This document provides information on the phylum Echinodermata. It discusses the general characteristics of echinoderms and provides classifications and descriptions of key examples including Antedon (feather star), Asterias (starfish), Ophiothrix (spiny brittle star), Echinus (sea urchin), Holothuria (sea cucumber), and Cucumaria (sea cucumber). The summary describes the phylum, notes that it includes starfish, sea urchins, sea cucumbers and others, and states that examples and their characteristics are provided.
Chordata is the last phylum of kingdom Animalia.
Which is further subdivided into subphylums, divisions and classes.
The Slides shows the classification of the phylum along with the basis on which it is classified.
(includes examples along with pictures for easy understanding and memorizing)
are worm-like parasites. The clinically relevant groups are separated according to their general external shape and the host organ they inhabit. There are both hermaphroditic and bisexual species.
The definitive classification is based on the external and internal morphology of egg, larval, and adult stages.
Helminth is a general term meaning worm. The helminths are invertebrates characterized by elongated, flat or round bodies.
In flatworms or platyhelminths (platy from the Greek root meaning “flat”) include flukes and tapeworms.
Roundworms are nematodes (nemato from the Greek root meaning “thread”).
Chordata is the last phylum of kingdom Animalia.
Which is further subdivided into subphylums, divisions and classes.
The Slides shows the classification of the phylum along with the basis on which it is classified.
(includes examples along with pictures for easy understanding and memorizing)
are worm-like parasites. The clinically relevant groups are separated according to their general external shape and the host organ they inhabit. There are both hermaphroditic and bisexual species.
The definitive classification is based on the external and internal morphology of egg, larval, and adult stages.
Helminth is a general term meaning worm. The helminths are invertebrates characterized by elongated, flat or round bodies.
In flatworms or platyhelminths (platy from the Greek root meaning “flat”) include flukes and tapeworms.
Roundworms are nematodes (nemato from the Greek root meaning “thread”).
It discusses basic information regarding a hemichordate animal called Balanoglossus or Acorn worm, which is also a good connecting link between the non-chordates and chordates.
The invertebrates, or invertebrates, are animals that do not contain bony structures, such as the cranium and vertebrae. The simplest of all the invertebrates are the Parazoans, which include only the phylum Porifera: the sponges.
Parazoans (“beside animals”) do not display tissue-level organization, although they do have specialized cells that perform specific functions. Sponge larvae are able to swim; however, adults are non-motile and spend their life attached to a substratum.
Since water is vital to sponges for excretion, feeding, and gas exchange, their body structure facilitates the movement of water through the sponge. Structures such as canals, chambers, and cavities enable water to move through the sponge to nearly all body cells.
The chordates are named for the notochord: a flexible, rod-shaped structure that is found in the embryonic stage of all chordates and also in the adult stage of some chordate species.
It is located between the digestive tube and the nerve cord, providing skeletal support through the length of the body.
In some chordates, the notochord acts as the primary axial support of the body throughout the animal's lifetime.
Sponges,are pore bearing,multicellular,diploblastic animals that belong to phylum Porifera
Body of all sponges is perforated by large number of pores called ostia through which water enters Inside body and flows through a system of criss-crossing canals known as canal system
Three main types of canal systems in the order of increasing complexity are Asconoid, Syconoid and Leuconoid type.
Rotifers are microscopic aquatic animals of the phylum Rotifera. Rotifers can be found in many freshwater environments and in moist soil, where they inhabit the thin films of water that are formed around soil particles.
ORIGIN OF CHORDATES
Animal kingdom is basically divided into two sub kingdoms:
Non-chordata- including animals without notochord.
Chordata- This comprising animals having notochord or chorda dorsalis.
Chordates were evolved sometime 500 million years ago during Cambrian period (invertebrates were also began to evolve in this period) .
Chamberlain (1900) pointed out that all modern chordates possess glomerular kidneys that are designed to remove excess water from body.
It is believed that Chordates have originated from invertebrates.
It is difficult to determine from which invertebrate group the chordates were developed.
Chordate ancestors were soft bodied animals. Hence they were not preserved as Fossils.
However, early fossils of chordates have all been recovered from marine sediments and even modern protochordates are all marine forms.
Also glomerular kidneys are also found in some marine forms such as myxinoids and sharks. That makes the marine origin of chordates more believable.
Chordates evolved from some deuterostome ancestor (echinoderms, hemichordates, pogonophorans etc.) as they have similarities in embryonic development, type of coelom and larval stages.
Many theories infers origin of chordates, hemichordates and echinoderms from a common ancestor.
It discusses basic information regarding a hemichordate animal called Balanoglossus or Acorn worm, which is also a good connecting link between the non-chordates and chordates.
