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Timber and Wildlife Ecology 
Research Update from the Jones 
Center 
Forest Landowner’s Association 2014 Southeast Regional 
Forest Resource Owner and Manager Conference 
Valdosta, GA October 28, 2014 
Kevin McIntyre 
Education Coordinator 
& 
Steve Jack 
Conservation Ecologist
Joseph W. Jones Ecological Research Center 
at Ichauway 
Supported by the Robert W. Woodruff Foundation
Joseph W. Jones Ecological Research 
Ichauway, circa 1929 
Robert Woodruff 
Joseph W. Jones 
Center at Ichauway
Long-Term Research 
Longleaf Pine Ecosystems 
1) Productivity and biodiversity patterns of a 
longleaf pine ecosystem. 
2) Ecological forestry and restoration of longleaf 
pine ecosystems. 
Joseph W. Jones Ecological Research Center 
at Ichauway 
3) Ecological role of mesopredators, effects of 
control, and habitat approaches. 
Aquatic Ecology and Water Resources 
4) Hydrologic variation and human development in 
the lower Flint River Basin 
5) Depressional wetlands on the coastal plain 
landscape: maintenance of regional biodiversity
Conservation & Natural Resource 
Management 
• Stewardship of Ichauway & its natural 
resources 
• Management of forests, wildlife & 
wetlands, including prescribed fire 
• Monitoring of key environmental 
variables, plant & animal populations 
• New approaches for restoration & 
adaptive management 
• Key role in participation with education & 
research programs
Education and Outreach 
• Natural Resource 
Professionals 
• Policymakers 
• Land Owners/Managers 
• Undergraduate University 
Classes 
• Graduate Students
Regional Longleaf Pine Conservation 
and Restoration 
• Overall goal of doubling acreage of LLP 
from 4m to 8m by 2025 
• Supported by diverse coalition of 
Federal, State, NGO and private sector 
• Identified Significant Geographic Areas 
for longleaf restoration focus 
• Significantly increased funding for LLP 
• Decline of LLP reversed from low of 
3.84m to 4.28m as of 2010 
• 157,000 acres LLP established in 2013 
• 50% incentive funded 
• 50% funded by private landowners 
8
Forest Management Limitations 
Timber
Forest Management Limitations Tradeoffs 
Timber 
Timber
Wildlife Research 
Quail
Is Hard Mast a Limiting Factor?
Ichauway Turkey Track Count Data
Ichauway WTD Track Counts
Ichauway WTD Spotlight Data
Season of Burn & Wildlife Impacts
Growing-Season Fire Impacts 
Nest Success: 
• Growing-season fires had an effect (p-value=0.03) 
– 11% of nests burned 
– Majority burned in April 
– 80% of hens renested 
Poult Survival: 
• One newly hatched brood lost to fire
Poult Survival Data 
Year Site n 
Day 0-9 
(%lost) 
Day 10-16 
(%lost) 
Day 17-30 
(%lost) 
Day 30+ 
(%lost) 
2011 JC 6 67 17 17 N/A 
SL 3 33 0 33 0 
Pooled 9 56 11 22 0 
2012 JC 9 78 11 
SL 5 40 0 20 40 
Pooled 14 64 7 7 21 
Both Years/Sites 23 61 17 26 22 
• Average # of poults per hen at hatch: 8.6 
• 5 of the 23 brood hens killed by predators
Management Implications 
Predation much stronger driver of turkey population 
dynamics than season of burn 
Manage for habitat quality and population vs. individual nests 
Use rotating small-scale growing-season burns 
More to Wild Turkey Population Dynamics than Burns…
Predator Exclusion Experiment
Do Exclosures Exclude?
Annual Coyote Diet
Indirect Effects? 
Photo credit: Austin Baggarley
Deer use of predator exclosures 
70 
60 
50 
40 
30 
20 
10 
0 
Gestation Fawning Breeding 
White-tailed deer detections 
Control 
Exclosure
22 fawns/year 
0.41 vs. 0.77 fawns/doe
Longleaf Management and 
Restoration Research
Timber Research 
Timber
Conventional Wisdom 
USDA Silvics Manual – Volume 1: 
Conifers (Agriculture Handbook 654, 1990) 
Pinus palustris (Mill.) 
(from chapter by Boyer) 
“Reaction to Competition- 
Longleaf pine is intolerant of 
competition, whether for light or 
for moisture and nutrients. The 
species will grow best in the 
complete absence of all 
competition, including that from 
other members of the species.” 
