photoperiodism its discovery,significance,classifications,mechanism,critical day length,quality of light, night break phenomenon,phytochrome.florigen,floering genes, circadian rhythm
molecular and genetic analysis of floral induction is an integrated approach, taking into consideration various genes involved in the four major pathways of flowering process
photoperiodism its discovery,significance,classifications,mechanism,critical day length,quality of light, night break phenomenon,phytochrome.florigen,floering genes, circadian rhythm
molecular and genetic analysis of floral induction is an integrated approach, taking into consideration various genes involved in the four major pathways of flowering process
Everything about photoperiodism from scratch to smart, from the oldest models to the latest models as well as proposed one, exclusive and elusive illustrations and models for proper understanding
Self-incompatibility refers to the inability of a plant with functional pollen to set seeds when self pollinated. It is the failure of pollen from a flower to fertilize the same flower or other flowers of the same plant.
This presentation includes, Single-locus self-incompatibility- {Gametophytic self-incompatibility (GSI) and Sporophytic self-incompatibility (SSI)},2-locus gametophytic self-incompatibility, Heteromorphic self-incompatibility,Cryptic self-incompatibility (CSI) and Late-acting self-incompatibility (LSI).
flowering is perhaps the most important physiological phenomenon in the life-cycle of higher plants. it is a resultant of a range of internal and external factors, that leads to the activity of a plethora of genes, that leads to the development of flowers
Everything about photoperiodism from scratch to smart, from the oldest models to the latest models as well as proposed one, exclusive and elusive illustrations and models for proper understanding
Self-incompatibility refers to the inability of a plant with functional pollen to set seeds when self pollinated. It is the failure of pollen from a flower to fertilize the same flower or other flowers of the same plant.
This presentation includes, Single-locus self-incompatibility- {Gametophytic self-incompatibility (GSI) and Sporophytic self-incompatibility (SSI)},2-locus gametophytic self-incompatibility, Heteromorphic self-incompatibility,Cryptic self-incompatibility (CSI) and Late-acting self-incompatibility (LSI).
flowering is perhaps the most important physiological phenomenon in the life-cycle of higher plants. it is a resultant of a range of internal and external factors, that leads to the activity of a plethora of genes, that leads to the development of flowers
Annual plants is influenced by temperature for flowering is secondary to that of light.
Biennials and perennials remain vegetative during first growing season
After prolonged exposure to cold temperature of winter flowering occurs during following season.
Without cold exposure majority plants remain vegetative
Introduction:
Vernalization is the process whereby flowering is promoted by a cold treatment given to a fully hydrated seed or to a growing plant.
Dry seeds do not respond to the cold treatment.
Due to vernalization, the vegetative period of the plant is cut short resulting in an early flowering.
Also called yarovization.
Without the cold treatment, plants that require vernalization show delayed flowering or remain vegetative.
In many cases, these plants grow as rosettes with no elongation of the stem.
History:
Klippart,1857- first noticed the low-temperature requirement for flowering while working with winter wheat and spring wheat.
Lysenko,1938-used the term vernalization for a low-temperature promotion of flowering in plants.
Chourad ,1960- defined vernaliZation as “acquisition or acceleration of the ability to flower by a chilling treatment”.
Vernalization
For vernalization the seeds are allowed to germinate for some time and then are given cold treatment from 0 ̊C to 5 ̊C.
The period of cold treatment, varies from few days to many weeks.
After the cold treatment the seedlings are allowed to dry for some time and then sown.
Vernalization prepares the plant for flowering.
The cold stimulus usually perceived by the apical meristems. But in some species, all dividing cells of roots and leaves may be the potential sites of vernalization eg. Leennario biennis.
Vernalization induces the plant to produce a hormone called vernalin. It was discovered by Melcher(1936).
The vernalization stimulus can be transmitted from one plant to another through grafting.
The age of the plant is an important factor in determining the responsiveness of the plant to the cold stimulus and it differs in different species.
The suitable temperatures for vernalization ranges between 1 to 6 ̊c.
At higher temperature from 7 ̊c onwards response of the plant is decreased.
A temperature of about 12 to 14 ̊c is most ineffective in vernalizing the plant.