The invertebrates, or invertebrates, are animals that do not contain bony structures, such as the cranium and vertebrae. The simplest of all the invertebrates are the Parazoans, which include only the phylum Porifera: the sponges.
Parazoans (“beside animals”) do not display tissue-level organization, although they do have specialized cells that perform specific functions. Sponge larvae are able to swim; however, adults are non-motile and spend their life attached to a substratum.
Since water is vital to sponges for excretion, feeding, and gas exchange, their body structure facilitates the movement of water through the sponge. Structures such as canals, chambers, and cavities enable water to move through the sponge to nearly all body cells.
The chordates are named for the notochord: a flexible, rod-shaped structure that is found in the embryonic stage of all chordates and also in the adult stage of some chordate species.
It is located between the digestive tube and the nerve cord, providing skeletal support through the length of the body.
In some chordates, the notochord acts as the primary axial support of the body throughout the animal's lifetime.
Sponges,are pore bearing,multicellular,diploblastic animals that belong to phylum Porifera
Body of all sponges is perforated by large number of pores called ostia through which water enters Inside body and flows through a system of criss-crossing canals known as canal system
Three main types of canal systems in the order of increasing complexity are Asconoid, Syconoid and Leuconoid type.
Rotifers are microscopic aquatic animals of the phylum Rotifera. Rotifers can be found in many freshwater environments and in moist soil, where they inhabit the thin films of water that are formed around soil particles.
ORIGIN OF CHORDATES
Animal kingdom is basically divided into two sub kingdoms:
Non-chordata- including animals without notochord.
Chordata- This comprising animals having notochord or chorda dorsalis.
Chordates were evolved sometime 500 million years ago during Cambrian period (invertebrates were also began to evolve in this period) .
Chamberlain (1900) pointed out that all modern chordates possess glomerular kidneys that are designed to remove excess water from body.
It is believed that Chordates have originated from invertebrates.
It is difficult to determine from which invertebrate group the chordates were developed.
Chordate ancestors were soft bodied animals. Hence they were not preserved as Fossils.
However, early fossils of chordates have all been recovered from marine sediments and even modern protochordates are all marine forms.
Also glomerular kidneys are also found in some marine forms such as myxinoids and sharks. That makes the marine origin of chordates more believable.
Chordates evolved from some deuterostome ancestor (echinoderms, hemichordates, pogonophorans etc.) as they have similarities in embryonic development, type of coelom and larval stages.
Many theories infers origin of chordates, hemichordates and echinoderms from a common ancestor.
Insects, spiders, crabs, shrimp, millipedes, and centipedes are all arthropods. Arthropods have jointed feet, a segmented body, and an exoskeleton, a cuticle on the outside of their body. Arthropods have by far the greatest number of species of any animal group, at around 900,000 species
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3. PHYLLUM :- ECHINODERMATA
General Characters :-
1. They are marine animals.
2. Tribloblastic animals.
3. True coelom is present.
4. Their space is generally pentamerous.
5. The exoskeleton is formed of calcareous plates or ossicles.
6. Generally, spine are found over the body and calcareous plate below the
skin.
7. In Adult condition Body :- Radially symmetrical, but their larvae are
Bilaterally symmetrical.
8. Tube feet are present which are organ of locomotion and also helpful in
food catching & respiration.
4. 9. A water vascular system is found in them.
10. Respiration take place by branchiae.
11. Excretory organs are absent.
12. Sexes are separate.
CLASSIFICATION
• Phylum has been classified into four sub- phylum. (Marshall & Williams)
SUB-PHYLUM :- ECHINOZOA
1. Globoid and cylindrical echinoderm which do not develop arms.
2. Hydrocael forms a ring about the giving rise to meriodional
water vessels.
5. • Class 1 – Helicoplacodea
1. Free living, fusiform placoid echinozoans.
2. Plates of the body form the test.
3. Anal & Oral aperture at opposite poles.
Example :- Helicoplacus.
• Class 2 – Holothuroidea.
1. Body elongated in oral – aboral axis.
2. Skin leathery.
3. Body is long and cylindrical or pentamerous.
4. Mouth and anus are present on opposite surfaces.
5. Mouth surrounded by tentacles.
6. Tube feet are found & helpful in locomotion.
7. Sexes are separated.
6. • Sub-Class 1 – Dendrochioracea
1. Respiratory tree present.
2. Introvert presents or pharyngeal refractor muscles present.
• ORDER 1 – Dactylochirotida
1. Digitate and feebly branched tentacles.
Example :- Ypsilothuridae.
• ORDER 2 – Dendrochirotida
1. Tentacles are elongated, branched to form tree like structure.
Example :- Cucumaria.
