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Successful EEssttaabblliisshhmmeenntt ooff LLoonngglleeaaff 
RReeggeenneerraattiioonn iinn RReedd HHiillllss ooff GGAA && FFLL
co The couunnttrryy aarroouunndd TTaallllaahhaasssseeee aanndd 
TThhoommaassvviillllee iiss ttoo qquuaaiill hhuunnttiinngg wwhhaatt tthhee 
bblluueeggrraassss ooff KKeennttuucckkyy iiss ttoo tthhoorroouugghhbbrreeddss 
aanndd BBoorrddeeaauuxx iiss ttoo wwiinnee.. 
------ FFoorrbbeess MMaaggaazziinnee
TThhee RReedd HHiillllss 
hhaass bbeeeenn oonnee ooff 
tthhee ffeeww ppllaacceess iinn 
tthhee ssoouutthheeaasstt 
wwhheerree ffiirree hhaass 
ppeerrssiisstteedd iinn tthhee 
ccuullttuurree aatt tthhee 
llaannddssccaappee lleevveell 
ffoorr aa lloonngg ppeerriioodd 
ooff ttiimmee..
The ddeennssiittyy ooff ttiimmbbeerr ssttaannddss sshhoouulldd 
rreecceeiivvee ccoonnssttaanntt aatttteennttiioonn oonn tthhee 
wweellll--mmaannaaggeedd qquuaaiill pprreesseerrvvee.. 
—— HHeerrbbeerrtt LL.. SSttooddddaarrdd,, TThhee BBoobbwwhhiittee 
QQuuaaiill,, IItt’’ss HHaabbiittss,, PPrreesseerrvvaattiioonn aanndd 
IInnccrreeaassee
Relationship between timber basal area and bobwhite 
abundance 
SH 
TT 
SH 
SH 
TT TT 
TT 
TT 
LL 
LL 
LL 
PH 
PH 
PH 
PH 
DE 
DE 
DE 
SW 
SW 
DE 
LL 
SW 
4 6 8 10 12 14 16 18 20 22 
Basal Area (m 2 / hectare) 
8 
6 
4 
2 
18 
16 
14 
12 
10 
Bobwhite Covey Count 
r = -0.6118, p = 0.0019
Herbert L. Stoddard, Sr., Pioneering Wildlife Ecologist and 
Founder of Tall Timbers Research Station
Herbert L. Stoddard, Sr. 
Wade Tract - circa 1950 
DDeevveellooppeedd tthhee 
SSttooddddaarrdd--NNeeeell MMeetthhoodd 
ooff FFoorreesstt MMaannaaggeemmeenntt 
Leon Neel 
Greenwood - circa 2001
Traditional UEM
SSttooddddaarrdd--NNeeeell UUEEMM
Natural Disturbance Regime(s) of 
the Coastal Plain 
• Fire (natural and anthropogenic origin) 
– Lightning strikes kill individual trees 
– Frequent low intensity (1-5 yr interval) 
– Fire intensity can intensify during drought, 
mortality resulting 
• Wind 
– Significant player in coastal regions 
– Less frequent but larger scale events (Hurricanes) 
• Insects/pathogens 
– Southern pine beetle 
(LL is fairly resistant) 
– Brown spot
Patterns of Canopy Disturbance 
Mature Forest 
Stand Replacing 
Gap Creating 
Partial 
Disturbance
Silvicultural Practices For EAM 
Two-aged Potential
Silvicultural Practices For UEAM 
Balanced 
Unbalanced
Longleaf Pine 
Population Dynamics 
(Platt et al. 1988)
Actual Diameter Distributions
Growing 
Season 
EEssttaabblliisshhmmeenntt 
WWiitthh FFiirree 
Dormant Season
FFrreeqquueennccyy ooff FFiirree:: 
FFiirree ffrreeqquueennccyy iiss oonnee ooff tthhee mmoosstt iiff 
nnoott tthhee mmoosstt iimmppoorrttaanntt ooff aallll tthhee 
ffiirree vvaarriiaabblleess 
• 33 yyeeaarrss iiss aa eeccoollooggiiccaall tthhrreesshhoolldd 
• SShhiifftt ffrroomm wwooooddyy ttoo hheerrbbaacceeoouuss 
ddoommiinnaannccee 
Is important because it is a system driver
Fire Scars (Huffman, 2007) 
72% of the sample
Brockway and Boyer 
5-7+ yr interval
Morriss and Mills 1948 
Age of rough 0 1 2 0 1 2 0 1 2 0 1 
Seed source 
VG G F P
WWaaddee TTrraacctt EExxaammppllee 
Old-growth lloonngglleeaaff ppiinnee,, hhaadd 
bbeeeenn bbuurrnneedd aannnnuuaallllyy iinn tthhee 
ddoorrmmaanntt sseeaassoonn ffoorr oovveerr 110000 
yyeeaarrss.. 
