter 8 the skeletal system   appendicular skeleton]08 [chapter 8 the skeletal system   appendicular skeleton]
08 [chapter 8 the skeletal system appendicular skeleton]
Sompoch Thanachaikan
 
Spinal cord reflexes
Spinal cord reflexesSpinal cord reflexes
Spinal cord reflexesfarhan_aq91
 
Cerebral cortex ppt
Cerebral cortex pptCerebral cortex ppt
Cerebral cortex ppt
fiza rizvi
 

What's hot (20)

09 [chapter 9 joints]
09 [chapter 9 joints]09 [chapter 9 joints]
09 [chapter 9 joints]
 
20 [chapter 20 the cardiovascular system the heart]
20 [chapter 20 the cardiovascular system   the heart]20 [chapter 20 the cardiovascular system   the heart]
20 [chapter 20 the cardiovascular system the heart]
 
10 [chapter 10 muscular tissue]
10 [chapter 10 muscular tissue]10 [chapter 10 muscular tissue]
10 [chapter 10 muscular tissue]
 
Chapter 13
Chapter 13Chapter 13
Chapter 13
 
23 [chapter 23 the respiratory system]
23 [chapter 23 the respiratory system]23 [chapter 23 the respiratory system]
23 [chapter 23 the respiratory system]
 
06 [chapter 6 the skeletal system bone tissue]
06 [chapter 6 the skeletal system   bone tissue]06 [chapter 6 the skeletal system   bone tissue]
06 [chapter 6 the skeletal system bone tissue]
 
19 [chapter 19 the cardiovascular system the blood]
19 [chapter 19 the cardiovascular system   the blood]19 [chapter 19 the cardiovascular system   the blood]
19 [chapter 19 the cardiovascular system the blood]
 
MOTOR SYSTEM IV REGULATION OF POSTURE
MOTOR SYSTEM IV REGULATION OF POSTUREMOTOR SYSTEM IV REGULATION OF POSTURE
MOTOR SYSTEM IV REGULATION OF POSTURE
 
Physiology of memory and learning
Physiology of memory and learning  Physiology of memory and learning
Physiology of memory and learning
 
Reflex action
Reflex actionReflex action
Reflex action
 
Physiology of movement
Physiology of movementPhysiology of movement
Physiology of movement
 
12.postural reflexes kjg
12.postural reflexes kjg12.postural reflexes kjg
12.postural reflexes kjg
 
Action Potential
Action Potential Action Potential
Action Potential
 
Organisation Of Cns
Organisation Of CnsOrganisation Of Cns
Organisation Of Cns
 
Physiology of Sensory system
Physiology of Sensory systemPhysiology of Sensory system
Physiology of Sensory system
 
Muscle physiology
Muscle physiologyMuscle physiology
Muscle physiology
 
03 [chapter 3 the cellular level of organization]
03 [chapter 3 the cellular level of organization]03 [chapter 3 the cellular level of organization]
03 [chapter 3 the cellular level of organization]
 
08 [chapter 8 the skeletal system appendicular skeleton]
08 [chapter 8 the skeletal system   appendicular skeleton]08 [chapter 8 the skeletal system   appendicular skeleton]
08 [chapter 8 the skeletal system appendicular skeleton]
 
Spinal cord reflexes
Spinal cord reflexesSpinal cord reflexes
Spinal cord reflexes
 
Cerebral cortex ppt
Cerebral cortex pptCerebral cortex ppt
Cerebral cortex ppt
 

Similar to 16 [chapter 16 sensory, motor, and integrative systems]

Chapter 12 somatic senses and special
Chapter 12  somatic senses and specialChapter 12  somatic senses and special
Chapter 12 somatic senses and special
Nada G.Youssef
 
Receptor
ReceptorReceptor
Receptor
Sado Anatomist
 
sensitivity and coordination
sensitivity and coordinationsensitivity and coordination
sensitivity and coordinationCrelgo
 
SENSORY NERVOUS SYSTEM,drjma.pdf
SENSORY NERVOUS SYSTEM,drjma.pdfSENSORY NERVOUS SYSTEM,drjma.pdf
SENSORY NERVOUS SYSTEM,drjma.pdf
James Malce Alo, PhD, MAN, MAPsych, RN, OSHA
 
Concept of sensation with reference to organizational communication
Concept of sensation with reference to organizational communicationConcept of sensation with reference to organizational communication
Concept of sensation with reference to organizational communication
bp singh
 
