CHARA
CHARA
Division Chlorophyta
Class Chlorophyceae
Order Charales
Family Characeae
Genus Chara
Occurrence
Freshwater
Plants encrusted with calcium carbonate - Stone
warts
Vegetative structure
Multicellular, macrosopic
filamentous, branched
Main axis , rhizoids
Rhizoids
- lower end, branched, multicellular,
uniseriate, obliquely septate,
no nodes and internodes,
Arise from rhizoidal plates
fixation
Main axis
Erect, branched
Differentiated into nodes and internodes
Internodes – long, single celled, enveloped
by corticating threads
Nodes – short, regularly arranged cells
Bears branches
- Branches of limited growth (short laterals)
- Branches of unlimited growth (long laterals)
Branches of limited growth (short laterals)
primary laterals, leaves or branchlets
Arise from peripheral cell of the node
Limited growth
Limited number of nodes and internodes
Arise shorter, unicellular branches from nodes -
--secondary laterals
bear stipules
Fertile branches bear sex organs
Branches of unlimited growth (long laterals)
arise from axils of primary laterals
Arise singly
unlimited growth/indefinitely nodes and internodes,
Cortication bears primary laterals
Cell structure
Two types of cells
At the apex- & at the nodes
Small & isodiametric
Central nucleus
Granular cytoplasm
Chloroplst- numerous, discoid, evenly distributed,
no pyrenoids
Cell structure
Internodal cells
elongated
large vacuole, cytoplasm & nuclei- peripheral
Chloroplast- discoid, numerous, longitudinally
Cytoplsm – peripheral stationary layer-exoplasm
inner fluidy endoplasm
Movement- cyclosis
Cell wall – cellulose, hemicellulose, lignin, pectin
Mucilage covering, Calcium carbonate
Cortication
Internodal cell ensheathed by elongated corticating
threads
Thread consist of smaller cells
Half of corticating arise from uppernode, others from
lower node
Reproduction
Vegetative & sexual methods
Vegetative – amylum stars, bulbils, 20
protenema
a)Amylum stars
star shaped aggregate of vegetative cells
Lower nodes
Store reserve food
Detach from parent ----- new plants
Vegetative reproduction
b)Bulbils
Small, rounded, tuber-like structures, develop on
rhizoids
c) Secondary protenema
nodes/rhizoids
Protenema like structures ----new plants
Sexual reproduction
Oogamous
Sex organs large & visible
Male sex organ- antheridia/globules
Female sex organ – oogonia/nucules
Homothallic (monoecious) – both on same plant
Heterothallic ( Dioecious)- different plants
Nucule is always above the globule
Globule/antheridium
Large, spherical, Red/bright orange coloured
Stalk – pedicel
Outermost wall -8 large closely fitted curved cells-shield
cells encloses an internal cavity
Globule/antheridium
Manubrium
– center of shield cell- rod shaped handle like cell
1o
capitula- more or less rounded cells at the end
Inturn posess 2o
capitula
branched or unbranched antheridial filaments/
spermatogenous filaments
Matre Globule/antheridium
Schield cells move apart
Expose antheridial filaments
200 androytes
Elongated coiled biflagellated antherozoid
OOGONIUM (Nucule)
Macroscopic
Short stalk
Node
Homothallic-Nucule is always above the globule
Tube cell- 5 , spirally twisted, cover, except at the tip
Corona cell- 5, at the tip
Egg- center, uninucleate
Rich of food materials- oil drops & starch grains
Receptive spot- tip of the egg, hyaline
Fertilization
Numerous biflagellate antherozoids liberated outside
Swim- -water –reach nucule
Tube cells elongate near the tip- form neck
Narrow central slit in the corona – passage for
antherozoids
Only one penetrate the wall- fuse with the egg-zygote
Germination
Diploid nucleus migrate towards apex
Secrete cellulosic cell wall
Yellow/ brown
Wall of the tube cell & oogonium -thick & sculpturing
Zygote settles at the bottom
Undergo resting period
Germination of Zygote
vegetative and reproductive structures of Chara
vegetative and reproductive structures of Chara

vegetative and reproductive structures of Chara

  • 1.
