The nitrogen cycle is the biogeochemical cycle by which nitrogen is converted into various chemical forms as it circulates among the atmosphere and terrestrial and marine ecosystems. The conversion of nitrogen can be carried out through both biological and physical processes. Important processes in the nitrogen cycle include fixation, ammonification, nitrification, and denitrification. The majority of Earth's atmosphere (78%) is nitrogen, making it the largest pool of nitrogen. However, atmospheric nitrogen has limited availability for biological use, leading to a scarcity of usable nitrogen in many types of ecosystems. The nitrogen cycle is of particular interest to ecologists because nitrogen availability can affect the rate of key ecosystem processes, including primary production and decomposition. Human activities such as fossil fuel combustion, use of artificial nitrogen fertilizers, and release of nitrogen in wastewater have dramatically altered the global nitrogen cycle.
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The nitrogen cycle is the biogeochemical cycle by which nitrogen is converted into various chemical forms as it circulates among the atmosphere and terrestrial and marine ecosystems. The conversion of nitrogen can be carried out through both biological and physical processes. Important processes in the nitrogen cycle include fixation, ammonification, nitrification, and denitrification. The majority of Earth's atmosphere (78%) is nitrogen, making it the largest pool of nitrogen. However, atmospheric nitrogen has limited availability for biological use, leading to a scarcity of usable nitrogen in many types of ecosystems. The nitrogen cycle is of particular interest to ecologists because nitrogen availability can affect the rate of key ecosystem processes, including primary production and decomposition. Human activities such as fossil fuel combustion, use of artificial nitrogen fertilizers, and release of nitrogen in wastewater have dramatically altered the global nitrogen cycle.
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Nitrogen Fixation and the Nitrogen CycleIn a symbiotic relatio.docxhenrymartin15260
Nitrogen Fixation and the Nitrogen Cycle
In a symbiotic relationship with the soil bacteria known as 'rhizobia', legumes form nodules on their roots (or stems, see figure below) to 'fix' nitrogen into a form usable by plants (and animals). The process of biological nitrogen fixation was discovered by the Dutch microbiologist Martinus Beijerinck. Rhizobia (e.g., Rhizobium, Mesorhizobium, Sinorhizobium) fix atmospheric nitrogen or dinitrogen, N2, into inorganic nitrogen compounds, such as ammonium, NH4+, which is then incorporated into amino acids, which can be utilized by the plant. Plants cannot fix nitrogen on their own, but need it in one form or another to make amino acids and proteins. Because legumes form nodules with rhizobia, they have high levels of nitrogen available to them. Their abundance of nitrogen is beneficial not only to the legumes themselves, but also to the plants around them. There are other sources of nitrogen in the soil, but are not always provided at the levels required by plants, making the symbiotic relationship between legumes and rhizobia highly beneficial. In return for the fixed nitrogen that they provide, the rhizobia are provided shelter inside of the plant's nodules and some of the carbon substrates and micronutrients that they need to generate energy and key metabolites for the cellular processes that sustain life (Sprent, 2001). Nodulation and nitrogen fixation by rhizobia is not exclusive to legumes; rhizobia form root nodules on Parasponis Miq., a genus of five species in the Ulmaceae (see 'Rosales').
The picture on the left shows typical root nodules, these from bur clover (Medicago). The picture on the right shows "stem" nodules on Sesbania rostrata - stem nodules are produced from lateral or adventitious roots and are typically found in those few water-tolerant legume groups (Neptunia, Sesbania) that prefer wet or water-logged soils (Goormachtig et al., 2004).
The nitrogen cycle (shown below) describes the series of processes by which the element nitrogen, which makes up about 78% of the Earth’s atmosphere, cycles between the atmosphere and the biosphere. Plants, bacteria, animals, and manmade and natural phenomena all play a role in the nitrogen cycle. The fixation of nitrogen, in which the gaseous form dinitrogen, N2) is converted into forms usable by living organisms, occurs as a consequence of atmospheric processes such as lightning, but most fixation is carried out by free-living and symbiotic bacteria. Plants and bacteria participate in symbiosis such as the one between legumes and rhizobia or contribute through decomposition and other soil reactions. Bacteria like Rhizobium, or the actinomycete Frankia which nodulates members of the plant families Rosaceae and Betulaceae, utilize atmospheric nitrogen and convert it to an inorganic form (usually ammonium, NH4+) that plants can use. The plants then use the fixed nitrogen to produce vital cellular products such as proteins. The plants are then.
Nitrogen is important element of life. In importance it comes only next to carbon, hydrogen, and oxygen. The composition of protein, nucleic acid, growth hormones, and vitamins requires Nitrogen. Leaves consist of about 1 to 15% nitrogen of their dry weight but lesser % in another vegetative organ.
