Strain markers are objects in deformed rocks that can be used to measure strain. Good strain markers include reduction spots, pebbles, fossils, fold sets, and lineations. Spherical markers originally circular in cross-section become elliptical due to homogeneous deformation, with the ratio of major to minor axes indicating strain. Fossils with lines of symmetry like trilobites and brachiopods can indicate the principal strain directions. Fold sets allow comparing initial and final layered sequences to analyze strain. Schistosity and lineations in metamorphic rocks can also act as strain markers.