Thus, general recommendation 
was to manage using even-aged 
systems (Boyer and Peterson, 1983, 
Agriculture Handbook 445)
Natural Stands – Multi-aged
Regeneration Dynamics Questions 
• Below-canopy light 
environment 
• Competition (both inter-and 
intra-specific) 
• Role of fire 
• Management implications
Competition 
Pine Seedling Response 
to Canopy Gap Size 
Seedling growth 
Total biomass (g) 
30 
25 
20 
15 
10 
5 
0 
Seedling survival 
Large group 
Overstory treatment 
Uncut 
Single-tree 
Small group 
Mean survival (%) 
85 
80 
75 
70 
65 
60 
a 
ab 
c 
bc 
a 
a a 
b 
Pecot et al. 2007. CJFR
Three Stage 
Model for 
Seedling Survival 
and Growth
Canopy Gap Size & Advanced 
Regeneration 
Think in terms of releasing 
advanced regeneration rather 
than creating opening to 
secure regeneration
Canopy Gap Size & Advanced 
Regeneration 
O’Brien et al. 2008. Ambio
Diverse Ground Cover Community
Functionally important ground cover species 
Lower decomposition rate of elevated fine fuels 
(wiregrass & pine needles) 
0 200 400 600 800 1000 1200 
Days in Field 
Mass Remaining (%) 
100 
90 
80 
70 
60 
50 
40 
30 
20 
10 
0 
Hendricks et al. 2002 
Wiregrass (soil) 
Wiregrass (elevated) 
Longleaf pine (soil) 
Longleaf pine (elevated)
Functionally important species: 
Legume Species 
Foliar Ndfa (%) 
100 
90 
80 
70 
60 
50 
40 
30 
20 
10 
0 
Legumes 
No. of species identified: 43 
Average density: 115,000 
stems/ha 
Wet-Mesic Site 
Intermediate Site 
Cassia 
nictitans 
Crotalaria 
rotundifolia 
Centrosema 
virginiana 
Lespedeza 
virginiana 
Potential N2-fixation by legumes 
Hainds et al. 1999 
Hiers et al. 2001 
Cathey et al. 2010
How Do We Best Manage to 
Maintain and Restore This Forest 
Type While Utilizing Resources?
Financial Comparison 
Harvest Scenario Low Medium High 
20-year IRR w/ land value (%) 3.21 3.29 3.48 
Total accumulated net cash flow ($) 1,796,027 3,732,401 4,986,258 
Ending Total Value (Cash flow + ending value) ($) 6,703,877 7,445,119 8,104,815 
Conversion Scenario Moderate Rapid 
20-year IRR w/ land value (%) 4.21 5.77 
Total accumulated net cash flow ($) 5,171,745 4,928,824 
Ending Total Value (Cash flow + ending value) ($) 6,612,212 6,569,429
Managing Longleaf Forests 
Using natural disturbance regime as guide to 
management  uneven-aged approaches
Which Silvicultural System? 
• Disturbance regime and regeneration dynamics 
point toward uneven-aged systems to meet 
objectives 
• Group or individual tree selection are feasible 
approaches for restoring toward reference 
objective
Long-Term Management and 
Restoration Experiments
Fuels, Fire & Silviculture 
Attentuation of Needle Cast with Distance from Gap Edge 
-2 0 2 4 6 8 10 12 14 16 
Distance from Gap Edge 
Needle Cast (grams per 1/4 m2 trap) 
40 
30 
20 
10 
0 
grams of needles/ trap
Regeneration Dynamics
Ground Cover Recovery
Harvest Treatments 
Uncut Control 
Single-tree Selection 
Group Selection 
Group Selection 
with Retention
Stem Maps
Skidder Traffic – Operational Example 
* Also have “experimental” skidder treatments
Soil and Ground Cover Disturbance 
1 Pass 2 Passes 
4 Passes 8 Passes
Soil Compaction 
Operational Skid 
Trails
Ground Cover Changes 
Wiregrass Dominated 
Pre-harvest, 2009 Post-harvest, 2010
Ground Cover Changes 
Old-Field Dominated 
Pre-harvest, 2009 Post-harvest, 2010
Planted 
Seedling 
Biomass
Management Project Summary 
• Still preliminary results, early in study 
• Short-term weather patterns can affect results 
• Few statistically significant results due to high 
variability, though are some evident trends 
• Harvesting operations have impacts but do 
not appear severe, recovery occurs over long 
time span
Restoration Research 
Restoration – timing and pathways 
– Species conversion 
– When to introduce ground cover 
– Wildlife habitat and which species present at 
different stages of development
Plantations to “Natural” Stands
Plantation Species Conversion 
Underplanting longleaf seedlings in gaps 
Thinning planted slash pine with gap creation 
Reintroduction of ground cover? 
- when and what species
Old Field Conversion
Ground Cover Restoration
Hardwood Encroachment in Uplands 
Legacy of past management 
How to reconnect the landscape?
