1. Systematics is the study of the diversity and relationships among organisms, both living and extinct. It involves classifying organisms and determining their evolutionary relationships through tools like cladograms and phylogenetic trees.
2. Taxonomy is the practice of naming and grouping organisms, which is a key part of systematics. However, systematics also examines broader aspects of evolution through fields like genetics and molecular biology.
3. Systematics has many important applications, including aiding agriculture and forestry by identifying pests, enabling biological control of pests by introducing natural predators, and assisting in public health by identifying disease vectors.
1.Definition and basic concepts of Biosystematics, , Historical perspectives of Biosystematics and Taxonomy, Stages of taxonomic procedures-alpha taxonomy, Beta taxonomy and Gamma taxonomy,
Neo taxonomy.
Classical and molecular taxonomic parameters, species concept, systematic gradation of animals, nomenclature, modern scheme of animal classification into sub-Kingdom, division, section, phyla and minor phyla
1.Definition and basic concepts of Biosystematics, , Historical perspectives of Biosystematics and Taxonomy, Stages of taxonomic procedures-alpha taxonomy, Beta taxonomy and Gamma taxonomy,
Neo taxonomy.
Classical and molecular taxonomic parameters, species concept, systematic gradation of animals, nomenclature, modern scheme of animal classification into sub-Kingdom, division, section, phyla and minor phyla
Iczn(The International Commission on Zoological Nomenclature )Al Nahian Avro
The International Commission on Zoological Nomenclature (ICZN) acts as adviser and arbiter for the zoological community by generating and disseminating information on the correct use of the scientific names of animals. The ICZN is responsible for producing the International Code of Zoological Nomenclature - a set of rules for the naming of animals and the resolution of nomenclatural problems.
Kinds of taxonomic publications,taxonomic review ,revision, monograph,atlas,s...Anand P P
kinds of taxonomic publication mainly deals with different types of taxonomic publications.the taxonomy deals with several types of publications mainly that help to over all exchange of taxonomic information,its is a world wide taxonomic communication.
One major challenge is the time consumed by the interplay between the taxonomist and the publisher in preparing taxonomic data and going to print. Breaking this bottleneck requires seamless integration between compilation of the descriptive taxonomic data and the publication upon which the data are based
Taxonomic Collections, Preservation and Curating of InsectsKamlesh Patel
Taxonomy: Taxonomy is the science of defining and naming groups of biological organisms on the basis of shared characteristics.
The classification of organisms is according to hierarchal system or in taxonomic ranks (eg; domain, kingdom, phylum class, order, family, genus and species) based on phylogenetic relationship established by genetic analysis.
Taxonomic Collection : Biological collection are typically preserved plant or animals specimens along with specimen documentations such as labels and notations.
Dry Collection - Dry collections consist of those specimens that are preserved in a dry state.
Wet Collection - Wet collections are specimens kept in a liquid preservative to prevent their deterioration.
Iczn(The International Commission on Zoological Nomenclature )Al Nahian Avro
The International Commission on Zoological Nomenclature (ICZN) acts as adviser and arbiter for the zoological community by generating and disseminating information on the correct use of the scientific names of animals. The ICZN is responsible for producing the International Code of Zoological Nomenclature - a set of rules for the naming of animals and the resolution of nomenclatural problems.
Kinds of taxonomic publications,taxonomic review ,revision, monograph,atlas,s...Anand P P
kinds of taxonomic publication mainly deals with different types of taxonomic publications.the taxonomy deals with several types of publications mainly that help to over all exchange of taxonomic information,its is a world wide taxonomic communication.
One major challenge is the time consumed by the interplay between the taxonomist and the publisher in preparing taxonomic data and going to print. Breaking this bottleneck requires seamless integration between compilation of the descriptive taxonomic data and the publication upon which the data are based
Taxonomic Collections, Preservation and Curating of InsectsKamlesh Patel
Taxonomy: Taxonomy is the science of defining and naming groups of biological organisms on the basis of shared characteristics.
The classification of organisms is according to hierarchal system or in taxonomic ranks (eg; domain, kingdom, phylum class, order, family, genus and species) based on phylogenetic relationship established by genetic analysis.
