The vestibular sensory organs of the inner ear responds to physical stimuli related to movement and orientation of the head in three-dimensional space. In response to mechanical forces acting on the inner ear, neural messages regarding head motion and position are generated by the vestibular apparatus and relayed to the brain. That information along with visual and proprioceptive input, is used by the central nervous system (CNS) to maintain clear vision during head movement, to control muscles responsible for upright posture and to provide a sense of orientation of the body with respect to the surrounding environment. Although the vestibular system is one of our major sensory modalities, it differs somewhat from other senses such as vision and hearing in that it operates largely in the service of motor reflexes, outside the field of conscious perception. Thus we are not aware of vestibular sensory input unless the system is subjected to unusually high levels of stimulation or is compromised by disease, in which importance of vestibular function becomes acutely obvious. Some fundamental aspects of vestibular anatomy and physiology, emphasis on features of the system that provide the foundation for clinical testing of vestibular function.