NNoo rreeggeenneerraattiioonn,, ccaannooppyy 
bbrreeaakkddoowwnn,, bbrraacckkeenn ffeerrnn 
uunnddeerrssttoorryy
In 1982 mmoovveedd ttoo ggrroowwiinngg 
sseeaassoonn ((MMaayy--JJuunnee)) bbuurrnnss oonn aa 
22--yyeeaarr ccyyccllee.. 
RReeggeenneerraattiioonn,, ssttiillll ooppeenn 
ccaannooppyy,, wwiirreeggrraassss uunnddeerrssttoorryy 
NNoo SSooiill DDiissttuurrbbaannccee ttoo 
CCaappttuurree LLLLPP
160 
120 
80 
40 
0 15 30 45 60 75 90 
Size Class (cm) 
0 
PIPA Stems/4 ha 
HA4_76 
HA4_96 
1979 
1996
HHiigghh DDeennssiittyy QQuuaaiill PPrrooppeerrttyy 
2-year Fire Frequency 
March-April Fires 
Selecting for LLP- Mowers
Conversion of Old Field Pines 
Under Planting and Gap 
Planting 
UUNNDDEERR PPLLAANNTTIINNGG LLLLPP 
BBaassaall AArreeaass << 6600 fftt^^22//aaccrree 
BBuurrnn LLaattee SSpprriinngg oorr SSuummmmeerr 
PPllaanntt tthhaatt FFaallll 
uussee aa 44xx44’’ ssppaacciinngg 
aaccrree pplloottss oorr 9900--112200’’ wwiiddee ssttrriippss
UUNNDDEERR PPLLAANNTTIINNGG LLLLPP CCoonntt 
HHeerrbbiicciiddeess ((GGaarrlloonn)) ccaann bbee uusseedd ttoo iimmpprroovvee GGrroouunndd ccoovveerr 
SSpprraayy ssuummmmeerr pprriioorr ttoo ppllaannttiinngg 
ffrroomm ffoorrbb//hhaarrddwwooooddss ttoo ggrraasssseess
First Burn 18 months post planting 
Two Year Fire Return Interval naturally pruning and quality trees
Planting Forest Openings 
Planting Density and Fire
Pruned LL Pines
Conversion of Crop Fields 
Poorly Formed Trees – higher fertility sites? 
Generally Higher Stocking Rate - 605 TPA 
Pruning will be needed at lower TPA 
Ground cover to permit burning – first MAY avoid issues
Establish Ground CCoovveerr –– BBuurrnniinngg 
CClluummpp ppllaannttiinngg vveerrssuuss rrooww ppllaannttiinngg
In a Nutshell 
• Stoddard-Neel System is a form of 
UEA Management 
• Bimodal Distribution versus Reverse J 
• Existing LLP – Fire and Gaps 
• Works well with wildlife Values 
• Retains significant old growth portion 
• Under-planting with frequent fire 
– May require herbicide as site prep 
• Field systems require heavier stocking 
– Establish burnable ground cover

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“Successful Establishment of Longleaf Regeneration in the Red Hills of GA and FL” Bill Palmer, Director, Tall Timbers Research Center, Tallahassee, FL

  • 1. Successful EEssttaabblliisshhmmeenntt ooff LLoonngglleeaaff RReeggeenneerraattiioonn iinn RReedd HHiillllss ooff GGAA && FFLL
  • 2.