Taxonomic Collection : Biological collection are typically preserved plant or animals specimens along with specimen documentations such as labels and notations.
Dry Collection - Dry collections consist of those specimens that are preserved in a dry state.
Wet Collection - Wet collections are specimens kept in a liquid preservative to prevent their deterioration.
Contribution to Systematic biology. KUSTKashif Obaid
Its all about general taxonomic characters and about the contribution to systematic biology...
Be ready dear KUSTIANS....
follow me and comment below in comment box to encourage me. thanks!
This lesson discusses Biodiversity and Evolution
define biodiversity and evolution;
cite the contributions of Charles Darwin to the theory of evolution;
account for the evidence of evolution;
explain how biodiversity and evolution affect life;
demonstrate how biodiversity and evolution help an ecosystem to function;
explain the role of natural selection in the evolutionary process; and
relate evolution and speciation.
define what an ecosystem is;
identify the components of ecological structures in an ecosystem;
explain how diversity contributes to stability and survival;
cite examples of what helps and what disrupts the interaction in an ecosystem;
analyze how the human population affects the different ecosystems; and
apply the knowledge of biodiversity in the maintenance of an ecosystem and vice versa.
BOTANY IS THE SCIENCE THAT DEAL WITH THE LIVING THING (PLANT), IN THE PHARMACY , THIS SCIENCES IS SO SPECIAL AS IT IS HELPFUL IN IDENTIFICATION OF PLANT WITH PHARMACOLOGICAL EFFECT TO HUMAN , APPLIED IN TRADTIONAL MEDICINES
Production of live food (Aquatic micro animals)for the rearing of fish fry at...Hafiz M Waseem
Production of live food (Aquatic micro animals)for the rearing of fish fry at hatcheries ROTIFERSBrachionus sp. CRUSTACEANS (CLADOCERANS)Moina sp. Daphnia sp.
green water production at fish hatcheries and its uses to enhance primary pro...Hafiz M Waseem
green water production at fish hatcheries and its uses to enhance primary productivity.ppt
Chlorella sp.
Scenedesmus sp.
Tetraselmis chuii
Skeletonemia sp
Spirulina sp.
Chaetoceros sp.
Nitzschia sp.
Determination of p h of waste water sample .....................................Hafiz M Waseem
ecologyDetermination of pH of Waste Water Sample ..................................................... 4
Determination Dissolved Oxygen within Water ................................................... 5
Adaptive Features of Animals in Relation to Food and Environment .................. 7
Study the Plant Population Density ................................................................... 10
Experimental Design and Approaches to Ecological Research ........................ 12
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
Trophic levels and energy variation with increasing trophic levels.food chain...Hafiz M Waseem
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
I AM HAFIZ MUHAMMAD WASEEM from mailsi vehari
BSc from science college Multan
MSC university of education Lahore
i love Pakistan and my teachers and my parents
Unit 8 - Information and Communication Technology (Paper I).pdfThiyagu K
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The Indian economy is classified into different sectors to simplify the analysis and understanding of economic activities. For Class 10, it's essential to grasp the sectors of the Indian economy, understand their characteristics, and recognize their importance. This guide will provide detailed notes on the Sectors of the Indian Economy Class 10, using specific long-tail keywords to enhance comprehension.
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The people of Punjab felt alienated from main stream due to denial of their just demands during a long democratic struggle since independence. As it happen all over the word, it led to militant struggle with great loss of lives of military, police and civilian personnel. Killing of Indira Gandhi and massacre of innocent Sikhs in Delhi and other India cities was also associated with this movement.
2024.06.01 Introducing a competency framework for languag learning materials ...Sandy Millin
http://sandymillin.wordpress.com/iateflwebinar2024
Published classroom materials form the basis of syllabuses, drive teacher professional development, and have a potentially huge influence on learners, teachers and education systems. All teachers also create their own materials, whether a few sentences on a blackboard, a highly-structured fully-realised online course, or anything in between. Despite this, the knowledge and skills needed to create effective language learning materials are rarely part of teacher training, and are mostly learnt by trial and error.
