Ecology is the study of interactions between organisms and their environment. The biosphere is the global ecosystem, encompassing all living things found in air, land, and water on Earth. Ecologists study how both biotic factors (living things) and abiotic factors (non-living things) influence organisms and populations within various levels of organization, from individuals to ecosystems. Energy and matter flow through ecosystems via food chains and food webs, with only about 10% of available energy being transferred between trophic levels. Key biogeochemical cycles such as the water, carbon, nitrogen, and phosphorus cycles recycle critical elements and compounds.