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2.6 – Changes and some 2.5 Function
2.6.1 - 2.6.4
And 2.5.5 and 2.5.6

5/11/2013
IB/ESS
Author-Guru

Topic 2 – The
Ecosystem

1
Specifications

IB/ESS
Author-Guru

2.6.2 – Describe and explain ‘S’ and ‘J’ population growth curves.
Population curves should be sketched, described, interpreted
and constructed from given data.

5/11/2013

2.6.1 – Explain the concepts of limiting factors and carrying
capacity in the context of population growth.

2
Population Growth

IB/ESS
Author-Guru

• Nearly 1.6 million people
join the human population
each week.
• 84 million people join every
year.
• In three years the human
population grows by an
amount nearly equivalent to
the entire U.S population.
• By 2025 the world
population could exceed 8
billion

5/11/2013

Some Facts

3
• We are going to concentrate on population
control of ecosystems but these theories can
also be applied to human populations.

IB/ESS

• Studies on both human populations and smaller
ecosystem populations are carried out in depth.

Author-Guru

• The study of any population is an important
aspect of science.

5/11/2013

Population Studies

4
• The factors affecting a population
size may be biotic or abiotic.

IB/ESS
Author-Guru

• By taking samples and counting the
numbers of organisms in a
particular habitat, ecologists can
study the affects of any factor on
the size of a population.

5/11/2013

Population Size

• Together they affect the rate at
which population grows, and also
it’s final size.
5
IB/ESS

• How many abiotic factors can you think of that might affect
population size?

Author-Guru

• How many biotic factors can you think of that might affect
population size?

5/11/2013

Biotic Factors Affecting Population Size

6
Biotic and Abiotic Factors
5/11/2013

1. Temperature – higher
temperatures speed up
enzyme-catalyzed
reactions and increase
growth.
2. Oxygen Availability – affect
the rate of energy
production by respiration.
3. Light Availability – for
photosynthesis and
breeding cycles in animals
and plants.
4. Toxins and pollutants –
tissue growth may be
reduced.

IB/ESS

1. Food – both quantity
and quality of food are
important.
2. Predators – refer back
to predator prey
relationships.
3. Competitors – other
organisms may require
the same resources
from an environment.
4. Parasites – may cause
disease and slow down
the growth of an
organism.

Abiotic

Author-Guru

Biotic

7
8

Author-Guru

IB/ESS

5/11/2013
IB/ESS
Author-Guru

All of these things come under the category of ‘Limiting Factors’

5/11/2013

Biotic and Abiotic Factors

9
• Look at the graph of population growth.
• This shows how population growth is eventually
inhibited by environmental resistance and the
environment reaches it’s carrying capacity.

IB/ESS

• This is usually because there is plenty of food and no
accumulation of poisonous wastes.

Author-Guru

• When a small population grows in a particular
environment, the environmental resistance is almost
non-existent.

5/11/2013

Carrying Capacity

10
• Think of your brine shrimps!?

IB/ESS

• Once the carrying capacity is reached, unless the
environmental resistance is changed, e.g. by a
new disease, the size of the population will only
fluctuate slightly.

Author-Guru

• The carrying capacity (K) is the maximum
number of a species that the habitat can hold.

5/11/2013

Carrying Capacity

11
‘S’ Curves

IB/ESS
Author-Guru

• This is the type of graph that is almost always seen in nature.
• As the energy resources become more scarce the population
size levels off at the carrying capacity (K).

5/11/2013

• The graph we have just been looking at is an example of an ‘S’
curve.

12
Author-Guru

IB/ESS

5/11/2013

‘J’ Curves

13
• If the resources of the new habitat were endless
then the population would continue to increase
at this rate.

IB/ESS

• This type of growth produces a J shaped growth
curve.

Author-Guru

• Just as in the ‘S’ curve example, a population
establishing themselves in a new area will
undergo rapid exponential growth.

5/11/2013

‘J’ Curves

14
• Can you think of any examples where ‘J’ curve
population growth would be extremely
desirable.