10. Chp 11-12 PPT Sensory receptors.pptx
10. Chp 11-12 PPT Sensory receptors.pptx10. Chp 11-12 PPT Sensory receptors.pptx
10. Chp 11-12 PPT Sensory receptors.pptx
amanullahb42
 
The human nervous system and sense organ
The human nervous system and sense organThe human nervous system and sense organ
The human nervous system and sense organ
Anik Yudianto
 
Sense receptors
Sense receptors Sense receptors
Sense receptors
Lucknow University India
 
Classification of Sensory receptor 2
Classification of Sensory receptor  2Classification of Sensory receptor  2
Classification of Sensory receptor 2
Surjya Kumar Saikia
 
Balance for artifact
Balance for artifactBalance for artifact
Balance for artifactzs4033bn
 
Sensitivity and coordination by María
Sensitivity and coordination by MaríaSensitivity and coordination by María
Sensitivity and coordination by MaríaCrelgo
 
Physiology of sensory & pain.pdf
Physiology of sensory & pain.pdfPhysiology of sensory & pain.pdf
Physiology of sensory & pain.pdf
qahtanali2
 
Receptors classification
Receptors classificationReceptors classification
Receptors classification
Dr Sara Sadiq
 
Somatic Nervous System.pptx
Somatic Nervous System.pptxSomatic Nervous System.pptx
Somatic Nervous System.pptx
Karuna Yoga Vidya Peetham
 
The nervous system
The nervous systemThe nervous system
The nervous system
Alifya F
 
Bio 201 chapter 16 lecture
Bio 201 chapter 16 lectureBio 201 chapter 16 lecture
Bio 201 chapter 16 lectureMatt
 
Does that make sense
Does that make senseDoes that make sense
Does that make sense
cz0634bn
 
Does that make sense
Does that make senseDoes that make sense
Does that make sense
cz0634bn
 

Similar to 16 [chapter 16 sensory, motor, and integrative systems] (20)

Chapter 16
Chapter 16Chapter 16
Chapter 16
 
Chapter 12 somatic senses and special
Chapter 12  somatic senses and specialChapter 12  somatic senses and special
Chapter 12 somatic senses and special
 
Receptor
ReceptorReceptor
Receptor
 
sensitivity and coordination
sensitivity and coordinationsensitivity and coordination
sensitivity and coordination
 
SENSORY NERVOUS SYSTEM,drjma.pdf
SENSORY NERVOUS SYSTEM,drjma.pdfSENSORY NERVOUS SYSTEM,drjma.pdf
SENSORY NERVOUS SYSTEM,drjma.pdf
 
Concept of sensation with reference to organizational communication
Concept of sensation with reference to organizational communicationConcept of sensation with reference to organizational communication
Concept of sensation with reference to organizational communication
 
10. Chp 11-12 PPT Sensory receptors.pptx
10. Chp 11-12 PPT Sensory receptors.pptx10. Chp 11-12 PPT Sensory receptors.pptx
10. Chp 11-12 PPT Sensory receptors.pptx
 
The human nervous system and sense organ
The human nervous system and sense organThe human nervous system and sense organ
The human nervous system and sense organ
 
Sense receptors
Sense receptors Sense receptors
Sense receptors
 
Classification of Sensory receptor 2
Classification of Sensory receptor  2Classification of Sensory receptor  2
Classification of Sensory receptor 2
 
Balance for artifact
Balance for artifactBalance for artifact
Balance for artifact
 
Sensitivity and coordination by María
Sensitivity and coordination by MaríaSensitivity and coordination by María
Sensitivity and coordination by María
 
Sensory deprivation
Sensory deprivationSensory deprivation
Sensory deprivation
 
Physiology of sensory & pain.pdf
Physiology of sensory & pain.pdfPhysiology of sensory & pain.pdf
Physiology of sensory & pain.pdf
 
Receptors classification
Receptors classificationReceptors classification
Receptors classification
 
Somatic Nervous System.pptx
Somatic Nervous System.pptxSomatic Nervous System.pptx
Somatic Nervous System.pptx
 
The nervous system
The nervous systemThe nervous system
The nervous system
 
Bio 201 chapter 16 lecture
Bio 201 chapter 16 lectureBio 201 chapter 16 lecture
Bio 201 chapter 16 lecture
 
Does that make sense
Does that make senseDoes that make sense
Does that make sense
 