Knowledge and skills frameworks, generally called competency frameworks, for ELT teachers, trainers and managers have existed for a few years now. However, until I created one for my MA dissertation, there wasn’t one drawing together what we need to know and do to be able to effectively produce language learning materials.
This webinar will introduce you to my framework, highlighting the key competencies I identified from my research. It will also show how anybody involved in language teaching (any language, not just English!), teacher training, managing schools or developing language learning materials can benefit from using the framework.
We all have good and bad thoughts from time to time and situation to situation. We are bombarded daily with spiraling thoughts(both negative and positive) creating all-consuming feel , making us difficult to manage with associated suffering. Good thoughts are like our Mob Signal (Positive thought) amidst noise(negative thought) in the atmosphere. Negative thoughts like noise outweigh positive thoughts. These thoughts often create unwanted confusion, trouble, stress and frustration in our mind as well as chaos in our physical world. Negative thoughts are also known as “distorted thinking”.
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The Roman Empire A Historical Colossus.pdfkaushalkr1407
The Roman Empire, a vast and enduring power, stands as one of history's most remarkable civilizations, leaving an indelible imprint on the world. It emerged from the Roman Republic, transitioning into an imperial powerhouse under the leadership of Augustus Caesar in 27 BCE. This transformation marked the beginning of an era defined by unprecedented territorial expansion, architectural marvels, and profound cultural influence.
The empire's roots lie in the city of Rome, founded, according to legend, by Romulus in 753 BCE. Over centuries, Rome evolved from a small settlement to a formidable republic, characterized by a complex political system with elected officials and checks on power. However, internal strife, class conflicts, and military ambitions paved the way for the end of the Republic. Julius Caesar’s dictatorship and subsequent assassination in 44 BCE created a power vacuum, leading to a civil war. Octavian, later Augustus, emerged victorious, heralding the Roman Empire’s birth.
Under Augustus, the empire experienced the Pax Romana, a 200-year period of relative peace and stability. Augustus reformed the military, established efficient administrative systems, and initiated grand construction projects. The empire's borders expanded, encompassing territories from Britain to Egypt and from Spain to the Euphrates. Roman legions, renowned for their discipline and engineering prowess, secured and maintained these vast territories, building roads, fortifications, and cities that facilitated control and integration.
The Roman Empire’s society was hierarchical, with a rigid class system. At the top were the patricians, wealthy elites who held significant political power. Below them were the plebeians, free citizens with limited political influence, and the vast numbers of slaves who formed the backbone of the economy. The family unit was central, governed by the paterfamilias, the male head who held absolute authority.
Culturally, the Romans were eclectic, absorbing and adapting elements from the civilizations they encountered, particularly the Greeks. Roman art, literature, and philosophy reflected this synthesis, creating a rich cultural tapestry. Latin, the Roman language, became the lingua franca of the Western world, influencing numerous modern languages.
Roman architecture and engineering achievements were monumental. They perfected the arch, vault, and dome, constructing enduring structures like the Colosseum, Pantheon, and aqueducts. These engineering marvels not only showcased Roman ingenuity but also served practical purposes, from public entertainment to water supply.
2. Contents:
Definition of Systematics
Cladograms and Phylogenetic
Trees
Branches of Systematics
1. Numerical Systematics
2. Biochemical Systematics
3. Experimental Systematics
Importance of Systematics
Application of Systematics
3. Systematics:
The term systematics is derived from the Latinised Greek word and ‘systema’ means
‘together’. The systematics partly overlaps with taxonomy and originally used to des-
cribe the system of classification prescribed by early biologists. Linnaeus applied the
word “Systematics” in the system of classification in his famous book ‘Systema Natu-
rae’ published in 1735.
Biological systematics is the study of the diversification of living forms both past and
present and the relationships among living things through time. Relationships are
visualized as evolutionary trees.
According to Blackwelder and Boyden, “systematics is the entire field dealing with
the kinds of animals, their distinction, classification and evolution”.
According to Christoffersen Systematics, is the theory, principles and practice of
identifying systems, i.e., of ordering the diversity of organisms (parts) into
more general systems of taxa according to the most general causal processes”.