IB/ESS

• Initially exponential growth will occur but
eventually the increase in numbers will not be
supported by the environment.

Author-Guru

• This type of population growth is rarely seen in
nature.

5/11/2013

‘J’ Curves

15
• It is very natural to ask the linked questions - does humanity
have a carrying capacity and, if so, what is it - and when will
we reach or overshoot this

IB/ESS

• Furthermore, experience with other species tells us that,
ultimately, resource limitations and/or habitat degradation
will force the human population curves to approach an upper
limit - the carrying capacity, often symbolized as " K" by
ecologists.

Author-Guru

• ‘As we have seen, the human population growth curve is
currently following an exponential curve or a "J-shape”.
Common sense tells us that such growth cannot continue otherwise within a few hundred years every square foot of
the Earth's surface would be taken up by a human.

5/11/2013

Is there a Carrying Capacity for Homo
sapiens?

16
Activity

IB/ESS
Author-Guru

2.6.3 – Describe the role of density-dependent and densityindependent factors, and internal and external factors, in the
regulation of populations.

5/11/2013

• Complete the activity – The new zoos

17
• These factors exert their effect irrespective of
the size of the population when the catastrophe
struck.

IB/ESS
Author-Guru

• The following factors are classed as densityindependent factors:
• Drought
• Freezes
• Hurricanes
• Floods
• Forest Fires

5/11/2013

Density Independent Factors

18
The drought ended in 1978, but even
with ample food once again available
the finch population recovered only
slowly.

IB/ESS

The decline (from 1400 to 200
individuals) occurred because of a
severe drought that reduced the
quantity of seeds on which this
species feeds.

Author-Guru

This graph shows the decline in the
population of one of Darwin's finches
(Geospiza fortis) on Daphne Major, a
tiny (100-acre) member of the
Galapagos Islands.

5/11/2013

Density Independent Factors

19
• Many rodent populations (e.g.,
lemmings in the Arctic) also go
through such boom-and-bust
cycles.

IB/ESS

• Read the information about the
gypsy moth.

Author-Guru

• Intraspecific Competition competition between members of
the same species.

5/11/2013

Density Dependant Factors

20
• This occurs when two species share overlapping
ecological niches, they may be forced into
competition for the resource(s) of that niche.

IB/ESS

• This can include food, nesting sites, sunlight.

Author-Guru

• Interspecific Competition – this is competition
between different species for different
resources.

5/11/2013

Density Dependant Factors

21
Specifications

Author-Guru

IB/ESS

5/11/2013

• 2.6.4 – Describe the principles associated with survivorship
curves including K- and r -strategists.

22
• Ragweed is well adapted to
exploiting it’s environment
in a hurry – before
competitors can become
established!

IB/ESS
Author-Guru

• “I once ploughed up an old
field and allowed it to lie
fallow. In the first season it
grew a large crop of
ragweed.”

5/11/2013

R-Strategists

23
R-Strategists

• Can you think of any other animals that may be r-strategists?

IB/ESS
Author-Guru

• We say that they have a high value of ‘r’
• They are called r-strategists

5/11/2013

• Ragweed’s approach to continued survival is through rapid
reproduction.

24
2.
3.
4.
5.
6.
7.

IB/ESS

1.

In general, r-strategists share a number of
features:
Usually found in disturbed and/or transitory
habitats
Have short life spans
Begin breeding early in life
Have short gestation times
Produce large numbers of offspring
Take little care of their offspring (infant
mortality large)
Have efficient means of dispersal to new
habitats

Author-Guru

•

5/11/2013

R-Strategists

25
• K-strategists have a stable population that is
close to K.
• There is nothing to be gained from a high r.

• The species will benefit the most by a close
adaptation to the conditions of the environment.

IB/ESS
Author-Guru

• When a habitat become filled with a diverse
collection of creatures competing with one
another for resources, the advantage shifts to KStrategists

5/11/2013

K-Strategists

26
IB/ESS

K-strategists share these qualities:
Found in a stable habitat
Long life spans
Begin breeding later in life
Long gestation times
Produce small numbers of offspring
Take good care of their young – infant
mortality low
7. Have evolved to become increasingly efficient
at exploiting an ever-narrower slice of their
environment.

