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Preparing	Scien,fic	Papers	
for	Publica,on	
A.	Townsend	Peterson	
University	of	Kansas
Overall	Publishing	Process	
•  Write	the	paper	
•  Edit/proof	the	manuscript	
•  Format	for	the	journal	
•  Submit	to	the	journal	
•  Editor	decides	iniAally	and	sends	out	to	review	
•  Reviewers	provide	suggesAons	and	criAcisms	
•  Editor	decides	whether	to	accept	
–  …	can	repeat	several	Ames	
•  Author	revises	
•  Final	acceptance	
•  Proofs	sent	to	author	
•  Author	corrects	
•  Author	signs	copyright	transfer	agreement	
•  PublicaAon
WRITING	–	STRUCTURE	AND	ORDER
Scoping	the	Paper	
Short	and	Sweet	
•  Advantages	
–  Easy	to	write	
–  Simple	message	
–  “More”	publicaAons	on	your	
CV	
•  Disadvantages	
–  Dilutes	the	message	
–  Reader	has	to	search	for	the	
whole	story	across	many	
papers	
–  Less	impact	per	publicaAon	
Long	and	Complete	
•  Advantages	
–  Whole	story	is	there	in	one	
place	
–  Greater	potenAal	impact	
•  Disadvantages	
–  More	difficult	to	organize	and	
write	
–  Message	can	get	complex	
–  Fewer	publicaAons	on	your	
CV
Picking	a	Journal	
•  With	advisors	and	colleagues,	decide	on	the	opAmal	
outlet	for	your	paper	
•  Aim	too	high:	
–  Waste	your	Ame	
–  Waste	reviewers’	and	editors’	Ame	
•  Aim	too	low:		
–  Less	credit	for	you	
–  Less	aUenAon	to	your	work	
•  Journal	characterisAcs	
–  Subject	focus	
–  Previous	papers	published	
–  Impact	factor	
–  Access	poliAcs	(see	the	end	of	this	presentaAon)
WriAng	–	Structure	and	Order	
•  Suggested	order	of	wriAng:	
–  Figures	/	Tables	
–  Methods	/	Results	
–  IntroducAon	/	Discussion	
–  Abstract	
•  Last	paragraph	of	IntroducAon	should	begin	“This	
paper	…”	and	should	state	clearly	the	purpose	of	
the	paper	
•  Length	–	care	not	to	be	overly	long	in	
IntroducAon	and	Discussion	
•  RepeAAon	–	no	need	to	repeat	parts	of	
IntroducAon	in	Discussion,	or	parts	of	Results	in	
Discussion
TECHNOLOGY	ASSISTS
Technology	Assists	–	Google	Docs
Technology	Assists	–	Google	Groups
Technology	Assists	-	EndNote
Free	Technology	Assists	-	Mendeley	
Another	possibility:	hUp://www.zotero.org/	-	reference	manager,	also	free
WORDS	TO	AVOID
Words	to	Avoid	I	
•  While:	Don’t	use	this	word	to	mean	although,	whereas,	and,	or	but,	and	don’t	
use	it	where	only	a	semicolon	is	necessary.	While	means	“during	the	Ame	that”	
and	should	be	used	to	link	simultaneous	occurrences	in	instances	in	which	
simultaneity	is	the	point.	
•  E.g.,	i.e.:	The	abbreviaAon	e.g.	means	“for	example”	(exempli	gra4a)	and	
introduces	an	illustraAve	instance	or	a	short	list	of	names	or	other	items.	The	
abbreviaAon	i.e.	means	“that	is”	(id	est)	and	introduces	a	repeAAon	in	different	
words	of	the	ideas	just	discussed	
•  Compose,	comprise:	Comprise	expresses	the	relaAon	of	the	larger	to	the	
smaller,	not	the	other	way	around	(think	of	comprise	as	meaning	to	embrace	or	
take	in).	The	whole	comprises	the	parts;	the	whole	is	composed	of	its	parts.	The	
parts	compose	the	whole	and	are	comprised	in	it.	Do	not	use	comprised	of.	
•  Because,	since,	as:	Because	is	the	most	specific	of	the	conjuncAons	used	to	
express	reason	or	cause.	It	always	indicates	an	unequivocal	causal	relaAonship.	