The systematics includes both taxonomy and evolution. Taxonomy includes classifi-
cation and nomenclature but inclines heavily on systematics for its concepts. So the
study of systematics includes a much broader aspect that includes not only
morphology and anatomy but also genetics, molecular biology, behavioral aspects
and evolutionary biology. The recent approach to the science of biology has added a
new dimension to the science of classification and the new systematics has emerged
as a synthesis of progress in all the major disciplines of Biology.
Systematics uses taxonomy as a primary tool in understanding as nothing about an
organism's relationships with other living things can be understood without it first
being properly studied and described in sufficient detail to identify and classify it
correctly. Scientific classifications are aids in recording and reporting information to
other scientists and to laymen. The systematist is a scientist who specializes in
4. systematics must therefore, be able to use existing classification systems, or at least
know them well enough to skillfully justify not using them.
Difference between Systematics and Taxonomy:
Taxonomy and systematics are two concepts related to the study of diversification of living
forms and the relationships of living things through time. The main difference between
taxonomy and systematics is that taxonomy is involved in the classification and naming of
organisms whereas systematics is involved in the determination of evolutionary relationships of
organisms. This means systematics ascertain the sharing of the common ancestry by different
organisms. In taxonomy, different organisms are scientifically named and grouped in different
taxonomic levels. Organisms are grouped based on their evolutionary relationships. Taxonomy
can be considered as a branch of systematics. Both taxonomy and systematics use
morphological, behavioral, genetics, and the biochemical observation.
5. Cladograms:
A cladogram is a branching diagram which shows the evolutionary relationship among a group
of clades. A clade is a group of organisms, comprised of all the evolutionary descendants of a
common ancestor. A cladogram does not depict the amount of evolutionary change in the
group, nor does it indicate the evolutionary time or the genetic distance. Each branch of the
cladogram ends with a clade. It starts from a last common ancestor. Cladograms are usually
formed based on the morphological characters.
Cladistics:
Cladistics refers to a biological classification system that involves the categorization of a
organisms based on shared traits. Organisms are typically grouped by how closely related
they are and thus cladistics can be used to trace ancestry back to shared common ancestors
and the evolution of various characteristics.
Phylogenetic Trees:
6. A phylogenetic tree is a branching diagram which shows the inferred relationship between
various biological species. The distance of the branches in the phylogenetic tree represents the
amount of inferred evolutionary change. In order to generate a phylogenetic tree several
characteristics like external morphology, internal anatomy, biochemical pathways, behavior,
DNA and protein sequences, as well as the evidence of fossils have to be used. But,
phylogenetic trees are hypotheses and do not indicate the exact relationships. The data
obtained from DNA sequencing increases the reliability of the relationships in the tree. There
are two components of phylogenetic tree.
Branch order shows the group relationships.
Branch length shows the amount of evolution.
General Phylogenetic Tree
7. Phylogenetic Tree of Vertebrate
Branches of Systematics:
1. Numerical systematics:
This type of systematics is based on bio-statistical method in identification and classification of
animals. This branch is called biometry. It aims to create a taxonomy using numeric algorithms
like cluster analysis rather than using subjective evaluation of their properties. The concept was
first developed by Robert R. Sokal and Peter H. A. Sneath in 1963 and later elaborated by the
same authors. They divided the field into phenetics in which classifications are formed based on
the patterns of overall similarities and cladistics in which classifications are based on the
branching patterns of the estimated evolutionary history of the taxa.
2. Biochemical Systematics:
This branch of systematics deals with classification of animals on the basis of biochemical
analysis of protoplasm.
3. Experimental Systematics:
8. This branch of systematics deals with identification of various evolutionary units within a
species and their role in the process of evolution. Here mutation is considered as evolutionary
unit.
Importance of Systematics:
A. Theoretical biology:
Systematics has played some important role in the field of theoretical biology such as:
It is responsible in making conceptual contribution like population thinking.
It is responsible in solving the problems of multiplication of species. It illustrates the
structure of species and evolutionary processes.