The vernalization is an aerobic process and requires metabolic energy.
In the absence of oxygen cold treatment becomes completely inefficient.
A sufficient amount of water is also essential.
Vernalization of dry seeds is not possible.
Factors affecting Vernalization:
Site of vernalization
Age of plants
Appropriate low temperature
Duration of exposure
Mechanism of vernalization:
Two theories..
1. Phasic development theory
2. Hormonal theories.
Epigenetic Changes in Gene Expression:
Vernalization May Involve Epigenetic Changes in Gene Expression.
Changes in gene expression that are stable even after the signal that induced the change (in this case cold) is removed are known as epigenetic regulation.
One model for how vernalization affects flowering is that there are stable changes in the pattern of gene expression in the meristem after cold treatment.
The involvement of epigenetic regulation in the vernalization process has been confirmed in the LDP Arabidopsis.
Photoperiodism is the phenomenon of physiological changes that occur in plants in
response to relative length of day and night (i.e. photoperiod). The response of the plants to
the photoperiod, expressed in the form of flowering is also called as photoperiodism. The
phenomenon of photoperiodism was first discovered by Garner and Allard (1920).Depending
upon the duration of photoperiod, the plants are classified into three categories.
1. Short day plants (SDP)
2. Long day plants (LDP)
3. Day neutral plants (DNP)
Vernalization is the induction of a plant's flowering process by exposure to the prolonged cold of winter, or by an artificial equivalent. After vernalization, plants have acquired the ability to flower, but they may require additional seasonal cues or weeks of growth before they will actually flower.The vernalization requirement ensures that plants do not flower in the fall when the environmental conditions are unfavorable for reproduction. The strength of the vernalization requirement can vary within plant species.
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June 3, 2024 Anti-Semitism Letter Sent to MIT President Kornbluth and MIT Cor...Levi Shapiro
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The US House of Representatives is deeply concerned by ongoing and pervasive acts of antisemitic
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This Congress will not stand idly by and allow an environment hostile to Jewish students to persist. The House believes that your institution is in violation of Title VI of the Civil Rights Act, and the inability or
unwillingness to rectify this violation through action requires accountability.
Postsecondary education is a unique opportunity for students to learn and have their ideas and beliefs challenged. However, universities receiving hundreds of millions of federal funds annually have denied
students that opportunity and have been hijacked to become venues for the promotion of terrorism, antisemitic harassment and intimidation, unlawful encampments, and in some cases, assaults and riots.
The House of Representatives will not countenance the use of federal funds to indoctrinate students into hateful, antisemitic, anti-American supporters of terrorism. Investigations into campus antisemitism by the Committee on Education and the Workforce and the Committee on Ways and Means have been expanded into a Congress-wide probe across all relevant jurisdictions to address this national crisis. The undersigned Committees will conduct oversight into the use of federal funds at MIT and its learning environment under authorities granted to each Committee.
• The Committee on Education and the Workforce has been investigating your institution since December 7, 2023. The Committee has broad jurisdiction over postsecondary education, including its compliance with Title VI of the Civil Rights Act, campus safety concerns over disruptions to the learning environment, and the awarding of federal student aid under the Higher Education Act.
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http://sandymillin.wordpress.com/iateflwebinar2024
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Knowledge and skills frameworks, generally called competency frameworks, for ELT teachers, trainers and managers have existed for a few years now. However, until I created one for my MA dissertation, there wasn’t one drawing together what we need to know and do to be able to effectively produce language learning materials.
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2. Content
• Introduction
• Technique of vernalization
• Vernalization and flowering
• Mechanism of vernalization
•Devernalization
•Importance of vernalization
3. Introduction
• in many plants the flowering is influenced not only by the
correct photoperiod but also by temperature.
• In annuals the flowering is primarily affected by the
photoperiod.
• The effect of temperature is secondary to light,a biennial
plant,on the other hand,grows only vegetatively during the
first season and will not initiate.
• Flowering until it has been exposed to prolonged period of
low temperature of the winter.
• The term vernalization was coined by T.D.Lysenko in 1920s.
• Other important scientists who have contributed to the
study on vernalization were F.G.Gregory and O.N.Purvis
(1961).