Author-Guru

•
1.
2.
3.
4.
5.
6.

5/11/2013

K-Strategists

27
Survivorship Curves

Author-Guru

IB/ESS

5/11/2013

• The graph shows 4 representative survivorship curves.

28
Survivorship Curves

IB/ESS
Author-Guru

• Curve B – typical of populations in which factors such
as starvation and disease inhibit the effects of aging
and infant mortality is high.

5/11/2013

• Curve A – characteristic of organisms that have low
mortality until late in life when aging takes its toll.

• Curve C – a theoretical curve for an organism whereby
the chance of death is equal at all stages
• Curve D – typical of organisms that produce huge
numbers of offspring accompanied by high rates of
mortality.

29
Survivorship Curves

• The Californian side-blotted lizard

Author-Guru

IB/ESS

• R-strategists usually have D survivorship curves.

5/11/2013

• K-strategists usually have survivorship curves somewhere
between A and C.

30
• 2.6.7 – Describe factors affecting the nature of
climax communities.

IB/ESS

• 2.6.6 – Explain the changes in energy flows,
gross and net productivity, diversity and mineral
cycling in different stages of succession.

Author-Guru

• 2.6.5 – Describe the concept and processes of
succession in a named habitat.

5/11/2013

Specifications

31
Succession – An intro

• Succession takes places as a result of complex
interactions of biotic and abiotic factors.
• Early communities modify the physical environment
causing it to change.
• This in turn alters the biotic community which
further alters the physical environment and so on.

IB/ESS
Author-Guru

• A forest following a disturbance such as a fire.

5/11/2013

• The gradual process by which the species population
of a community changes is called ecological
succession.

32
Succession – What happens?

IB/ESS
Author-Guru

• A succession (or sere) proceeds in seral stages, until the
formation of a climax community is reached.

5/11/2013

• Each successive community makes the environment more
favourable for the establishment of new species.

33
Primary Succession

IB/ESS

• You will be studying glacial moraines in detail as well as the
succession occurring on bare rock.

Author-Guru

• Can you think of examples where this would occur?

5/11/2013

• Refers to colonization of regions where there is no pre-existing
community.

34
Succession

IB/ESS
Author-Guru

• Study the information on glacial moraines and answer the
following questions:

5/11/2013

• Community changes on a glacial moraines

35
1.
2.

Explain what is meant by a climax community.
Explain each of the following changes which occur
during succession:
Species diversity increases
Gross production increases
Stability of the ecosystem increases
Give two reasons why farmland in the UK does not
reach a climax community.

a)
b)
c)
3.

IB/ESS

During succession there is a change in species
composition of a community. There are also changes
in species diversity, stability of the ecosystem, and in
gross and net production until a climax community is
reached.

Author-Guru

•

5/11/2013

Questions – Glacial Moraines

36
IB/ESS

• Secondary Succession – occurs on sites that have previously
supported a community of some sort.
• Examples?

Author-Guru

• Primary Succession – occurs on newly formed habitats that
have not previously supported a community.
• Examples?

5/11/2013

Primary and Secondary Succession

37
Lichens,
bryophytes
and annual
herbs

After 100-200 years

Slower growing
broadleaf species
e.g. oak
Complex Community

Fast
growing
trees e.g.
Ash

Author-Guru

IB/ESS

Bare Rock

Mosses,
Grasses and
small
shrubs

5/11/2013

Primary Succession – Bare
Rock

38

Example for a Northern Hemisphere lithosere: a succession on bare rock
• As these species begin to grow well, they produce shade.
Their own seedlings grow more poorly than shade-adapted
plants.
• Plants that grow well under full sun are replaced by plants
that germinate and grow better in deeper shade.

IB/ESS

• Examples = lichens, grasses, herbs

Author-Guru

• These are usually fast growing plants that photosynthesize
well in full sunlight.
• We call these pioneer species making up the pioneer
community

5/11/2013

In Summary - the 1st Invaders!

