Plant growth and development involves two main processes: growth, which is an irreversible increase in size, and development, which refers to the changes an organism undergoes throughout its lifecycle. Growth occurs through cell division, cell enlargement, and cell differentiation, and can be measured by increases in length, area, or cell number. Development involves morphogenesis, the formation of shape and structure, and differentiation, when cells take on specialized functions. The processes of growth and development are controlled through intercellular, intracellular, and extracellular signals and allow a plant to complete its lifecycle from seed germination to senescence and death.