4. • It allows the plant to reach vegetative maturity
before reproduction can occur.
• The condition occurs in winter varieties of some
annual food plants (e.g., Wheat, Barley, and
Rye).
• some biennial (e.g., Cabbage, Sugar beet,
Carrot) and perennial plants (e.g.,
Chrysanthemum).
• The annual winter plants also possess spring
varieties. The spring varieties are planted in
spring.
• They come to flower and bear fruits prior to end
of growing season.
5. • They germinate and over winter as small
seedlings , resume growth in spring , and
harvested usually about mid summer.
• The over wintering cold treatment , or
vernalization , renders the plants sensitive to
long day.
• If seeds of the winter strain are sown in the fall ,
however, the germinated seedlings are
subjected to an over wintering low temperature
treatment.
Cont…
6.
7. • The effect of temperature is secondary to light.
A biennial plant, on the other hand, grows only Vegetatively
during the first season and will not initiate flowering until it
has been exposed to prolonged periods of low temperature
of the winter.
The biennials like cabbages, kales, carrots, foxglove and
beets need absolute cold requirement for flowering.
8. • Technique of vernalization:
Although differing in details, the technique of vernalization
in esentially very simple.
The seed is first soaked in water and is allowed to
germinate.
The seedlings resume growth in spring.
They bear flowers and fruits in summer.
9. • It was found by Lysenko (1928), a Russian worker
that the cold requiring annual and biennial plants
can be made to flower in one growing season by
providing low temperature treatment to young
plants or moistened seeds.
• He called the effect of this chilling treatment as
vernalization.
• Vernalization is, therefore, a process of
shortening of the juvenile or vegetative phase
and hastening flowering by a previous cold
treatment.
Cont…
10.
11. A similar phenomenon is to be seen in certain varieties of
cereals.
In cold countries, there are cereals of two physiological
kinds –the winter cereals and the spring cereals.
The winter variety is sown in early autumn i.e. in the
month of September or October to make them flower in
the following summer.
Cont…
12. If they are sown in the spring along with the spring variety,
they grow vegetatively but fail to produce ears or flowers.
Exposure to low temperature of the winter is evidently
essential for the flowering of winter varieties.
It has been shown by several worker that this requirement
of low temperature in nature can be satisfied artifically in
laboratories in the absence of the winter season and the
plant may be made to flower in summer season.
Cont…
13. For example, If a biennial seed is germinated and is then
exposed to low temperature ( 0 – 5 degree C ) for few
weeks, it will behave as if it has gone through the cold
winter after a year of growth.
Semilar treatment will enable the winter variety of wheat
or rye to flower even if they are shown in the spring.
Cont…
14. Vernalization and flowering:
In nature, vernalization takes place in the seed stage in the
annuals.
Biennials and Perennials are however vernalized at a much
later stage.
In Biennials, vegetative growth takes place in the first
season, and the vernalization takes place in the following
winter.
15. • The site of vernalization, in the case of biennials and
perennials,is believed to be the growing point(apical
bud).
• According to chailakhyan (1968)there are two types of
chemicals responsible for flowering: Gibberellins and
anthesin.
• he assumed that the cold treatment resulted in the
production of vernalin.
• Gibberellins:if the cold-treated plants are then sown
in long days the vernalin is converted into……..
• Anthesin:during grafting the vernalised one produces
vernalin while the non-vernalized one produced……
• Then flowering is believed to occur due to cobbined
effect of both.
16. The flowering stimulus resulting from appropriate vernalization of the
apex may pass from one plant to a grafted unvernalized partner.
17.
18.
19.
20.
21.
22.
23. Importance of vernalization
(i) Vernalization can help in shortening the juvenile
or vegetative period of plant and bring about
early flowering. It is not only applicable to
temperate plants but also to some tropical
plants, e.g., Wheat, Rice, Millets, Cotton,
(ii) It increases yield, resistance to cold and
diseases, and
(iii) Kernel wrinkles of Triticale can be removed by
vernalization.
24. Reference :
Text book of Plant physiology:
- V.Verma
funadamentals and applications of Plant
physiology
– Arvind kumar/ S.S Purohit
Plant physiology
- Taiz and Zeiger
www.vernalization image.in