  • 2.
    CHARA Division Chlorophyta Class Chlorophyceae OrderCharales Family Characeae Genus Chara
  • 3.
    Occurrence Freshwater Plants encrusted withcalcium carbonate - Stone warts
  • 4.
    Vegetative structure Multicellular, macrosopic filamentous,branched Main axis , rhizoids Rhizoids - lower end, branched, multicellular, uniseriate, obliquely septate, no nodes and internodes, Arise from rhizoidal plates fixation
  • 5.
    Main axis Erect, branched Differentiatedinto nodes and internodes Internodes – long, single celled, enveloped by corticating threads Nodes – short, regularly arranged cells Bears branches - Branches of limited growth (short laterals) - Branches of unlimited growth (long laterals)
  • 6.
    Branches of limitedgrowth (short laterals) primary laterals, leaves or branchlets Arise from peripheral cell of the node Limited growth Limited number of nodes and internodes Arise shorter, unicellular branches from nodes - --secondary laterals bear stipules Fertile branches bear sex organs
  • 7.
    Branches of unlimitedgrowth (long laterals) arise from axils of primary laterals Arise singly unlimited growth/indefinitely nodes and internodes, Cortication bears primary laterals
  • 8.
    Cell structure Two typesof cells At the apex- & at the nodes Small & isodiametric Central nucleus Granular cytoplasm Chloroplst- numerous, discoid, evenly distributed, no pyrenoids
  • 9.
    Cell structure Internodal cells elongated largevacuole, cytoplasm & nuclei- peripheral Chloroplast- discoid, numerous, longitudinally Cytoplsm – peripheral stationary layer-exoplasm inner fluidy endoplasm Movement- cyclosis Cell wall – cellulose, hemicellulose, lignin, pectin Mucilage covering, Calcium carbonate
  • 10.
    Cortication Internodal cell ensheathedby elongated corticating threads Thread consist of smaller cells Half of corticating arise from uppernode, others from lower node
  • 11.
    Reproduction Vegetative & sexualmethods Vegetative – amylum stars, bulbils, 20 protenema a)Amylum stars star shaped aggregate of vegetative cells Lower nodes Store reserve food Detach from parent ----- new plants
  • 12.
    Vegetative reproduction b)Bulbils Small, rounded,tuber-like structures, develop on rhizoids c) Secondary protenema nodes/rhizoids Protenema like structures ----new plants
  • 13.
    Sexual reproduction Oogamous Sex organslarge & visible Male sex organ- antheridia/globules Female sex organ – oogonia/nucules Homothallic (monoecious) – both on same plant Heterothallic ( Dioecious)- different plants Nucule is always above the globule
  • 14.
    Globule/antheridium Large, spherical, Red/brightorange coloured Stalk – pedicel Outermost wall -8 large closely fitted curved cells-shield cells encloses an internal cavity
  • 15.
    Globule/antheridium Manubrium – center ofshield cell- rod shaped handle like cell 1o capitula- more or less rounded cells at the end Inturn posess 2o capitula branched or unbranched antheridial filaments/ spermatogenous filaments
  • 16.
    Matre Globule/antheridium Schield cellsmove apart Expose antheridial filaments 200 androytes Elongated coiled biflagellated antherozoid
  • 17.
    OOGONIUM (Nucule) Macroscopic Short stalk Node Homothallic-Nuculeis always above the globule Tube cell- 5 , spirally twisted, cover, except at the tip Corona cell- 5, at the tip Egg- center, uninucleate Rich of food materials- oil drops & starch grains Receptive spot- tip of the egg, hyaline
  • 18.
    Fertilization Numerous biflagellate antherozoidsliberated outside Swim- -water –reach nucule Tube cells elongate near the tip- form neck Narrow central slit in the corona – passage for antherozoids Only one penetrate the wall- fuse with the egg-zygote
  • 19.
    Germination Diploid nucleus migratetowards apex Secrete cellulosic cell wall Yellow/ brown Wall of the tube cell & oogonium -thick & sculpturing Zygote settles at the bottom Undergo resting period
  • 20.