• Sub-Class 2 – Aspidochiroatcea
1. Tentacles 10-20 shield shaped.
2. Test vestigial .
3. Tube feet present on ventral side.
7. • ORDER 1 – Aspidochiroatida
1. Large branched respiratory tree.
Example :- Holothuria.
• ORDER 2 – Elasipodida
1. Respiratory tree absent.
Example :- Elpidia, Pelagothuria.
• Sub-Class 3 – Apodacea
1. Holothurian with simple digitate tentacles.
2. Tube feet reduced or absent.
3. No spicules in the skin, no refractor muscle.
8. • Class 3 – Edriosteroidea
1. Free swimming placoid echinozoan with mouth and anus on the upper side
of the test.
2. Five ambulacra radiate from the mouth. Hinged ambulacral ossicles border
the ambulacra, which carry food groove.
3. Body discoidal, dorsovantrally flattened .
4. Extinct forms.
Example :- Edrioaster.
• Class 4 – Exhinoidea
1. Body is oval round or heart shaped without arms but with ambulacral areas.
2. Body enclosed in a test made up of calcareous plates.
3. Pedicilarie are stalked & tree – jawed.
4. Introvert or lantern present.
Example :- Sea urchin.
9. • Sub-Class 1 – Perischoechinoidea
• ORDER 1 – Exhinocytitoida , Example :- Eothuria.
• ORDER 2 – Bothricidaroida , Example :- Bothriocidaris.
• ORDER 3 – Palaechinoidea , Example :- Palaechinus.
• ORDER 4 – Cidaroida , Example :- Miocidaris.
• Sub-Class 2 – Euechinodea
• Super Order 1 – Diaematacea
• ORDER 1 – Echibothuroida
• ORDER 2 – Diadamatoida
• ORDER 3 – Padinoida
• ORDER 4 – Pygasteroida
10. • Super Order 2 – Echinancea
• ORDER 1 – Salenioida
• ORDER 2 – Hemidaroida
• ORDER 3 – Phymosomatoida
• ORDER 4 – Arbacioida
• ORDER 5 – Temnopkuroida.
• ORDER 6 – Schinoida.
• Super Order 3 – Gnathostomata
• ORDER 1 – Holectypoida
• ORDER 2 – Clypeasteroida
• Super Order 4 – Atelostomata
• ORDER 1 – Classiduloida
• ORDER 2 – Holastaoida
• ORDER 3 – Spatangoida
11. • Class 5 –
• Sub – Phylum 1 – Hc
• Includes extinct, dorsovantrally flattened echinoderms, with no signs of radial
symmetry .
• Sub – Phylum 2 – Cri
1. Body aboral, cup shaped.
2. Ambulacra are restricted to the adoral surface.
3. Sessile, atleast in any part of their life.
4. Mouth and anus both are situated on the upper side. Most of the member
are now extinct.
Example :- Antedon.
• Sub – Phylum 3 – Homalozoa
• Sub – Phylum 4 – Crinozoa
• Sub – Phylum 5 – Asterozoa
12. ANTEDON
General Characters :-
1. Commonly known as – “Feather star”
or “Sea lily”.
2. The pentamerous body is disc-shaped
with flower-like appearance.
3. The body is divisible into a central disc
or calyx and a series of 10 radiating
arms.
4. The central disc is differentiated into
convax and upwardly directed oral &
flat ventrally directed aboral surfaces.
5. The animals are highly coloured and
may be green, purple, yellow & red.
13. 6. The oral surface is covered by soft, leathery skin the oral membranes are the
tegmen in which are embedded minute calcareous plates in the form of a
continuous sheet.
7. The mouth is the situated in the central of the oral disc and is surrounded by
five small triangular oral valve.
8. In the between of oral valves radiate five narrow, open and ciliated food
grooves.
9. The anus is situated near the mouth in one inner ambulacrum on a
prominence.
10. A series of five movable long slender arms often branched and bearing
small alternating branches , the pinnules originated from central body.
11. Ambulacral groove ciliated, open runs from mouth on the oral surface of
arms & pinnules their tips.
12. Tube feet reduced generally tentacle like present along the edge of
ambulacral groove.
14. 10. Madreporite is absent.
11. The aboral surface is partly covered by tagmen and partly by a large
flat, pentagonal central sketelal plate , the centro-dorsal ossicles.
12. On the aboral surface are slender curved and cylidrical appendages
known as cirri meant for gripping the substratum.
13. The sexes are separate but there is no sexual dimorphism .
14. The development is indirect with pentacrinoid larva with jointed stalk.
15. They have the power of regeneration .
15. ASTERIAS
General Characters :-
1. It is commonly known as ‘stare fish’.
2. Dorsoventrally flattened star –shaped ,
pentagonal body .
