Sexual selection leads to the evolution of traits that enhance mating success but do not necessarily help survival. These traits can include elaborate ornamentation that requires significant energy or reduces mobility. Darwin proposed sexual selection to explain the evolution of such extravagant but potentially detrimental traits seen in male peacocks and other species. Sexual selection operates through either intersexual selection, where female mate choice shapes male traits, or intrasexual selection, such as male-male combat over mates. Elaborate male traits may have evolved because they honestly signal genetic quality to females, since only males in good condition can afford the costs of such ornamentation.