Does that make sense
Does that make senseDoes that make sense
Does that make sense
 

More from Sompoch Thanachaikan

22 [chapter 22 the lymphatic system and immunity]
22 [chapter 22 the lymphatic system and immunity]22 [chapter 22 the lymphatic system and immunity]
22 [chapter 22 the lymphatic system and immunity]
Sompoch Thanachaikan
 
28 [chapter 28 the reproductive system]
28 [chapter 28 the reproductive system]28 [chapter 28 the reproductive system]
28 [chapter 28 the reproductive system]
Sompoch Thanachaikan
 
29 [chapter 29 development and inheritance]
29 [chapter 29 development and inheritance]29 [chapter 29 development and inheritance]
29 [chapter 29 development and inheritance]
Sompoch Thanachaikan
 
27 [chapter 27 fluid, electrolyte and acid base homeostasis]
27 [chapter 27 fluid, electrolyte and acid base homeostasis]27 [chapter 27 fluid, electrolyte and acid base homeostasis]
27 [chapter 27 fluid, electrolyte and acid base homeostasis]
Sompoch Thanachaikan
 
26 [chapter 26 the urinary system]
26 [chapter 26 the urinary system]26 [chapter 26 the urinary system]
26 [chapter 26 the urinary system]
Sompoch Thanachaikan
 
24 [chapter 24 the digestive system][11e]
24 [chapter 24 the digestive system][11e]24 [chapter 24 the digestive system][11e]
24 [chapter 24 the digestive system][11e]
Sompoch Thanachaikan
 
21 [chapter 21 the cardiovascular system blood vessels and hemodynamics][11e]
21 [chapter 21 the cardiovascular system   blood vessels and hemodynamics][11e]21 [chapter 21 the cardiovascular system   blood vessels and hemodynamics][11e]
21 [chapter 21 the cardiovascular system blood vessels and hemodynamics][11e]
Sompoch Thanachaikan
 
11 [chapter 11 the muscular system][11e]
11 [chapter 11 the muscular system][11e]11 [chapter 11 the muscular system][11e]
11 [chapter 11 the muscular system][11e]
Sompoch Thanachaikan
 
07 [chapter 7 the skeletal system the axial skeleton]
07 [chapter 7 the skeletal system   the axial skeleton]07 [chapter 7 the skeletal system   the axial skeleton]
07 [chapter 7 the skeletal system the axial skeleton]
Sompoch Thanachaikan
 
05 [chapter 5 the integumentary system]
05 [chapter 5 the integumentary system]05 [chapter 5 the integumentary system]
05 [chapter 5 the integumentary system]
Sompoch Thanachaikan
 
04 [chapter 4 the tissue level of organization][11e]
04 [chapter 4 the tissue level of organization][11e]04 [chapter 4 the tissue level of organization][11e]
04 [chapter 4 the tissue level of organization][11e]
Sompoch Thanachaikan
 
02 [chapter 2 the chemical level of organization]
02 [chapter 2 the chemical level of organization]02 [chapter 2 the chemical level of organization]
02 [chapter 2 the chemical level of organization]
Sompoch Thanachaikan
 
01 [chapter 1 an introduction to the human body]
01 [chapter 1 an introduction to the human body]01 [chapter 1 an introduction to the human body]
01 [chapter 1 an introduction to the human body]
Sompoch Thanachaikan
 

More from Sompoch Thanachaikan (13)

22 [chapter 22 the lymphatic system and immunity]
22 [chapter 22 the lymphatic system and immunity]22 [chapter 22 the lymphatic system and immunity]
22 [chapter 22 the lymphatic system and immunity]
 
28 [chapter 28 the reproductive system]
28 [chapter 28 the reproductive system]28 [chapter 28 the reproductive system]
28 [chapter 28 the reproductive system]
 
29 [chapter 29 development and inheritance]
29 [chapter 29 development and inheritance]29 [chapter 29 development and inheritance]
29 [chapter 29 development and inheritance]
 
27 [chapter 27 fluid, electrolyte and acid base homeostasis]
27 [chapter 27 fluid, electrolyte and acid base homeostasis]27 [chapter 27 fluid, electrolyte and acid base homeostasis]
27 [chapter 27 fluid, electrolyte and acid base homeostasis]
 
26 [chapter 26 the urinary system]
26 [chapter 26 the urinary system]26 [chapter 26 the urinary system]
26 [chapter 26 the urinary system]
 