39
Secondary Succession

• Humans may deflect the natural course of succession
in these circumstances (e.g. by mowing or farming).
• This leads to the development of a different climax
community than would otherwise develop naturally.

IB/ESS
Author-Guru

• These events do not involve loss of the soil.
• Secondary succession therefore occurs more rapidly
than primary succession.

5/11/2013

• This type of succession takes place after a land
clearance (e.g. from fire or landslide).

40
1-2
3-5

Young broad
leaved woodland
31-150

Mature
woodland:
mainly oak

Scrub: shrubs
and small trees

IB/ESS

Time to develop: Years

Grasses and low
growing
perennials

Author-Guru

Primary Bare
Earth

Open pioneer
community
(annual grasses)

5/11/2013

Secondary Succession – Cleared
Land

16-30
41

150+ = climax community
Succession Continues

• Animal species have a profound affect on the plant species
occurring within a habitat.
• Changing conditions in the present community allows for new
species to become established (the future community).
• Succession continues until the climax community is reached.

IB/ESS
Author-Guru

• Decomposers will join the community as well as animal
species.

5/11/2013

• As the plant community changes, the soil will also undergo
changes (abiotic factors will change).

42
Wetland Succession
5/11/2013

• Wetland areas present a special case of ecological succession.

Open water

Plant invasion

Siltation and
Infilling

Author-Guru

IB/ESS

• Wetlands are constantly changing:

• Wetland ecosystem may develop in a variety of ways:
43
Wetland Succession

• In special circumstances, a an acid peat bog may develop.
(may take 5000+ years).

IB/ESS
Author-Guru

• In non-acidic, poorly drained areas, a swamp will eventually
develop into a fen.

5/11/2013

• In well drained areas, pasture or heath may develop as a
result of succession from fresh water to dry land.

44
IB/ESS

• This is called productivity

Author-Guru

• Think back to the work on food webs/chains
• It is often useful to know how much energy is
passing through a trophic level over a period of
time.

5/11/2013

Productivity

• Productivity is a measure of the amount of
energy incorporated into the organisms in a
trophic level, in an area, over a certain period of
time.

45
IB/ESS

• 2.5.6 – Define and Calculate the values of gross and net
productivity from given data

Author-Guru

• 2.5.5 – Define the terms gross productivity, net productivity,
primary productivity, secondary productivity, gross primary
productivity and net primary productivity.

5/11/2013

Specifications

46
Productivity

• The rate at which producers convert light energy into chemical
energy is called primary productivity.

IB/ESS
Author-Guru

• It is therefore measured in units of kilojoules per square metre
per year (kJm-2year-1)

5/11/2013

• The area is normally one square metre and the time is usually
one year.

47
Gross Productivity

• The producers use some of this energy during respiration and
energy needs which is eventually lost to the environment as
heat.
• The remaining energy is available to the herbivores and is
known as net primary productivity (NPP)

IB/ESS

• It is related to the total amount of chemical energy
incorporated into the producers.

Author-Guru

• This is sometimes shown as GPP – Gross Primary Productivity

5/11/2013

• Gross Productivity (GP) – is the total gain in energy or
biomass per unit time.

48
Recap of Definitions!

• Gross Productivity (GP) = the total gain in energy/biomass per unit time.
• Gross Primary Productivity (GPP) = the total gain in energy of the producers.
• Net Productivity (NP) = the gain in energy/biomass per unit time remaining
after allowing for respiration (R) loses.
• Net Primary Productivity (NPP) = the gain in energy/biomass per unit time
remaining after allowing for respiration loses which is passed onto the
herbivores.

IB/ESS

• Secondary Productivity = The rate at which primary material is synthesised
into animal tissue per unit area in a given time.

Author-Guru

• Primary Productivity = The rate at which energy/biomass is formed through
photosynthesis

5/11/2013

• Productivity = production per unit time

49
IB/ESS

• The rate at which plants can convert light energy
into chemical energy is affected by many factors:
• Sunlight
• Water
• Temperature
• Amount of nutrients

Author-Guru

• Primary productivity varies greatly in different
environments.