Since	is	oaen	a	weak	form	of	because.	It	also	contains	a	noAon	of	duraAon	over	
Ame	that	because	does	not.	Use	since	when	the	meaning	of	what	follows	it	is	
implied	by	what	precedes	it.	Using	as	to	mean	since	or	because	is	always	feeble.	
It	makes	whatever	follows	sound	trivial.		
•  Although,	though:	In	most	uses	these	words	are	interchangeable,	but	it	may	be	
best	to	begin	clauses	with	although,	since	it	is	more	emphaAc,	and	to	use	
though	to	connect	elements	within	a	clause.	(Although	intelligent,	he	could	not	
apply	himself.	She	was	born	of	poor	though	well-educated	parents.)
Words	to	Avoid	II	
•  Very:	This	word	is	almost	always	completely	unnecessary	in	sentences.	
That	is,	the	statements	“values	of	the	index	were	very	high”	and	“values	
of	the	index	were	high”	communicate	the	same	informaAon	without	the	
hyperbole.	
•  There	is/there	are:	This	sort	of	construcAon.	E.g.,	“there	are	thirty	birds	
sicng	on	the	wire”	can	be	replaced	by	a	more	direct	statement	…	“thirty	
birds	are	sicng	on	the	wire.”		
•  This/these/those	as	nouns:	“This,”	“these,”	and	“those”	are	modifiers,	
and	should	not	be	used	as	nouns.		
•  Passive	versus	ac,ve	verbs:	Most	(but	not	all)	uses	of	the	passive	form	
of	verbs	can	and	should	be	transformed	into	acAve	forms.	For	instance	
“this	bird	was	processed	by	XJV	approaches”	can	be	restated	as	“we	
used	XJV	approaches	to	process	this	bird.”	
•  Et	al.:	This	construcAon	is	derived	from	the	LaAn	et	altera	for	“and	
others.”	It	is	frequently	misused	in	a	variety	of	uncareful	ways:	“Johnson	
et	al	2004”	(forgecng	the	abbreviaAon	of	altera),	“Johnson	et	all	2004,”	
etc.
Words	to	Avoid	III	
•  Due	to:	Most	uses	of	this	construcAon	should	be	replaced	with	“owing	to.”	
“Due	to”	implies	that	some	quanAty	should	be	paid	to	or	is	owed	to,	whereas	
the	correct	meaning	is	one	of	causaAon.	
•  Spaces	aPer	period	and	colon:	Although	tradiAonal	typesecng	(and	probably	
many	of	your	composiAon	classes)	required	two	spaces	aaer	periods	and	
colons,	modern	typesecng	is	not	in	need	of	these	extra	spaces.	Most	
importantly,	however,	is	TO	BE	CONSISTENT	…	
•  Bri,sh	versus	US	spelling:	Perhaps	neither	is	universally	correct,	but	one	or	the	
other	should	be	used	consistently	throughout	the	enAre	document.	Ideally,	use	
the	style	demanded	by	the	journal	for	which	you	are	wriAng.	
–  Centre/center					Theatre/theater	
–  Behaviour/behavior	
–  Acknowledgement/acknowledgment	
–  Whilst/while					Analyse/Analyze	
•  Hyphenated	compound	modifiers:	Hyphens	connect	the	words	of	a	compound	
modifier	that	come	before	the	word	being	modified.	Hyphens	are	not	used	this	
way	with	compound	parts	ending	in	-ly	or	made	up	of	proper	nouns	or	proper	
adjecAves.		“He	is	a	well-respected	man”,	“That	man	is	well	respected”,	“That	
was	a	badly	punctuated	sentence”,	and	“The	South	American	rain	forest	is	
home	to	hundreds	of	species	of	hummingbirds”	are	correct.
Words	to	Avoid	IV	
•  Where:	Where	implies	a	locality,	posiAon,	or	direcAon.		Do	not	use	for	
“in	which”	or	“for	which”.		“PredaAon	on	small	prey,	where	many	
individuals	must	be	captured	per	dive,	is	limited	to	shallow	depths.”		
Replace	“where”	with	“in	which”.	
•  Predate:	Predate	means	to	“date	in	advance”.		The	verb	derived	from	
“predaAon”	is	“depredate”.	
