Shark evolution has been studied through their fossil record, as shark teeth are mineralogically stable and can provide details about species. The oldest shark fossils date back 455 million years to scales, while the Devonian era saw sharks becoming truly abundant. Ancient sharks such as Cladoselache differed from modern sharks in having rounded snouts, mouths in front, and equally sized tail lobes. Neoselachian sharks expanded 100 million years ago, inhabiting a variety of habitats as nearshore predators. Modern sharks demonstrate a diversity of forms like cow sharks and hammerheads.