Mimicry and other evolutionary areas also have been a clearly understood through
taxonomy.
It has also played important role in the development of behavioral science.
Taxonomy is the key to the study of ecology, as no ecological survey can be undertaken
unless all the species of ecological importance are identified.
B. Applied Biology:
Systematics provides basic understanding about the components of biodiversity which is
necessary for effective decision making about conservation and sustainable use. The most
important uses are:
1. Agriculture and Forestry:
Presently we are faced with the acute problem of saving our crops and trees from the attack of
various kinds of pests. So, it is necessary to know the correct names of such pests; before their
proper control, and eradication. Taxonomists can give correct identification of pest species,
which is vital for its effective control. Similarly, many of the plant diseases are caused by certain
9. vectors. The correct identification of a particular vector is vital for bringing the vector under
control by killing its transmitters.
2. Biological Control:
Natural enemies of pests can be introduced for biological control of pests. The biological
control is much more economical than the chemical control. In 1940s a
parasite Archytusincertus was introduced from Uruguay and Argentina into the U.S.A. to
control armyworms. The systematics is presently greatly involved in designing and
implementing the biological control programmes of pests and diseases most effectively.
3. Public health:
Taxonomy plays an important role in public health program also. There are number of diseases,
which are spread by many Arthropods. So, our controlled measures should be planned to attack
the target species. As for example, All Anopheles maculipennis are not responsible for
transmitting malaria. This species consists of several sibling species, of which a few were
10. responsible for transmitting malaria. An expert taxonomist can identify this particular sibling
species. A correct identification ensures a maximum of effective control at minimum cost.
4. Quarantine:
Many new pests and diseases of plants, animals and human beings can spread from one
country to another through transportation. Respective Governments have established
quarantine laboratories at aerodromes, ports etc. to check such transmissions. Taxonomists
play a vital role here in prompt identification of these pests and diseases.
5. Wild life management:
Presently great attention is being paid to conserve and propagate wild life. The indiscriminate
killing and felling of trees have already resulted in great disturbance in the natural
environment. Taxonomists can help all environmental protectors by identifying the
economically and ecologically important wild life. The task is important for the preservation and
protection of our biodiversity.
6. Mineral prospecting:
11. The identification of fauna and flora in sedimentary rocks gives a clear picture of the sequence
of geological events, which helps in search for fuels and mineral deposits. The paleontologists
play a major role in the identification of such fossil specimens of the sedimentary rocks and
thus give us a clear picture of the correct sequence of geological events. Such works have been
great success in the industrialization in America.
7. National defense:
Information concerning disease vectors and parasites is an obvious application of
systematics to national defense. The use of biological means in the war is economical and
requires fewer efforts in their operation. During World War II, Japanese paper balloons carrying
paper balloons created havoc in the forest of north east America. Eventually a balloons was
recovered with sand contained a large number of shells of micro-organisms. The taxonomists of
America observed the shells of micro-organisms and confirmed that this type of sand represent
mainland island of Japan. Subsequent bombing of this beach area destroyed the balloon
launching site. Moreover, the identification of potential disease vectors is vital to the health of
both military and civilian populations all over the world.
12. 8. Environmental problem:
Taxonomists have played an important role in detecting some of the environmental
problems. Certain pesticides are entered in the food chain of ecosystem and biomagnifications
of pesticide takes place at certain tropic level. Here a taxonomist can play important role in
detecting such problem and can take effective measures to control it. Presently water pollution
is considered as a major environmental problem. Certain planktons are reliable indicator of the
degree of water pollution. The identification of such organism by taxonomists gives rapid
information for detecting pollution.
13. 9. Soil fertility:
Some organisms play important role in increasing the fertility of soil. So, it is necessary
to know such animals for their proper management in agriculture.
10. In Commerce:
Many animals and animal’s products are used commercially by human beings such as,
honey, silk, lace, dyes etc. Systematics play important role in increasing and improving the
qualities of these products by manipulating the useful species.
Systematists and taxonomists are presently employed by universities, research institute,
museum, central and state govt. agencies, industries, and zoos. A well trained taxonomist is
well qualified teacher to teach course of zoology or biology as he has a great background in
morphology, physiology, genetics and ecology.