5/11/2013

Environmental Productivity

50
Environmental Productivity

• In the oceans however, the most productive areas are in cold
regions due to the up-welling of water bringing plant nutrients
with it.

IB/ESS
Author-Guru

• This is due to good light levels and high temperatures in the
tropics.

5/11/2013

• In natural ecosystems primary productivity tends to be highest
in tropical regions.

51
IB/ESS

• We can calculate NPP for both producers and
consumers as:
NPP = GPP – energy used in respiration

Author-Guru

• We can calculate GPP as follows:
GPP = NPP + R

5/11/2013

Important Calculations

• In addition, the equation for consumers only is:
GP = food eaten – faecal losses
52
Calculating Productivity Values

• What is the GPP of an ecosystem if the NPP is 1660 kJm2yr-1 and the energy lost during respiration is 573 kJm-2yr1?

• What is the NPP if the GPP is 2700 kJm-2yr-1 and the
energy used in respiration is 1850 kJm-2yr-1?

IB/ESS
Author-Guru

• What is the % energy from sunlight that is fixed as GPP if
the total energy from the sun in 3 x 106 and the gross
primary productivity = 2.8 x 104?

5/11/2013

• Some easy ones to start you off!

53
IB/ESS
Author-Guru

Now for some slightly harder ones!

5/11/2013

Calculating Productivity Values

54
Therefore, for photoautotrophs, photosynthetic
efficiency is determined as:

Photosynthetic Efficiency =
Net production ÷ Light Energy Absorbed

IB/ESS

Ecological Efficiency is the net production of new
biomass at each trophic level as a percentage of the
total energy flowing through that trophic level

Author-Guru

Energy flow diagrams illustrate energy flow
through communities and include values for
respiratory losses and energy flow through
the decomposers
Information from energy flow diagrams can be used to
calculate ecological efficiencies

5/11/2013

Energy Flow Diagrams

55
5/11/2013
IB/ESS
Author-Guru

Use information from the energy flow diagram to:
• Explain the meaning of the term Gross Primary Production
• Explain the meaning of the term Net Primary Production
• Calculate the Photosynthetic Efficiency of the phytoplankton

56
5/11/2013
IB/ESS
Author-Guru

Gross Primary Production is the total energy fixed
by photoautotrophs during photosynthesis
Net Primary Production is the energy stored as
biomass (gross production – energy lost as heat
in respiration)

Photosynthetic
Efficiency =
57
3.7 x 104
------------ x 100
172 x 104
= 2.15%
58

Author-Guru

IB/ESS

5/11/2013
114 x 103 kJ m-2year-1

NPP =
69.7 x 103 kJ m-2year-1

NPP =
200 kJ m-2year-1

Author-Guru

GPP =
24 x106 kJ m-2year-1

IB/ESS

5/11/2013

Photosynthetic
efficiency = 1.3%

NPP =
800 kJ m-2year-1

59
• This is because climax communities are better
adapted to an efficient rate of utilisation of their
resources.
• They become stable.

IB/ESS

• As ecosystems become more diverse, the overall
GPP is also going to increase.

Author-Guru

• The NPP and GPP of any ecosystem is going to
fluctuate. This is especially the case during each
seral stage.

5/11/2013

Finally Back To Succession!

60
The Early Stages

• This is due to low respiration rates of the initial
producers and therefore a lot of energy available to be
passed on.
• This allows the system to grow and biomass to
accumulate.

IB/ESS

• Net Productivity = High

Author-Guru

• This is due to the initial conditions and the relatively low
density of producers.

5/11/2013

• Gross Productivity = Low

61
The Later Stages

• Net Productivity = Low
• Increased rates of respiration and other energy
sapping activities by consumers means that NP
will begin approaching zero.

IB/ESS
Author-Guru

• This is due to an increase in the consumer
community who can synthesise a lot of energy
from the food they eat.

