Thomas Morgan observed that genes on the same chromosome can be linked during meiosis. He conducted test crosses on Drosophila with traits for eye color and wing type. The results showed that some offspring received combinations of traits that were not expected if the genes assorted independently, demonstrating genetic linkage. Genetic linkage occurs because homologous chromosomes occasionally exchange sections during meiosis through the process of crossover. Crossover results in new combinations of genes that can become separated from their original linked combinations on the chromosome. The frequency of crossover between genes determines how far apart they will be on the genetic map of the chromosome.