•  Monophyle,c	clade:	A	clade	is,	by	definiAon,	monophyleAc.	Save	space	
and	omit	this	redundancy.	
•  Data:	The	word	data	is	plural.	The	singular	is	datum.		
•  The:	The	is	overused.	When	ediAng,	think	about	whether	or	not	the	
placement	of	every	“the”	is	necessary.		
•  Split	infini,ves:	To	blatantly	divide	the	word	“to”	from	the	verb	that	it	
accompanies	by	an	adverb	is	an	ugly	construcAon.	
•  Hyphens	and	dashes:	Hyphens	are	used	within	text	to	build	compound	
modifiers	(e.g.,	“early-career	professionals),	n-dashes	are	used	to	
indicate	ranges	of	numbers	(e.g.,	4–5),	m-dashes	are	used	within	
sentences	(e.g.,	“I	ate—more	than	anything—because	I	was	hungry”).
FIGURES
Figures	-	Technology	
•  Save	AI	version	(or	GIS	original	or	…)	
•  Export	JPG	version	for	review
Color	versus	Black-and-white	
•  Color	is	much	beUer	for	interpretability	
•  Can	use	color	for	review,	and	subsAtute	B&W	
for	publicaAon	
•  Can	use	color	in	electronic	publicaAon	version,	
and	B&W	in	hard-copy	version	(if	there	is	one)	
•  However,	print	in	B&W	to	make	sure	it	is	
interpretable	without	color,	in	case	reviewers	
do	not	print	in	color	…	you	want	them	to	be	
able	to	read	the	figure!
Figures	–	Colors	and	Colorblindness	
•  hUp://colorbrewer2.org/
Structure	–	Making	Readers	Refer	to	CapAons	
Good	
	
	
	
	
																								Bad
Figures	–	Format	
•  At	end	of	manuscript	
•  Figure	Legends	page	
•  Then	one	figure	per	page
TABLES
Tables	–	Try	Not	To	
•  Minimal	use	of	tables	is	best		
•  Most	informaAon	in	tables	can	be	expressed	
beUer	in	a	visual,	eyecatching	figure	
•  Readers	tend	not	to	pay	much	aUenAon	to	the	
contents	of	tables	
•  Tables	take	up	lots	of	space	
•  Consider	posAng	the	full	data	set	as	part	of	
the	publicaAon,	or	on	a	well-documented	data	
site
Tables	-	Format	
Header	informaAon	
	
	
Body	of	table	
Table	1.	The	capAon	goes	here,	immediately	above	the	
table	itself
Tables	–	Placement	
•  At	end	of	manuscript	
•  Aaer	figures	
•  Pages	not	numbered	
•  One	table	per	page
PROOFING	AND	EDITING
Proofing	and	EdiAng	-	Example	
Scale	effects	such	as	distribuAon	amplitude	and	spaAal	
resoluAon	may	also	introduce	contradictory	results	when	
interpreAng	LPSR	(Rahbeck	2006).	For	example,	in	
smaller	geographic	areas,	monotonic	trends	in	species	
richness	are	usually	observed,	whereas	across	larger	
areas,	however,	hump-shape	paUerns	are	generally	
observed	(Rahbeck,	2006;	Sánchez-Cordero,	2001).	
SpaAal	resoluAon	can	also	affect	paUerns	of	species	
richness:	studies	involving	too	coarse-grained	resoluAon	
may	produce	misleading	results	by	registering	high	
richness	at	localiAes	where,	in	fact,	low	numbers	of	
species	occur,	averaging	over	diverse	distribuAonal	
phenomena.
Proofing	and	EdiAng	-	Example	
Scale	effects	such	as	range	size	distribuAon	amplitude	and	
spaAal	resoluAon	may	also	introduce	contradictory	results	
when	interpreAng	LPSR	(Rahbeck,	2006).	For	example,	in	
smaller	geographic	extentsareas,	monotonic	trends	in	
species	richness	are	usually	observed,	whereas;	across	
larger	areas,	however,	hump-shape	paUerns	are	generally	
observed	(Rahbeck,	2006;	Sánchez-Cordero,	2001).	SpaAal	
resoluAon	can	also	affect	species	richness	paUerns	of	
species	richness:	studies	at	overly	involving	too	coarse-
grained	resoluAon	may	produce	misleading	results	by	
finding	registering	high	richness	at	‘localiAes’	where,	in	fact,	
lower	numbers	of	species	occur.,	The	difference	results	
from	averaging	over	diverse	distribuAonal	phenomena.