Hardwood Encroachment in Uplands 
• Silvicultural treatment is species conversion – removal 
followed by planting 
• Focused on trees, other things not immediately restored
• Explicitly factor time into considerations – 
thinking of time as an ecological factor that 
cannot be purchased or replaced 
• In restoration, make use results of time 
already “invested”
Summary 
• Integration between research and operational 
management and between research subject 
areas 
• Research is focused on long-term projects 
with operational scale 
• Attempting to examine multiple resource 
responses to manipulations – trade-offs from 
maintaining all ecosystem characteristics

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“Timber and Wildlife Ecology Research Update from the Jones Center” Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

  • 1. Timber and Wildlife Ecology Research Update from the Jones Center Forest Landowner’s Association 2014 Southeast Regional Forest Resource Owner and Manager Conference Valdosta, GA October 28, 2014 Kevin McIntyre Education Coordinator & Steve Jack Conservation Ecologist
  • 2. Joseph W. Jones Ecological Research Center at Ichauway Supported by the Robert W. Woodruff Foundation
  • 3. Joseph W. Jones Ecological Research Ichauway, circa 1929 Robert Woodruff Joseph W. Jones Center at Ichauway
  • 4.
  • 5. Long-Term Research Longleaf Pine Ecosystems 1) Productivity and biodiversity patterns of a longleaf pine ecosystem. 2) Ecological forestry and restoration of longleaf pine ecosystems. Joseph W. Jones Ecological Research Center at Ichauway 3) Ecological role of mesopredators, effects of control, and habitat approaches. Aquatic Ecology and Water Resources 4) Hydrologic variation and human development in the lower Flint River Basin 5) Depressional wetlands on the coastal plain landscape: maintenance of regional biodiversity
  • 6. Conservation & Natural Resource Management • Stewardship of Ichauway & its natural resources • Management of forests, wildlife & wetlands, including prescribed fire • Monitoring of key environmental variables, plant & animal populations • New approaches for restoration & adaptive management • Key role in participation with education & research programs
  • 7. Education and Outreach • Natural Resource Professionals • Policymakers • Land Owners/Managers • Undergraduate University Classes • Graduate Students
  • 8. Regional Longleaf Pine Conservation and Restoration • Overall goal of doubling acreage of LLP from 4m to 8m by 2025 • Supported by diverse coalition of Federal, State, NGO and private sector • Identified Significant Geographic Areas for longleaf restoration focus • Significantly increased funding for LLP • Decline of LLP reversed from low of 3.84m to 4.28m as of 2010 • 157,000 acres LLP established in 2013 • 50% incentive funded • 50% funded by private landowners 8
  • 10. Forest Management Limitations Tradeoffs Timber Timber
  • 12. Is Hard Mast a Limiting Factor?
  • 13. Ichauway Turkey Track Count Data
  • 16.
  • 17. Season of Burn & Wildlife Impacts
  • 18. Growing-Season Fire Impacts Nest Success: • Growing-season fires had an effect (p-value=0.03) – 11% of nests burned – Majority burned in April – 80% of hens renested Poult Survival: • One newly hatched brood lost to fire
  • 19. Poult Survival Data Year Site n Day 0-9 (%lost) Day 10-16 (%lost) Day 17-30 (%lost) Day 30+ (%lost) 2011 JC 6 67 17 17 N/A SL 3 33 0 33 0 Pooled 9 56 11 22 0 2012 JC 9 78 11 SL 5 40 0 20 40 Pooled 14 64 7 7 21 Both Years/Sites 23 61 17 26 22 • Average # of poults per hen at hatch: 8.6 • 5 of the 23 brood hens killed by predators
  • 20. Management Implications Predation much stronger driver of turkey population dynamics than season of burn Manage for habitat quality and population vs. individual nests Use rotating small-scale growing-season burns More to Wild Turkey Population Dynamics than Burns…
  • 24. Indirect Effects? Photo credit: Austin Baggarley
  • 25. Deer use of predator exclosures 70 60 50 40 30 20 10 0 Gestation Fawning Breeding White-tailed deer detections Control Exclosure
  • 26. 22 fawns/year 0.41 vs. 0.77 fawns/doe
  • 27. Longleaf Management and Restoration Research
  • 29. Conventional Wisdom USDA Silvics Manual – Volume 1: Conifers (Agriculture Handbook 654, 1990) Pinus palustris (Mill.) (from chapter by Boyer) “Reaction to Competition- Longleaf pine is intolerant of competition, whether for light or for moisture and nutrients. The species will grow best in the complete absence of all competition, including that from other members of the species.” Thus, general recommendation was to manage using even-aged systems (Boyer and Peterson, 1983, Agriculture Handbook 445)