24 [chapter 24 the digestive system][11e]
24 [chapter 24 the digestive system][11e]24 [chapter 24 the digestive system][11e]
24 [chapter 24 the digestive system][11e]
 
21 [chapter 21 the cardiovascular system blood vessels and hemodynamics][11e]
21 [chapter 21 the cardiovascular system   blood vessels and hemodynamics][11e]21 [chapter 21 the cardiovascular system   blood vessels and hemodynamics][11e]
21 [chapter 21 the cardiovascular system blood vessels and hemodynamics][11e]
 
11 [chapter 11 the muscular system][11e]
11 [chapter 11 the muscular system][11e]11 [chapter 11 the muscular system][11e]
11 [chapter 11 the muscular system][11e]
 
07 [chapter 7 the skeletal system the axial skeleton]
07 [chapter 7 the skeletal system   the axial skeleton]07 [chapter 7 the skeletal system   the axial skeleton]
07 [chapter 7 the skeletal system the axial skeleton]
 
05 [chapter 5 the integumentary system]
05 [chapter 5 the integumentary system]05 [chapter 5 the integumentary system]
05 [chapter 5 the integumentary system]
 
04 [chapter 4 the tissue level of organization][11e]
04 [chapter 4 the tissue level of organization][11e]04 [chapter 4 the tissue level of organization][11e]
04 [chapter 4 the tissue level of organization][11e]
 
02 [chapter 2 the chemical level of organization]
02 [chapter 2 the chemical level of organization]02 [chapter 2 the chemical level of organization]
02 [chapter 2 the chemical level of organization]
 
01 [chapter 1 an introduction to the human body]
01 [chapter 1 an introduction to the human body]01 [chapter 1 an introduction to the human body]
01 [chapter 1 an introduction to the human body]
 

Recently uploaded

Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists
Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists  Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists
Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists
Saeid Safari
 
263778731218 Abortion Clinic /Pills In Harare ,
263778731218 Abortion Clinic /Pills In Harare ,263778731218 Abortion Clinic /Pills In Harare ,
263778731218 Abortion Clinic /Pills In Harare ,
sisternakatoto
 
Gram Stain introduction, principle, Procedure
Gram Stain introduction, principle, ProcedureGram Stain introduction, principle, Procedure
Gram Stain introduction, principle, Procedure
Suraj Goswami
 
Knee anatomy and clinical tests 2024.pdf
Knee anatomy and clinical tests 2024.pdfKnee anatomy and clinical tests 2024.pdf
Knee anatomy and clinical tests 2024.pdf
vimalpl1234
 
Basavarajeeyam - Ayurvedic heritage book of Andhra pradesh
Basavarajeeyam - Ayurvedic heritage book of Andhra pradeshBasavarajeeyam - Ayurvedic heritage book of Andhra pradesh
Basavarajeeyam - Ayurvedic heritage book of Andhra pradesh
Dr. Madduru Muni Haritha
 
Tom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness Journey
Tom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness JourneyTom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness Journey
Tom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness Journey
greendigital
 
planning for change nursing Management ppt
planning for change nursing Management pptplanning for change nursing Management ppt
planning for change nursing Management ppt
Thangamjayarani
 
Flu Vaccine Alert in Bangalore Karnataka
Flu Vaccine Alert in Bangalore KarnatakaFlu Vaccine Alert in Bangalore Karnataka
Flu Vaccine Alert in Bangalore Karnataka
addon Scans
 
Pharma Pcd Franchise in Jharkhand - Yodley Lifesciences
Pharma Pcd Franchise in Jharkhand - Yodley LifesciencesPharma Pcd Franchise in Jharkhand - Yodley Lifesciences
Pharma Pcd Franchise in Jharkhand - Yodley Lifesciences
Yodley Lifesciences
 
Evaluation of antidepressant activity of clitoris ternatea in animals
Evaluation of antidepressant activity of clitoris ternatea in animalsEvaluation of antidepressant activity of clitoris ternatea in animals
Evaluation of antidepressant activity of clitoris ternatea in animals
Shweta
 
Ophthalmology Clinical Tests for OSCE exam
Ophthalmology Clinical Tests for OSCE examOphthalmology Clinical Tests for OSCE exam
Ophthalmology Clinical Tests for OSCE exam
KafrELShiekh University
 