CITING	OTHER	AUTHORS
GeneraliAes	
•  Cite	other	studies	to	place	your	work	in	the	
context	of	work	already	done	
•  Avoid	self-citaAons,	unless	your	work	is	the	key	
and	best	citaAon	for	the	point	being	made	
•  Cite	others’	work	broadly	and	fairly:	
– Cover	what	is	‘out	there’	in	the	literature	
– Don’t	play	favorites	
– Try	not	to	have	enemies,	or	at	least	try	to	“cite	thy	
enemy”	
•  CitaAons	are	not	rewards	or	symbols	of	friendship	
or	alliance
LITERATURE	CITED
GeneraliAes	
•  Literature	Cited.	This	sector	of	scienAfic	
wriAng	is	almost	universally	rife	with	errors	
and	inconsistencies.	Here	are	a	few:	
– CapitalizaAon	aaer	a	colon	in	journal	Atles	–	
either	way	is	acceptable	and	journals	vary	in	their	
preferences,	BUT	pick	one	style	and	be	consistent	
– Spaces	between	iniAals	in	authors’	names	–	again,	
journals	vary	in	their	preferences,	but	be	
consistent	
– Inclusion	of	issue	number	–	be	consistent
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
•  BIRDLIFE	INTERNATIONAL.	2012.	IUCN	Red	List	for	birds.	Downloaded	from	hUp://www.birdlife.org	on	
03/07/2012.	
•  BROWN,	R.	M.,	AND	A.C.	DIESMOS.	2002.	ApplicaAon	of	lineage-based	species	concept	to	oceanic	island	frog	
populaAons:	The	effects	of	differing	taxonomic	philosophies	on	the	esAmaAon	of	Philippine	biodiversity.	
Silliman	Journal	42,	133–162.	
•  CLEMENT,	M.,	POSADA,	D.,	AND	K.A.	CRANDALL.	2000.	TCS:	a	computer	program	to	esAmate	gene	genealogies.	
Molecular	Ecology	9:1657–1659.		
•  COLLAR,	N.J.	2006.	A	parAal	revision	of	the	Asian	babblers,	Timaliidae.	Forktail	22:85–112.	
•  COLLAR,	N.	J.	AND	C.	ROBSON.	2007.	Family	Timaliidae	(babblers).	70–291.	In	Handbook	of	the	birds	of	the	
world.	Volume	12.	Picathartes	to	4ts	and	chickadees.	Del	Hoyo,	J.,	A.	EllioU,	and	D.	A.	ChrisAe.	Editors.	Lynx	
Edicions.	Barcelona,	Spain.		
•  COYNE,	J.A.	AND	T.D.		PRICE.	2000.	LiUle	evidence	for	sympatric	speciaAon	in	island	birds.	Evolu4on,	54,	2166–
2171.		
•  DE	ROEVER,	J.	1990.	Notes	on	the	Luzon	Wren-Babbler.	Bull.	Oriental	Bird	Club	12:33-–34.	
•  DIAMOND,	J.M.	1977.	ConAnental	and	insular	speciaAon	in	Pacific	landbirds.	Systema4c	Zoology,	26,	263–
268.		
•  DUPONT,	J.E.	1971.	Notes	on	Philippine	birds	(No.	1).	Nemouria	3.	
•  DUPONT,	J.E.	1971.	Philippine	birds.	Delaware	Museum	of	Natural	History	Monographs	Series	2.	
•  HEANEY,	L.	R.,	1986.	Biogeography	of	mammals	in	SE	Asia:	esAmates	of	rates	of	colonizaAon,	exAncAon	and	
speciaAon.	Biological	Journal	of	the	Linnean	Society	28:	127–165.	
•  HIJMANS,	R.J.,	CAMERON,	S.E.,	PARRA,	J.L.,	JONES,	P.G.,	AND	A.	JARVIS.	2005.	Very	high	resoluAon	interpolated	
climate	surfaces	for	global	land	areas.	InternaAonal	Journal	of	Climatology	25:1965–1978.		