3. Body is divisible into a central disc and five
radiating arms.
4. The animal are usually bright coloured yellow,
brown or orange.
5. The body is divisible into a flat , downwardly
directed and light coloured oral surface and a
convex, upwardly directed and light coloured
oral surface and a convex , upwardly directed
and much darker aboral surface.
16. 6. The pentagonal mouth lies in the centre of oral surface and surrounded by
oral spines.
7. The narrow ambulacral grooves arise from the five corners of mouth and
extend through the middle of each arm upto the tip.
8. Each ambulacral groove bears four rows (two double rows ) of soft thin
walled tubular retractile tube – feet with sucker – like extremities.
17. OPHIOTHRIX
General Characters :-
1. It is commonly known as ‘ spiny brittles
star’.
2. Brilliantly coloured with phosphorocence.
3. The body is divisible into a small rounded
central disc and five greatly developed
fragile and flexible arms sharply marked
off from the disc.
4. The arms are covered by plates or shields
fringed with spines on all sides.
5. The mouth is on the oral surface and is
with five movable plates serving as jaws.
18. 6. Only the madreporite is present near the mouth on the oral surface.
7. The oral surface also bears five shields and podial pores.
8. Ambulacral grooves and pedicellariae are absent.
9. The tube feet without suckers are present on the lower plates of arms.
10. When distributed it readily breaks of one or more arms , which are
later regenerated.
11. The anus is absent.
12. The sexes are separate.
13. The development is indirect with pelagic ophiopluteus larva.
14. The genital bursac are present at the base of each arm.
15. They have great power of regeneration or autotomy.
19. ECHINUS
General Characters :-
1. It is commonly known as ‘sea urchin.’
2. The hared and prickly body is globular
in shape, slightly pentagonal in
equatorial outline with rounded
angles .
3. The body is more or less flattened at
the two poles, forming distinct oral .
4. The colour of the body may be brown,
black , purple, green, white or red.
5. The body is enclosed in a shell or
corons formed of closely fitting firmly
united calcaeous plates.
20. 6. Body surface is covered by numerous cylindrical , solid , spines.
7. Among spines are found stalked pedicellarise with three jaws and tube feet
arranged in five double rows.
8. The more flattened oral surface bear rounded central mouth surrounded by
thin, soft flexible peristome.
9. Through mouth project ten hard , white calcareous teeth from Aristotle’s
lantern.
10. At the outer edge of peristome lie ten little boshy, thin walled and branching
gills one pair opposite to each inter ambulacral area.
11. The aboral, end is somewhat dome-shaped and lie opposite to oral end.
12. At the aboral end lies a much smaller aperture , the anus , surrounded by
periproct.
13. The sexes are separate and there is no sexual dimorphism.
14. Fertilization is external.
21. HOLOTHURIA
General Characters :-
1. It is commonly known as ‘ cucumber’.
2. Body is large , cylindrical , elongated ,
vermiform and bilaterally symmetrical
with mouth and anus at opposite
ends.
3. The colour of the animal is brown or
yellowish and often reddish blue.
4. The animal rests on ground with its
long axis parallel to the surface.
5. The anus is placed at the extreme
aboral end.
22. 6. The retractile and locomotory podia are distributed all over the body
7. The surface of the animal is divided into five ambulacral and five
interambulacral zones which are not recognizable externally.
8. Body wall leathery and muscular with exoskeleton in the form of calcareous
ossicles.
9. The mouth is at the oral end and is surrounded by peristome and 20 or 30
pellate tentacles.
10. The respiratory tree is well developed .
11. The Cuvierian organ present.
12. Water vascular system is well developed.
13. The madreporite is internal.
14. The sexes are separate
15. The development is indirect with curricular larva.
23. CUCUMARIA
General Characters :-
1. It is commonly known as ‘ sea
cucumber’.
2. Body is elongated and cylindrical with
terminal mouth and anus opening at
opposite ends.
3. The anterior oral end is somewhat
thicker that the posterior aboral end.
4. The oral and aboral axis greatly
elongated.
5. Body is pentamerous , having five
longitudinal bands or ambulacra, each
is provided with double row of
locomotary podia.
24. 6. Three of these ambulacral areas are ventral and provided with numerous
tube feet with suckers.
7. The other ambulacral are dorsal and are with smaller tube feet lacking
sucker.
8. Mouth is surrounded by a lip thin, darkly pigmented peristomial
membrane and a circlet of ten dendritic , tree-like tentacles , two belonging
to each ambulacrum.
9. Each tentacle is enlarged, specially modified tube – feet of highly
sensitive and contractile nature.
10. The smooth , thin and collar like body at the base of tentacles is known
as introvert.
11. Two respiratory trees are present.
12. Water vascular system is well developed.