Gaseous exchange occurs through diffusion of oxygen from and carbon dioxide to the external environment. Respiratory structures have large surface areas, thin cell linings, and are kept moist to facilitate diffusion. Surface area increases with decreasing organism size, from entire cell membranes in unicellular organisms to branched tracheal systems in insects and gill filaments with countercurrent blood flow in aquatic organisms. Lungs and skin also enable gas exchange through networks of capillaries. Breathing involves rhythmic muscle movements that lower then raise pressure to draw air in and force it out.