•  IUCN	STANDARDS	AND	PETITIONS	SUBCOMMITTEE.	2010.	Guidelines	for	Using	the	IUCN	Red	List	Categories	and	
Criteria.	Version	8.1.	Prepared	by	the	Standards	and	PeAAons	SubcommiUee	in	March	2010.	Downloadable	
from	hUp://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf	
•  LAMBERT,	F.L.	1993.	Some	key	sights	and	significant	records	of	bird	from	the	Philippines	and	Sabah.	Bird	
ConservaAon	InternaAonal	3:281–297.	
•  GONZÁLEZ,	L.A.S.,	OLIVEROS,	C.,	PUNA,	N.	AND	R.G.	MOYLE.	2010.	Nests,	Nest	Placement,	and	Eggs	of	Three	
Philippine	Endemic	Birds.	Wilson	Journal	of	Ornithology	122:587–591.	
•  GOODMAN,	S.	M.	AND	P.	C.	GONZALES.	1990.	The	birds	of	Mt	Isarog	NaAonal	Park,	southern	Luzon,	Philippines,	
with	parAcular	reference	to	alAtudinal	distribuAon.	Fieldiana	Zoology	60:1–39.
COVER	LETTERS
Minimalist	Version	
To	The	Editor:	
	
Please	find	aUached	to	this	leUer	a	manuscript	enAtled	This	Is	
The	Title	of	My	Manuscript,	for	consideraAon	for	publicaAon	
in	Your	Journal.	This	manuscript	is	the	intellectual	product	
only	of	myself	and	my	coauthors,	and	is	not	currently	
submiUed	for	publicaAon	with	any	other	journal.	
	
I	hope	that	this	manuscript	will	prove	acceptable	for	
publicaAon	in	your	journal.	Please	do	not	hesitate	to	contact	
me	should	you	need	any	further	informaAon.	
	
Sincerely,
Should	You	Argue	For	Your	MS	In	Cover	LeUer?	
•  For	high-end	journals	(Science,	Nature,	PNAS),	it	
is	necessary	…	pre-review	editorial	decisions	may	
depend	on	it!	
•  For	‘normal’	journals,	it	is	a	maUer	of	choice	…	
many	people	do	include	a	paragraph	of	why	the	
paper	is	important;	I	do	not	
•  Be	VERY	careful	not	to	overstate	the	importance	
of	the	paper	or	the	impact	of	the	results
CHOOSING	AND	RECOMMENDING	
REVIEWERS
Reviewers	
•  Many	journals	permit	suggesAons	regarding	
appropriate	reviewers	for	a	manuscript	
•  These	suggesAons	may	be	opAonal	or	may	be	
required,	depending	on	the	journal	
•  The	editor	may	or	may	not	follow	your	
suggesAons
Suggested	Reviewers	
•  Ideally,	reviewers	will	have	experAse	and	
experience	in	the	area	of	the	paper	
•  Reviewers	should	not	have	any	conflict	of	
interest	with	any	of	the	authors	
•  Do	not	suggest	reviewers	who	are	close	
associates,	friends,	relaAves	(!),	or	involved	in	
the	project	…	quite	simply,	it	is	not	
appropriate,	and	it	is	unfair	to	the	editors	to	
make	them	have	to	discover	these	
connecAons	without	the	authors’	honesty
Non-suggested	Reviewers	
•  Occasionally,	conflicts	of	opinion,	personal	
disagreements,	nonfuncAonal	personaliAes,	and	
other	issues	can	cloud	the	objecAvity	of	peer	review	
•  Authors	can	request	editors	to	remove	such	
individuals	from	consideraAon	as	reviewers—either	
in	the	cover	leUer	or	via	lists	on	web	pages:	
–  In	cover	leUer:	“Based	on	past	experiences,	in	which	
XXXXX	showed	lack	of	objecAvity	in	peer	review	of	
manuscripts	by	myself	and	my	coauthors,	I	would	
respec{ully	that	XXXXX	not	be	considered	as	a	reviewer	
for	this	manuscript.”	