Applications of Systematics:
1. Studying the diversity of organisms and the differentiation between extinct and living
creatures. Biologists study the well-understood relationships by making many different
diagrams and "trees".
14. 2. Including the scientific names of organisms, species descriptions and overviews,
taxonomic orders and classifications of evolutionary and organism histories.
3. Explaining the biodiversity of the planet and its organisms. The systematic study is that
of conservation.
4. Manipulating and controlling the natural world. This includes the practice of 'biological
control', the intentional introduction of natural predators and disease.
Application of Systematics in Biology:
1. Systematics is the study of diversity of organisms including past and present and
relationships among living things. Relationships are established by making cladograms,
phylogenetic trees and phylogenies. The phylogeny is the evolutionary history of an
animal or plant, for a taxonomic group.
2. Phylogenies include two parts the first part shows the group relationships and the
second part indicates the amount of evolution. Phylogenetic trees of species and higher
taxa are established by morphological, physiological and molecular characteristics, and
the distribution of animals and their ancestors are related to geography. In this way the
systematics is used to understand the evolutionary history of organisms.
3. The field of systematics provides scientific names of the organisms, description of the
species, ordering the organisms into higher taxa, classification of the organisms and
evolutionary histories.
4. Systematics is also important in implementing the conservation issues because it
attempts to explain the biodiversity which is related to different kinds of species and
could be used in preservation and protect the endangered animals and plants. The loss
of biodiversity is related to the extreme harmful of the existence of mankind. The
unchecked human population destroys different kinds of plants and animals for food
and other factors.
5. The destruction or suppression of harmful pests or animals by the introduction and
increase of their natural enemies is called biological control. The natural enemies of
pests are often introduced for biological control for the advantage of agriculture and
15. forestry. The natural enemies include insectivorous spiders, centipedes, some insects,
frogs and birds which are much more economical than the chemical control because
they have no injurious side effects. The predaceous insects play a vital role in the natural
control of injurious insects. The adult and larval stages of predatory insects of lady bird
beetles are economically very important and are responsible for the destruction of the
colonies of plant lice, scale insects, mealy bugs and white flies which are found as
serious pests in various parts of the world.
6. Some chrysopids are also predatory enemies of mealy bugs and plant lice. An egg
parasite, Trichogramma sp. is utilized in India for the control of sugar cane borers and
boll worms of cotton. In all cases the proper identification of parasites and their hosts
are necessary for the control of the pests. The systematists are involved in implement-
ing the biological control programmes of the pests and diseases most effectively.
7. There are a lot of insects which act as vectors of various human diseases.
For example, some species of Anopheles sp. are the vector of malaria diseases, Aedes
aegypti spreads the virus of dengue fever and phlebotomies argentipes spreads the
pathogens of kala-azar fever. So taxonomists play a vital role in identification of the species
of vectors, and control strategy programmes of the vectors should be planned in such a way
that the target species is attacked.
References:
1. Jump up to "Systematics: Meaning, Branches and Its Application". Biology Discussion.
2016-05-27. Retrieved 2017-04-12.
2. Wilkins, J. S. What is systematics and what is taxonomy Archived 2016-08-27 at
the Wayback Machine. Available on http://evolvingthoughts.
3. Michener, Charles D., John O. Corliss, Richard S. Cowan, Peter H. Raven, Curtis W.
Sabrosky, Donald S. Squires, and G. W. Wharton (1970). Systematics In Support of
16. Biological Research. Division of Biology and Agriculture, National Research Council.
Washington, D.C. 25 pp.
4. Brusca, R. C., & Brusca, G. J. (2003). Invertebrates (2nd ed.). Sunderland, Mass. :
Sinauer Associates, p. 27
5. Fortey, Richard (2008), Dry Store Room No. 1: The Secret Life of the Natural History
Museum, London: Harper Perennial, ISBN 978-0-00-720989-7
6. Mayr, Ernst (1991). Principles of Systematic Zoology. New York: McGraw-Hill, p. 159.
7. Mayr, Ernst (1991), p. 162.