Cervical & Brachial Plexus By Dr. RIG.pptx
Cervical & Brachial Plexus By Dr. RIG.pptxCervical & Brachial Plexus By Dr. RIG.pptx
Cervical & Brachial Plexus By Dr. RIG.pptx
Dr. Rabia Inam Gandapore
 
Novas diretrizes da OMS para os cuidados perinatais de mais qualidade
Novas diretrizes da OMS para os cuidados perinatais de mais qualidadeNovas diretrizes da OMS para os cuidados perinatais de mais qualidade
Novas diretrizes da OMS para os cuidados perinatais de mais qualidade
Prof. Marcus Renato de Carvalho
 
ANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptx
ANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptxANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptx
ANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptx
Swetaba Besh
 
Hemodialysis: Chapter 3, Dialysis Water Unit - Dr.Gawad
Hemodialysis: Chapter 3, Dialysis Water Unit - Dr.GawadHemodialysis: Chapter 3, Dialysis Water Unit - Dr.Gawad
Hemodialysis: Chapter 3, Dialysis Water Unit - Dr.Gawad
NephroTube - Dr.Gawad
 
Top 10 Best Ayurvedic Kidney Stone Syrups in India
Top 10 Best Ayurvedic Kidney Stone Syrups in IndiaTop 10 Best Ayurvedic Kidney Stone Syrups in India
Top 10 Best Ayurvedic Kidney Stone Syrups in India
SwastikAyurveda
 
heat stroke and heat exhaustion in children
heat stroke and heat exhaustion in childrenheat stroke and heat exhaustion in children
heat stroke and heat exhaustion in children
SumeraAhmad5
 
NVBDCP.pptx Nation vector borne disease control program
NVBDCP.pptx Nation vector borne disease control programNVBDCP.pptx Nation vector borne disease control program
NVBDCP.pptx Nation vector borne disease control program
Sapna Thakur
 
Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...
Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...
Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...
Oleg Kshivets
 
Physiology of Chemical Sensation of smell.pdf
Physiology of Chemical Sensation of smell.pdfPhysiology of Chemical Sensation of smell.pdf
Physiology of Chemical Sensation of smell.pdf
MedicoseAcademics
 

Recently uploaded (20)

Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists
Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists  Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists
Ozempic: Preoperative Management of Patients on GLP-1 Receptor Agonists
 
263778731218 Abortion Clinic /Pills In Harare ,
263778731218 Abortion Clinic /Pills In Harare ,263778731218 Abortion Clinic /Pills In Harare ,
263778731218 Abortion Clinic /Pills In Harare ,
 
Gram Stain introduction, principle, Procedure
Gram Stain introduction, principle, ProcedureGram Stain introduction, principle, Procedure
Gram Stain introduction, principle, Procedure
 
Knee anatomy and clinical tests 2024.pdf
Knee anatomy and clinical tests 2024.pdfKnee anatomy and clinical tests 2024.pdf
Knee anatomy and clinical tests 2024.pdf
 
Basavarajeeyam - Ayurvedic heritage book of Andhra pradesh
Basavarajeeyam - Ayurvedic heritage book of Andhra pradeshBasavarajeeyam - Ayurvedic heritage book of Andhra pradesh
Basavarajeeyam - Ayurvedic heritage book of Andhra pradesh
 
Tom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness Journey
Tom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness JourneyTom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness Journey
Tom Selleck Health: A Comprehensive Look at the Iconic Actor’s Wellness Journey
 
planning for change nursing Management ppt
planning for change nursing Management pptplanning for change nursing Management ppt
planning for change nursing Management ppt
 
Flu Vaccine Alert in Bangalore Karnataka
Flu Vaccine Alert in Bangalore KarnatakaFlu Vaccine Alert in Bangalore Karnataka
Flu Vaccine Alert in Bangalore Karnataka
 
Pharma Pcd Franchise in Jharkhand - Yodley Lifesciences
Pharma Pcd Franchise in Jharkhand - Yodley LifesciencesPharma Pcd Franchise in Jharkhand - Yodley Lifesciences
Pharma Pcd Franchise in Jharkhand - Yodley Lifesciences
 
Evaluation of antidepressant activity of clitoris ternatea in animals
Evaluation of antidepressant activity of clitoris ternatea in animalsEvaluation of antidepressant activity of clitoris ternatea in animals
Evaluation of antidepressant activity of clitoris ternatea in animals
 