•  Editors	may	nonetheless	request	reviews	from	such	
individuals,	but	may	consider	the	resulAng	review	in	
light	of	the	knowledge	that	a	conflict	is	at	least	
perceived	to	exist
AUTHORSHIP
Reasons	For	Authorship	
•  Wrote	the	paper	
•  Provided	criAcal	intellectual	input	to	ideas	or	
methods	
•  Worked	hard	gathering	or	analyzing	data	
•  Led	the	lab	group	
•  Your	advisor	
•  In	sum,	should	be	people	who	made	significant	
contribuAons	to	the	overall	effort
RESPONSES	TO	REVIEWS
First	–	Understand	the	Response	
Thank	you	very	much	for	submicng	your	manuscript	
"Uses	and	misuses	of	bioclimaAc	envelope	modelling"	for	
review	by	Ecology.	The	reviewers	and	I	appreciate	the	
work	you	have	accomplished.	Based	on	the	reviews,	we	
will	not	be	able	to	accept	this	manuscript	for	publicaAon	
at	this	Ame.	We	would	be	willing	to	consider	a	much-
revised	version,	based	on	the	review	comments.	Indeed	
conceptual	papers,	as	this	one,	are	difficult	to	write.	Here	
both	reviewers	stressed	that	the	arguments	and	balance	
view	of	the	debate	should	be	beUer	presented	(not	as	
vague).	Both	reviewers	were	very	generous	in	their	
construcAve	comments	and	a	revised	version	could	make	
a	great	contribuAon.
In	General	
•  Whenever	possible,	say	“OK”	
•  When	not	possible	to	say	OK,	then	be	reasonable,	
make	some	changes	
•  When	you	really	disagree,	then	state	a	clear	
reasoning	as	to	WHY	
•  Finding	reviewers	is	a	hard	task	for	editors	
•  Your	objecAve	…	the	editor	accepts	the	paper	
without	further	review
Ideal	Responses	
Line	28.	"Offer	a	bridge	across	scales".	Statement	is	not	explained	or	
supported	in	any	way.	Should	be	removed.	
		
•  Done.	
		
Line	563.	"microscopic"	is	obviously	incorrect.	Do	you	mean	"small	areas"	
		
•  Changed.	
		
I	find	that	the	paper	is	very	long	relaAve	to	the	points	it	makes.	Many	of	the	
secAons	could	be	shorthened	considerably.	This	secAon	could	be	much	
shorter.	
		
•  The	manuscript	has	indeed	been	shortened	quite	a	bit,	with	2	major	
sec4ons	either	removed	en4rely	or	shortened	massively.
When	You	Disagree
When	You	Disagree
CHECKING	PROOFS
“Proof”	Reading	
•  Last	check	before	publicaAon	
•  What	problems	and	errors	are	not	detected	
are	your	fault,	not	the	journal’s	
•  Must	check	enAre	proof	in	minute	detail	
•  Special	aUenAon	to	equaAons,	tables,	
literature	cited	…	anywhere	that	
concentraAon	is	hard	
•  Some	people	even	read	backward	…	!
COPYRIGHT	TRANSFER	
AGREEMENTS	AND	OPEN	ACCESS
Copyright	is	A	Bundle	of	Rights	
84	
Rights	are	held	by	the	OWNER	of	the	work	(not	necessarily	the	
creator/author).		Owners	may:	
	
• Reproduce	-	make	copies	of	their	works	publicly	or	privately.	
• Adapt	-	prepare	addiAonal	works	derived	from	their	copyrighted	
work,	(aka,	derivaAve	works).	
• Distribute	-	Disseminate	copies	of	their	works,	to	the	public	by	sale	
or	other	transfer	of	ownership,	or	by	rental,	lease,	lending.		
• Perform	-	Perform	their	work	publicly	(at	locaAon	open	to	the	
public	or	to	persons	other	than	close	family	and	social	
acquaintances).	
• Display	-	Display	their	work	publicly	(applies	to	all	works	except	
sound	recordings	and	architectural	works).	
From,	SecAon	106	of	1976	Copyright	Act	and	see,	hUp://www.copyright.gov/circs/circ1.pdf
Why	Open	Access???
Conceive	
Research	
Idea	
Submit	
Paper	
Write	
Paper	
Do	
Research	
Secure	
Research	
Funding	
Paper	
Published	
Editor	
Accepts	
Reviewers	
Consider	
Editor	
Receives	
Subscrip,on	
Fees	
Access	
Charges	
Page	
Charges
How	Big	is	the	Access	Problem?		