  • 3. co The couunnttrryy aarroouunndd TTaallllaahhaasssseeee aanndd TThhoommaassvviillllee iiss ttoo qquuaaiill hhuunnttiinngg wwhhaatt tthhee bblluueeggrraassss ooff KKeennttuucckkyy iiss ttoo tthhoorroouugghhbbrreeddss aanndd BBoorrddeeaauuxx iiss ttoo wwiinnee.. ------ FFoorrbbeess MMaaggaazziinnee
  • 4. TThhee RReedd HHiillllss hhaass bbeeeenn oonnee ooff tthhee ffeeww ppllaacceess iinn tthhee ssoouutthheeaasstt wwhheerree ffiirree hhaass ppeerrssiisstteedd iinn tthhee ccuullttuurree aatt tthhee llaannddssccaappee lleevveell ffoorr aa lloonngg ppeerriioodd ooff ttiimmee..
  • 5.
  • 6.
  • 7. The ddeennssiittyy ooff ttiimmbbeerr ssttaannddss sshhoouulldd rreecceeiivvee ccoonnssttaanntt aatttteennttiioonn oonn tthhee wweellll--mmaannaaggeedd qquuaaiill pprreesseerrvvee.. —— HHeerrbbeerrtt LL.. SSttooddddaarrdd,, TThhee BBoobbwwhhiittee QQuuaaiill,, IItt’’ss HHaabbiittss,, PPrreesseerrvvaattiioonn aanndd IInnccrreeaassee
  • 8. Relationship between timber basal area and bobwhite abundance SH TT SH SH TT TT TT TT LL LL LL PH PH PH PH DE DE DE SW SW DE LL SW 4 6 8 10 12 14 16 18 20 22 Basal Area (m 2 / hectare) 8 6 4 2 18 16 14 12 10 Bobwhite Covey Count r = -0.6118, p = 0.0019
  • 9. Herbert L. Stoddard, Sr., Pioneering Wildlife Ecologist and Founder of Tall Timbers Research Station
  • 10. Herbert L. Stoddard, Sr. Wade Tract - circa 1950 DDeevveellooppeedd tthhee SSttooddddaarrdd--NNeeeell MMeetthhoodd ooff FFoorreesstt MMaannaaggeemmeenntt Leon Neel Greenwood - circa 2001
  • 13. Natural Disturbance Regime(s) of the Coastal Plain • Fire (natural and anthropogenic origin) – Lightning strikes kill individual trees – Frequent low intensity (1-5 yr interval) – Fire intensity can intensify during drought, mortality resulting • Wind – Significant player in coastal regions – Less frequent but larger scale events (Hurricanes) • Insects/pathogens – Southern pine beetle (LL is fairly resistant) – Brown spot
  • 14. Patterns of Canopy Disturbance Mature Forest Stand Replacing Gap Creating Partial Disturbance
  • 15. Silvicultural Practices For EAM Two-aged Potential
  • 16. Silvicultural Practices For UEAM Balanced Unbalanced
  • 17. Longleaf Pine Population Dynamics (Platt et al. 1988)
  • 19. Growing Season EEssttaabblliisshhmmeenntt WWiitthh FFiirree Dormant Season
  • 20. FFrreeqquueennccyy ooff FFiirree:: FFiirree ffrreeqquueennccyy iiss oonnee ooff tthhee mmoosstt iiff nnoott tthhee mmoosstt iimmppoorrttaanntt ooff aallll tthhee ffiirree vvaarriiaabblleess • 33 yyeeaarrss iiss aa eeccoollooggiiccaall tthhrreesshhoolldd • SShhiifftt ffrroomm wwooooddyy ttoo hheerrbbaacceeoouuss ddoommiinnaannccee Is important because it is a system driver