• The N2 is present in the atmosphere, in the form of gas. It is about 78%.
• Green plants unable to use this N2 directly in their metabolism. Only some micro-organism can convert this N2 gas directly into organic form.
• The N2 present in the soil is called soil nitrogen. The plants growing in the soil, mainly utilize the soil N2 for their metabolic requirements.
• In the soil the nitrogen is present in the form of nitrate nitrogen (NO3, NO2), ammonia nitrogen (ammonia, ammonium salt), organic nitrogen and molecular nitrogen (N2).
• The converging of the free nitrogen, by natural or physical process is called nitrogen fixation… when any biological system is involved in this process, then it is called as biological nitrogen fixation……
The Nitrogen cycle is defined as the biogeochemical cycle process that involves transforming the inert nitrogen that is available in the atmosphere, into a more usable or conventional form, that can be actively used by plants, and various living organisms. Enroll now at Tutoroot.
This is a comprehensive account of the nitrogen cycle in terrestrial environments. The nitrogen cycle is responsible for the circulation of nitrogen between inorganic and organic components of the environment.
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1. The circulation of nitrogen in nature, consisting
of a cycle of chemical reactions in which
nitrogen from the atmosphere is fixed in
Compounds in soil or water.
Assimilated by plants and animals.
Released to the soil and water
through decomposition
Returned to the
atmosphere through Denitrification
2. Nitrogen cycle
The continuous process by which nitrogen is excha
nged between organisms and the environment
Some of the atmosphere's free nitrogen combines
with other elements to form compounds that are
deposited in the soil.
These are then converted by bacteria, in process
called
nitrification, into nutrients that are absorbed by
the roots of green plants.
3. Nitrogen is then passed into the food
chain and returned to the soil by metabolism a
nd decay of plants and animals.
4.
5. The growth of all organisms depends on the
availability of mineral nutrients, and none is more
important than nitrogen, which is required in large
amounts as an essential component of proteins, nucleic
acids and other cellular constituents.
There is an abundant supply of nitrogen in the earth's
atmosphere - nearly 78% in the form of N2 gas.
However, N2 is unavailable for use by most organisms
because there is a triple bond between the two nitrogen
atoms, making the molecule almost inert.
In order for nitrogen to be used for growth it must be
"fixed" (combined) in the form of ammonium (NH4) or
nitrate (NO3).
6. Nitrogen-fixing bacteria, microorganisms
capable of transforming atmospheric nitrogen
into fixed nitrogen (inorganic compounds
usable by plants). More than 90 percent of all
nitrogen fixation is effected by these organisms,
which thus play an important role in
the nitrogen cycle.
7. Two kinds of nitrogen-fixing bacteria are
recognized.
The first kind, the free-living (nonsymbiotic)
bacteria, includes the cyanobacteria(or blue-
green algae) Anabaena and Nostoc and genera
such as Azotobacter and Clostridium.
The second kind comprises the mutualistic
(symbiotic) bacteria; examples
include Rhizobium, associated with leguminous
plants (e.g., various members of the pea family.
8. The symbiotic nitrogen-fixing bacteria present in
the root hairs of host plants, where they multiply
and stimulate formation of root nodules,
enlargements of plant cells and bacteria in intimate
association. Within the nodules the bacteria
convert free nitrogen to ammonia which the host
plant utilizes for its development.
Legumes (e.g., alfalfa, beans, peas, soybeans),
seeds are usually inoculated with commercial
cultures of appropriate Rhizobium species,
especially in soils poor or lacking in the required
bacterium.
9.
10.
11. The term nitrification refers to the conversion of
ammonium to nitrate (pathway 3-4). This is brought
about by the nitrifying bacteria, which are specialized
to gain their energy by oxidizing ammonium, while
using CO2 as their source of carbon to synthesize
organic compounds
The nitrifying bacteria are found in moist soils and
waters of moderate pH, but are not active in highly
acidic soils. They almost always are found as mixed-
species communities e.g. Nitrosomonasspecies - are
specialized to convert ammonium to nitrite (NO2
-)
while others - e.g. Nitrobacter species - convert nitrite to
nitrate (NO3
-
12. Denitrification refers to the process in which nitrate is
converted to gaseous compounds (nitric oxide, nitrous
oxide and N2) by microorganisms. The sequence
usually involves the production of nitrite (NO2
-)
.Several types of bacteria perform this conversion when
growing on organic matter in anaerobic conditions.
Because of the lack of oxygen for normal aerobic
respiration, they use nitrate in place of oxygen as the
terminal electron acceptor.
In the absence of oxygen, any reducible substance such
as nitrate (NO3
-) could serve the same role and be
reduced to nitrite, nitric oxide, nitrous oxide or N2.