An	Experiment	for	EEB	Scholarship	
•  Take	10	EEB	papers,	each	from	a	different	
journal	
•  Send	full	bibliographic	record	for	each	to	50	
colleagues	around	the	world	
•  Ask	each	colleague	to	go	to	his/her	office,	and	
to	aUempt	to	obtain	access	to	the	enAre	text	
of	the	publicaAon
How	Big	is	the	Access	Problem?		
An	Experiment	for	EEB	Scholarship	
•  Michael	J.	Farabee,	Edith	L.	Taylor,	Thomas	N.	Taylor,	CorrelaAon	of	permian	and	triassic	palynomorph	
assemblages	from	the	central	TransantarcAc	Mountains,	AntarcAca,	Review	of	Palaeobotany	and	Palynology,	
65:257-265.	
•  Claire	Horner-Devine,	M.,	Leibold,	M.	A.,	Smith,	V.	H.,	Bohannan,	B.	J.	M.	2003.	Bacterial	diversity	paUerns	
along	a	gradient	of	primary	producAvity.	Ecology	LeUers	6:613-622.	
•  Rafe	M.	Brown,	Jimmy	A.	McGuire	and	Arvin	C.	Diesmos.	2000.	Status	of	some	Philippine	frogs	referred	to	
Rana	evereY	(Anura:	Ranidae),	descripAon	of	a	new	species,	and	resurrecAon	of	Rana	igorota	Taylor	1922.	
Herpetologica	56:81-104.	
•  Billings,	S.	A.	2008.	Biogeochemistry:	Nitrous	oxide	in	flux.	Nature	456:888-889.	
•  Harald	Schneider,	Alan	R.	Smith,	Raymond	Cranfill,	Terri	J.	Hildebrand,	Christopher	H.	Haufler,	Tom	A.	Ranker.	
2004.	Unraveling	the	phylogeny	of	polygrammoid	ferns	(Polypodiaceae	and	GrammiAdaceae):	exploring	
aspects	of	the	diversificaAon	of	epiphyAc	plants.	Molecular	PhylogeneAcs	and	EvoluAon	31:1041-1063.		
•  Neal	Woodman,	Robert	M.	Timm,	Norman	A.	Slade	and	Terry	J.	Doonan.	1996.	Comparison	of	traps	and	baits	
for	censusing	small	mammals	in	Neotropical	lowlands.	Journal	of	Mammalogy	77:274-281.		
•  Leonard	Krishtalka	and	Philip	S.	Humphrey.	2000.	Can	natural	history	museums	capture	the	future?	BioScience	
50:611–617.	
•  Bryan	L.	Foster,	Timothy	L.	Dickson.	2004.	Grassland	diversity	and	producAvity:	The	interplay	of	resource	
availability	and	propagule	pools.	Ecology	85:1541-1547.	
•  Paul	O.	Lewis,	Mark	T.	Holder,	and	Kent	E.	Holsinger.	2005.	Polytomies	and	Bayesian	phylogeneAc	inference.	
SystemaAc	Biology	54:241-253.	
•  Lena	C.	Hileman,	Jens	F.	Sundstrom,	Amy	LiU,	Meiqin	Chen,	Takudzwa	Shumba,	and	Vivian	F.	Irish.	2006.	
Molecular	and	phylogeneAc	analyses	of	the	MADS-box	gene	family	in	tomato.	Molecular	Biology	and	EvoluAon	
23:2245-2258.
Percent	Success	
Note	that:	
-	only	2	colleagues	achieve		
				100%	success	
-	no	US	colleague	had	100%		
				success	
-	access	can	be	quite		
				unsaAsfactory,	even	in	the		
				US
How	Available	are	Different	Journals?	
i.e.,	percent	success	of	colleagues	in	accessing	papers	
Journal	
Success		
rate	
Systematic	Biology	 91.18	
Molecular	Biology	and	Evolution	 91.18	
Ecology	 73.53	
Nature	 70.59	
Ecology	Letters	 64.71	
BioScience	 64.71	
Molecular	Phylogenetics	and	Evolution	 55.88	
Journal	of	Mammalogy	 44.12	
Review	of	Palaeobotany	and	Palynology	 41.18	
Herpetologica	 41.18
ATP	Opinion	
Academic	publishing	has	been	
“bought	out”	by	an	opportunis4c	
commercial	publishing	world,	and	
is	fast	becoming	inaccessible	to	the	
academic	community,	and	
par4cularly	to	the	global	scholarly	
community.