  • 21. Fire Scars (Huffman, 2007) 72% of the sample
  • 22. Brockway and Boyer 5-7+ yr interval
  • 23. Morriss and Mills 1948 Age of rough 0 1 2 0 1 2 0 1 2 0 1 Seed source VG G F P
  • 24. WWaaddee TTrraacctt EExxaammppllee Old-growth lloonngglleeaaff ppiinnee,, hhaadd bbeeeenn bbuurrnneedd aannnnuuaallllyy iinn tthhee ddoorrmmaanntt sseeaassoonn ffoorr oovveerr 110000 yyeeaarrss.. NNoo rreeggeenneerraattiioonn,, ccaannooppyy bbrreeaakkddoowwnn,, bbrraacckkeenn ffeerrnn uunnddeerrssttoorryy
  • 25.
  • 26. In 1982 mmoovveedd ttoo ggrroowwiinngg sseeaassoonn ((MMaayy--JJuunnee)) bbuurrnnss oonn aa 22--yyeeaarr ccyyccllee.. RReeggeenneerraattiioonn,, ssttiillll ooppeenn ccaannooppyy,, wwiirreeggrraassss uunnddeerrssttoorryy NNoo SSooiill DDiissttuurrbbaannccee ttoo CCaappttuurree LLLLPP
  • 27. 160 120 80 40 0 15 30 45 60 75 90 Size Class (cm) 0 PIPA Stems/4 ha HA4_76 HA4_96 1979 1996
  • 28. HHiigghh DDeennssiittyy QQuuaaiill PPrrooppeerrttyy 2-year Fire Frequency March-April Fires Selecting for LLP- Mowers
  • 29. Conversion of Old Field Pines Under Planting and Gap Planting UUNNDDEERR PPLLAANNTTIINNGG LLLLPP BBaassaall AArreeaass << 6600 fftt^^22//aaccrree BBuurrnn LLaattee SSpprriinngg oorr SSuummmmeerr PPllaanntt tthhaatt FFaallll uussee aa 44xx44’’ ssppaacciinngg aaccrree pplloottss oorr 9900--112200’’ wwiiddee ssttrriippss
  • 30. UUNNDDEERR PPLLAANNTTIINNGG LLLLPP CCoonntt HHeerrbbiicciiddeess ((GGaarrlloonn)) ccaann bbee uusseedd ttoo iimmpprroovvee GGrroouunndd ccoovveerr SSpprraayy ssuummmmeerr pprriioorr ttoo ppllaannttiinngg ffrroomm ffoorrbb//hhaarrddwwooooddss ttoo ggrraasssseess
  • 31. First Burn 18 months post planting Two Year Fire Return Interval naturally pruning and quality trees
  • 32. Planting Forest Openings Planting Density and Fire
  • 34. Conversion of Crop Fields Poorly Formed Trees – higher fertility sites? Generally Higher Stocking Rate - 605 TPA Pruning will be needed at lower TPA Ground cover to permit burning – first MAY avoid issues
  • 35. Establish Ground CCoovveerr –– BBuurrnniinngg CClluummpp ppllaannttiinngg vveerrssuuss rrooww ppllaannttiinngg
  • 36. In a Nutshell • Stoddard-Neel System is a form of UEA Management • Bimodal Distribution versus Reverse J • Existing LLP – Fire and Gaps • Works well with wildlife Values • Retains significant old growth portion • Under-planting with frequent fire – May require herbicide as site prep • Field systems require heavier stocking – Establish burnable ground cover