Ophthalmology Clinical Tests for OSCE exam
Ophthalmology Clinical Tests for OSCE examOphthalmology Clinical Tests for OSCE exam
Ophthalmology Clinical Tests for OSCE exam
 
Cervical & Brachial Plexus By Dr. RIG.pptx
Cervical & Brachial Plexus By Dr. RIG.pptxCervical & Brachial Plexus By Dr. RIG.pptx
Cervical & Brachial Plexus By Dr. RIG.pptx
 
Novas diretrizes da OMS para os cuidados perinatais de mais qualidade
Novas diretrizes da OMS para os cuidados perinatais de mais qualidadeNovas diretrizes da OMS para os cuidados perinatais de mais qualidade
Novas diretrizes da OMS para os cuidados perinatais de mais qualidade
 
ANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptx
ANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptxANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptx
ANATOMY AND PHYSIOLOGY OF URINARY SYSTEM.pptx
 
Hemodialysis: Chapter 3, Dialysis Water Unit - Dr.Gawad
Hemodialysis: Chapter 3, Dialysis Water Unit - Dr.GawadHemodialysis: Chapter 3, Dialysis Water Unit - Dr.Gawad
Hemodialysis: Chapter 3, Dialysis Water Unit - Dr.Gawad
 
Top 10 Best Ayurvedic Kidney Stone Syrups in India
Top 10 Best Ayurvedic Kidney Stone Syrups in IndiaTop 10 Best Ayurvedic Kidney Stone Syrups in India
Top 10 Best Ayurvedic Kidney Stone Syrups in India
 
heat stroke and heat exhaustion in children
heat stroke and heat exhaustion in childrenheat stroke and heat exhaustion in children
heat stroke and heat exhaustion in children
 
NVBDCP.pptx Nation vector borne disease control program
NVBDCP.pptx Nation vector borne disease control programNVBDCP.pptx Nation vector borne disease control program
NVBDCP.pptx Nation vector borne disease control program
 
Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...
Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...
Lung Cancer: Artificial Intelligence, Synergetics, Complex System Analysis, S...
 
Physiology of Chemical Sensation of smell.pdf
Physiology of Chemical Sensation of smell.pdfPhysiology of Chemical Sensation of smell.pdf
Physiology of Chemical Sensation of smell.pdf
 

16 [chapter 16 sensory, motor, and integrative systems]