What	Is	Open	Access	(from	Peter	Suber)?	
•  Open-access	(OA)	literature	is	digital,	online,	free	of	
charge,	and	free	of	most	copyright	and	licensing	
restricAons.		
•  OA	is	possible	thanks	to	the	internet	and	consent	of	
the	copyright-holder.	
•  Scholarly	journals	do	not	generally	pay	authors,	who	
can	therefore	consent	to	OA	without	losing	revenue.		
•  OA	is	compaAble	with	peer	review;	all	major	OA	
iniAaAves	for	scienAfic	and	scholarly	literature	insist	on	
its	importance.		
•  OA	literature	is	not	free	to	produce;	the	goal	is	to	
make	scholarly	literature	costless	…	are	there	are	
beUer	ways	to	pay	the	bills	than	by	charging	readers	
and	creaAng	access	barriers.
SOLUTION	#1	–	PUBLISH	IN	OPEN	
ACCESS	JOURNALS
Open	Access	Journals	
•  Open	access	journals	have	removed	the	
“subscripAon	barrier”	to	access	to	journal	
scholarship	
•  As	with	closed-access	journals,	they	vary	in	
quality	and	editorial	policies	
•  Open	access	journals	have	the	same	potenAal	
to	provide	peer	review	
•  Many	open	access	journals	now	with	high	
circulaAon	and	impact
SoluAon	#1	–	OA	Journals
SOLUTION	#2	-	JOURNAL	SELECTION	…	
WORK	WITHIN	THE	LIMITS
SoluAon	#2	–	Work	Within	the	Limits
SoluAon	#2	–	Work	Within	the	Limits
SoluAon	#2	–	Work	Within	the	Limits
SOLUTION	#3	–	CHANGE	THE	LIMITS:	
THE	OA@KU	EXAMPLE
KU	Example:	
write	on	the	copyright	agreement:	
“Subject	to	
aeached	
addendum”
Addendum	to	PublicaAon	Agreement
Addendum	to	PublicaAon	Agreement
Or	Simply	Edit	the	CTA:	
If	the	publisher	looks	at	your	edits	and	says	
“NO!”	…	you	can:	
– Sign	the	original	agreement	and	the	paper	is	
published	
– Debate/negoAate	with	the	publisher	about	terms	
– Withdraw	the	paper(!)
Copyright	Transfer	Summary	
•  Access	is	a	major	problem	for	scholarship	
•  Open	access	journals	are	building,	but	are	not	
sufficiently	abundant	or	high-enough	impact	to	
sustain	scholarly	publishing	
•  Scholars	can	“select”	journals	based	on	access,	if	
they	choose	
•  InsAtuAons	can	pass	policies	regarding	open	
access	
•  Scholars	can	amend	and	debate	the	copyright	
transfer	agreements
OA	Key	Internet	Resources	
•  SHERPA/RoMEO	–	summary	of	OA	policies	for	
each	journal	and	publisher:	
hUp://www.sherpa.ac.uk/romeo/		
•  ROARMAP	–	summary	of	OA	policies	at	
insAtuAons	and	funding	agencies:	
hUp://www.eprints.org/openaccess/policysignup/			
•  KU	ScholarWorks	–	KU’s	OA	repository:	
hUp://kuscholarworks.ku.edu/		
•  Harvard	Office	of	Scholarly	Communica,on	–	
hUp://osc.hul.harvard.edu/osc.php
Overall	Publishing	Process	
•  Write	the	paper	
•  Edit/proof	the	manuscript	
•  Format	for	the	journal	
•  Submit	to	the	journal	
•  Editor	decides	iniAally	and	sends	out	to	review	
•  Reviewers	provide	suggesAons	and	criAcisms	
•  Editor	decides	whether	to	accept	
–  …	can	repeat	several	Ames	
•  Author	revises	
•  Final	acceptance	
•  Proofs	sent	to	author	
•  Author	corrects	
•  Author	signs	copyright	transfer	agreement	
•  PublicaAon
TOWN@KU.EDU

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