The synaptonemal complex forms between homologous chromosomes during meiosis and mediates chromosome pairing, synapsis, and recombination. It is a tripartite structure consisting of two parallel axes and a central element. Lampbrush chromosomes occur during oogenesis and are very large, with chromatin loops projecting from the chromosome axes that resemble the wick of a kerosene lamp, giving them their name. Lampbrush chromosomes are highly active in gene expression, with most genes in the chromatin loops being actively transcribed.