Polypodiophyta are vascular plants that reproduce via spores and range in size from less than 1 cm to 25 m tall tree ferns. Their leaves, called fronds, are megaphylls that are typically divided into smaller segments and require external water for reproduction. Spores are released and grow into small, flat gametophytes that bear archegonia and antheridia for sexual reproduction. A zygote develops into an independent sporophyte as the gametophyte dies off. Ferns were abundant as tree ferns during the Carboniferous period 320-250 million years ago. Some modern uses include using fern fronds in floral arrangements.