  • 30. Natural Stands – Multi-aged
  • 31. Regeneration Dynamics Questions • Below-canopy light environment • Competition (both inter-and intra-specific) • Role of fire • Management implications
  • 32. Competition Pine Seedling Response to Canopy Gap Size Seedling growth Total biomass (g) 30 25 20 15 10 5 0 Seedling survival Large group Overstory treatment Uncut Single-tree Small group Mean survival (%) 85 80 75 70 65 60 a ab c bc a a a b Pecot et al. 2007. CJFR
  • 33. Three Stage Model for Seedling Survival and Growth
  • 34. Canopy Gap Size & Advanced Regeneration Think in terms of releasing advanced regeneration rather than creating opening to secure regeneration
  • 35. Canopy Gap Size & Advanced Regeneration O’Brien et al. 2008. Ambio
  • 36. Diverse Ground Cover Community
  • 37. Functionally important ground cover species Lower decomposition rate of elevated fine fuels (wiregrass & pine needles) 0 200 400 600 800 1000 1200 Days in Field Mass Remaining (%) 100 90 80 70 60 50 40 30 20 10 0 Hendricks et al. 2002 Wiregrass (soil) Wiregrass (elevated) Longleaf pine (soil) Longleaf pine (elevated)
  • 38. Functionally important species: Legume Species Foliar Ndfa (%) 100 90 80 70 60 50 40 30 20 10 0 Legumes No. of species identified: 43 Average density: 115,000 stems/ha Wet-Mesic Site Intermediate Site Cassia nictitans Crotalaria rotundifolia Centrosema virginiana Lespedeza virginiana Potential N2-fixation by legumes Hainds et al. 1999 Hiers et al. 2001 Cathey et al. 2010
  • 39. How Do We Best Manage to Maintain and Restore This Forest Type While Utilizing Resources?
  • 40. Financial Comparison Harvest Scenario Low Medium High 20-year IRR w/ land value (%) 3.21 3.29 3.48 Total accumulated net cash flow ($) 1,796,027 3,732,401 4,986,258 Ending Total Value (Cash flow + ending value) ($) 6,703,877 7,445,119 8,104,815 Conversion Scenario Moderate Rapid 20-year IRR w/ land value (%) 4.21 5.77 Total accumulated net cash flow ($) 5,171,745 4,928,824 Ending Total Value (Cash flow + ending value) ($) 6,612,212 6,569,429
  • 41. Managing Longleaf Forests Using natural disturbance regime as guide to management  uneven-aged approaches
  • 42. Which Silvicultural System? • Disturbance regime and regeneration dynamics point toward uneven-aged systems to meet objectives • Group or individual tree selection are feasible approaches for restoring toward reference objective
  • 43. Long-Term Management and Restoration Experiments
  • 44. Fuels, Fire & Silviculture Attentuation of Needle Cast with Distance from Gap Edge -2 0 2 4 6 8 10 12 14 16 Distance from Gap Edge Needle Cast (grams per 1/4 m2 trap) 40 30 20 10 0 grams of needles/ trap
  • 47. Harvest Treatments Uncut Control Single-tree Selection Group Selection Group Selection with Retention
  • 49. Skidder Traffic – Operational Example * Also have “experimental” skidder treatments
  • 50. Soil and Ground Cover Disturbance 1 Pass 2 Passes 4 Passes 8 Passes
  • 52. Ground Cover Changes Wiregrass Dominated Pre-harvest, 2009 Post-harvest, 2010
  • 53. Ground Cover Changes Old-Field Dominated Pre-harvest, 2009 Post-harvest, 2010
  • 55. Management Project Summary • Still preliminary results, early in study • Short-term weather patterns can affect results • Few statistically significant results due to high variability, though are some evident trends • Harvesting operations have impacts but do not appear severe, recovery occurs over long time span
  • 56. Restoration Research Restoration – timing and pathways – Species conversion – When to introduce ground cover – Wildlife habitat and which species present at different stages of development
  • 58. Plantation Species Conversion Underplanting longleaf seedlings in gaps Thinning planted slash pine with gap creation Reintroduction of ground cover? - when and what species
  • 61. Hardwood Encroachment in Uplands Legacy of past management How to reconnect the landscape?
  • 62. Hardwood Encroachment in Uplands • Silvicultural treatment is species conversion – removal followed by planting • Focused on trees, other things not immediately restored
  • 63. • Explicitly factor time into considerations – thinking of time as an ecological factor that cannot be purchased or replaced • In restoration, make use results of time already “invested”
  • 64. Summary • Integration between research and operational management and between research subject areas • Research is focused on long-term projects with operational scale • Attempting to examine multiple resource responses to manipulations – trade-offs from maintaining all ecosystem characteristics