  • 1. CHAPTER 16 Sensory, Motor, and Integrative Systems Principles of Anatomy and Physiology 14th Edition Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 2.  Sensation is the conscious or subconscious awareness of changes in the external or internal environment.  Perception is the conscious interpretation of sensations performed mainly by the cerebral cortex. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 3.  Each type of sensation is called a sensory modality. This includes: touch, pain, vision and hearing.  Sensory modalities are grouped into either general senses or special senses.  General senses: somatic-(tactile, thermal, proprioceptive); visceral- pressure, chemicals, stretch, nausea, hunger, temperature. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 4.  Special senses: smell, taste, vision, hearing and equilibrium.  Different types of sensory receptors are involved in the different senses. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 5. Free nerve endings are used to detect pain, temperature, tickle, itch and some touch. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 6. Encapsulated nerve endings are used to detect pressure, vibration and some touch. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 7. Sensory receptors for some special senses include gustatory receptor cells in taste buds, photoreceptors in the retina of the eye and hair cells in the inner ear for hearing. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 8. Anatomy Overview Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.  The Nervous System You must be connected to the Internet and in Slideshow Mode to run this animation.
  • 9.  Receptors may also be grouped based on location of the receptors and the origin of the stimuli that activate them.  Exteroreceptors include: hearing, vision, smell, taste, touch, pressure, vibration and pain. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 10.  Interoreceptors monitor the body’s internal environment.  Proprioceptors provide information about body position, muscle length and tension and the position and movement of joints. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 11. Sensation Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 12.  Somatic sensations include tactile, thermal, pain and proprioceptive.  Tactile sensations: touch, pressure, vibration, itch and tickle. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 13. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 14.  There are two types of pain: fast and slow. Fast pain (acute, sharp or pricking) perceived within 0.1 second.  Slow pain (chronic, burning, aching or throbbing) is perceived a second or more after the stimulus.  Superficial somatic pain: arising from skin receptors. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 15.  Deep somatic pain: skeletal muscles, joints, tendons and fascia.  Stimulation of pain sensors in visceral organs is visceral pain. This type of pain usually presents in or just deep to the skin that overlies the stimulated organ. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 16. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 17.  Proprioception: recognizing position of body parts.  Proprioceptors: in muscles and tendons. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 18.  Two types of proprioceptors are muscle spindles and tendon organs. Muscle spindles: in skeletal muscles monitor their length and are involved in stretch reflexes.  Tendon organs: at the junction of a tendon and a muscle protect muscles and tendons from damage due to overstretching.  Joint kinesthetic receptors exist within and around the joint capsule of synovial joints. They respond to pressure and acceleration and deceleration during movement. Joint ligaments contain receptors to protect against excessive strain. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 19. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 20. Somatic Sensations Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 21. Somatic sensory pathways carry information from somatic sensory receptors to the primary somatosensory area in the cerebral cortex and to the cerebellum. The pathways to the cortex consist of thousands of sets of three neurons classified as first, second and third-order neurons. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 22.  First-order neurons: impulses from somatic receptors to the brain stem or spinal cord.  Second-order neurons: impulses from the brain stem and spinal cord to the thalamus.  Third-order neurons: impulses from the thalamus to the primary somatosensory area of the cortex on the same side. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 23.  Somatic sensory impulses ascend to the cerebral cortex along three general pathways:  The posterior column-medial lemniscus pathway (impulses from the limbs, trunk, neck and posterior head). Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 24. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 25. The anterolateral (spinothalamic) pathway (impulses for pain, temperature, itch, tickle and posterior head). Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 26. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 27.  The trigeminothalamic pathway (impulses for most somatic sensations— tactile, thermal and pain—from the face, nasal cavity, oral cavity and teeth).  Somatic sensory impulses reach the cerebellum via the spinocerebellar tracts. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 28. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 29.  Postcentral gyri located on both parietal lobes of the brain are the sites for the primary somatosensory area.  Each region in this area receives sensory input from a different part of the body on the opposite side. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 30. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 31. The posterior spinocerebellar tract and the anterior spinocerebellar tract carry proprioceptive impulses to the cerebellum. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 32. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 33. Somatic Sensory Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 34. Nerves that extend out of the brain stem and spinal cord are called lower motor neurons (LMNs). These nerves innervate skeletal muscles of the face and head through cranial nerves, and skeletal muscles of the limb and trunk through spinal nerves. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 35. Somatic motor pathways provide the input into lower motor neurons and are divided into four distinct circuits. 1. Local circuit neurons are located close to LMNs in the brain stem and spinal cord. 2. Upper motor neurons (UMNs): input to both lower circuit neurons and LMNs. 3. Basal nuclei neurons: assist movement by providing input to UMNs. 4. Cerebellar neurons: assist movement via control of activity of UMNs. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 36. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 37.  UMNs extend to LMNs via two types of pathways:  Direct motor pathways deliver signals to LMNs from the cerebral cortex.  Indirect motor pathways deliver signals to LMNs from motor centers in the basal nuclei, cerebellum and cerebral cortex. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 38. There are two direct motor pathways used for voluntary movement. These are the two corticospinal pathways (the lateral corticospinal tract and the anterior corticospinal tract) and the corticobulbar pathway. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 39. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 40. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 41. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 42. Indirect motor pathways (extrapyramidal pathways): rubrospinal, tectospinal, vestibulospinal, lateral reticulospinal and medial reticulospinal tracts. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 43. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 44. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 45. The cerebellum performs 4 activities: 1. Monitoring intentions for movement 2. Monitoring actual movement 3. Comparing command signals with sensory information 4. Sending out corrective feedback Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 46. Somatic Motor Pathways Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 47.  Wakefulness and sleep: relies on the reticular activating system (RAS).  Learning and memory: includes immediate, short-term and long-term memory. Integrative Functions of the Cerebrum Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
  • 48. Copyright 2014 John Wiley & Sons, Inc. All rights reserved. Reproduction or translation of this work beyond that permitted in section 117 of the 1976 United States Copyright Act without express permission of the copyright owner is unlawful. Request for further information should be addressed to the Permission Department, John Wiley & Sons, Inc. The purchaser may make back-up copies for his/her own use only and not for distribution or resale. The Publisher assumes no responsibility for errors, omissions, or damages caused by the use of these programs or from the use of the information herein. End of Chapter 16 Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.

Editor's Notes

  1. <number>