Swim bladder general organization and circulation and function
1. SCHOOL OF STUDIES IN ZOOLOGY
JIWAJI UNIVERSITY, GWALIOR
Dr. Ramkumar Lodhi
Conservation Biology Lab
TOPIC- SWIM BLADDER GENERAL ORGANIZATION AND CIRCULATION
AND FUNCTION
2. SYNOPSIS
Introduction
General organization and circulation
Types of air bladder
Gas secreting complex
Functions of the swim bladder
Conclusion
Reference
3. INTRODUCTION
Fishes possess various structural adaptations in
their body.
Swim bladder, also known as air bladder or gas
bladder, is a characteristic structure in all the
osteichthyes, excepting a few species in which it
has been lost secondarily.
It is situated between the alimentary canal and
kidneys, and is sac- like in appearance.
It contains air or gas, which is a mixture of
carbon dioxide, oxygen and nitrogen, and
develops as a small outgrowth from the dorsal or
lateral wall of the gut.
4. CIRCULATION
Blood supply of the air bladder-:
The air bladder is supplied with blood either directly
from the dorsal aorta or from branches of the coeliaco-
mesentric artery.
The venous blood is drained into the hepatic portal
system or into the posterior cardinal vein.
The gas bladder does not present a uniform pattern
and degree of vascularity in a various species.
In species belonging to the clupeidae and
salmonidae, there is a uniform distribution of blood
capillaries over the bladder wall, and red bodies or red
glands are not form.
5.
6. TYPES OF SWIM BLADDER
Depending on the presence of the duct (ductus
pneumaticus) between the swim-bladder and the
oesophagus, the swim-bladder in fishes can be
divided into two broad categories: Physostomus
and Physoclistous types-:
Physostomous Condition:
The swim-bladder develops from the
oesophagus. When the ductus pneumaticus is
present between the swim-bladder and the
oesophagus, the swim-bladder is called
physostomous type.
7. Physoclistous condition:
In this condition the ductus pneumaticus is either closed
or atrophied. This type of swim- bladder is observed in
spiny-rayed fishes.
8. GAS SECRETING COMPLEX
In all the fishes (physostomi and physoclisti) the
anterior part of the bladder is specialised for gas
secretion.
In physoclistic species, absorption of gas takes place in
the posterior region of the bladder.
In more specialised species as the mugil, gadus and
balistes, the posterior part of the bladder is modified
into a ‘oval’ for gas absorption.
Its opening is guarded by sphincter muscles.
A small part of the anterior region becomes specialised
for secreting gas, and is called the ‘red body’ or ‘red
gland’.
9. FUNCTIONS OF SWIM BLADDER
The swim bladder performs anyone or more of
the following functions:
Respiratory function
In a number of primitive teleostomes such as
polypterus, lepidosteus and amia, the gas
bladder has a lung-like structure and is highly
vascular.
In the dipnoi the bladder resembles closely with
the lung of amphibians.
In some species as perca fluviatilis, cyprinus, the
swim bladder is used to store oxygen; which is
available to the fish during emergancy.
10. This function is more important in deep sea
fishes in which the amount of oxygen stored is
pretty high.
Sound production
Swim bladder is an important organ for
producing sound in fishes.
It acts as a resonator for the sound produced by
other organs, as in Ballistidae and Triglidae.
In some species as the sciaenidae and in the
toadfish, sound is produced by the vibration of
the bladder wall, caused by the action of
muscles attached to it.
11. Auditory function
Air bladder is connected with the membranous
labyrinth in several species either directly or
through weberian ossicles.
In these fishes, the bladder serves to transmit
sound waves to the ear more efficiently than in
other.
Sensory function
The swim bladder works as a pressure receptor
like a manometer or a barometer.
When the fish moves from the depth to another,
change in pressure causes compression or
rarefaction of gas in the bladder, leading to the
deformation of its wall.
12. As a hydrostatic organ
The gas bladder is an important hydrostatic
organ and makes the fish weightless enabling
it to maintain itself in equilibrium with very
little effort and without loss of energy.
The swim bladder acts as a float for the fish.
This explains why bottom living species have
lost the bladder.
The volume of the bladder is about 5% of the
total volume of the fish in marine species,
and is about 7-10% in fresh water forms.
13. CONCLUSION
The air-bladder in fishes varies greatly in
structure, shape and size. It is essentially
a tough sac-like structure with an overlying
capillary network.
The air-bladder may be composed of one
or two or many chambers.
It serves as an important organ as it helps
in respiration, buoyancy, sound
production, audition and sensing.