Evolution is any change in the genes of a population over time. The origin of life theories propose that organic molecules assembled from inorganic elements in the primordial atmosphere and oceans around 4 billion years ago. Charles Darwin's theory of natural selection proposed that populations evolve over generations as individuals with favorable variations are more likely to survive and reproduce, passing on those variations. Evidence for evolution includes the fossil record showing sequences of simpler to more complex organisms over time, comparative anatomy revealing homologous and vestigial structures, and similarities in embryonic development across species.