GlusterFS uses "translators" to modify and route file requests between users and storage bricks. Translators can convert request types, modify request properties like paths or flags, intercept or block requests, and spawn new requests. This allows GlusterFS to provide features like replication, caching, and integration with other systems, but also enables custom file systems to be built by modifying the translators. The asynchronous programming model and shared context objects allow translators to cooperate complex workflows across multiple servers.