5/11/2013

• Gross Productivity = High

62
• Ultimately, the climate will be responsible for
affecting the nature of the climax community
unless human or other factors maintain an
equilibrium at a sub-climax community.

IB/ESS

• Climax communities are more stable that the
seral stages that preceded them.

Author-Guru

• Succession comes to an end with the
establishment of a mature, relatively stable
community – the climax

5/11/2013

The Climax

63

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Similar to 2.6.1 5 6 and7and some 2.5 gpp npp

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  • 1. 2.6 – Changes and some 2.5 Function 2.6.1 - 2.6.4 And 2.5.5 and 2.5.6 5/11/2013 IB/ESS Author-Guru Topic 2 – The Ecosystem 1
  • 2. Specifications IB/ESS Author-Guru 2.6.2 – Describe and explain ‘S’ and ‘J’ population growth curves. Population curves should be sketched, described, interpreted and constructed from given data. 5/11/2013 2.6.1 – Explain the concepts of limiting factors and carrying capacity in the context of population growth. 2
  • 3. Population Growth IB/ESS Author-Guru • Nearly 1.6 million people join the human population each week. • 84 million people join every year. • In three years the human population grows by an amount nearly equivalent to the entire U.S population. • By 2025 the world population could exceed 8 billion 5/11/2013 Some Facts 3
  • 4. • We are going to concentrate on population control of ecosystems but these theories can also be applied to human populations. IB/ESS • Studies on both human populations and smaller ecosystem populations are carried out in depth. Author-Guru • The study of any population is an important aspect of science. 5/11/2013 Population Studies 4
  • 5. • The factors affecting a population size may be biotic or abiotic. IB/ESS Author-Guru • By taking samples and counting the numbers of organisms in a particular habitat, ecologists can study the affects of any factor on the size of a population. 5/11/2013 Population Size • Together they affect the rate at which population grows, and also it’s final size. 5
  • 6. IB/ESS • How many abiotic factors can you think of that might affect population size? Author-Guru • How many biotic factors can you think of that might affect population size? 5/11/2013 Biotic Factors Affecting Population Size 6
  • 7. Biotic and Abiotic Factors 5/11/2013 1. Temperature – higher temperatures speed up enzyme-catalyzed reactions and increase growth. 2. Oxygen Availability – affect the rate of energy production by respiration. 3. Light Availability – for photosynthesis and breeding cycles in animals and plants. 4. Toxins and pollutants – tissue growth may be reduced. IB/ESS 1. Food – both quantity and quality of food are important. 2. Predators – refer back to predator prey relationships. 3. Competitors – other organisms may require the same resources from an environment. 4. Parasites – may cause disease and slow down the growth of an organism. Abiotic Author-Guru Biotic 7
  • 9. IB/ESS Author-Guru All of these things come under the category of ‘Limiting Factors’ 5/11/2013 Biotic and Abiotic Factors 9
  • 10. • Look at the graph of population growth. • This shows how population growth is eventually inhibited by environmental resistance and the environment reaches it’s carrying capacity. IB/ESS • This is usually because there is plenty of food and no accumulation of poisonous wastes. Author-Guru • When a small population grows in a particular environment, the environmental resistance is almost non-existent. 5/11/2013 Carrying Capacity 10
  • 11. • Think of your brine shrimps!? IB/ESS • Once the carrying capacity is reached, unless the environmental resistance is changed, e.g. by a new disease, the size of the population will only fluctuate slightly. Author-Guru • The carrying capacity (K) is the maximum number of a species that the habitat can hold. 5/11/2013 Carrying Capacity 11
  • 12. ‘S’ Curves IB/ESS Author-Guru • This is the type of graph that is almost always seen in nature. • As the energy resources become more scarce the population size levels off at the carrying capacity (K). 5/11/2013 • The graph we have just been looking at is an example of an ‘S’ curve. 12
  • 14. • If the resources of the new habitat were endless then the population would continue to increase at this rate. IB/ESS • This type of growth produces a J shaped growth curve. Author-Guru • Just as in the ‘S’ curve example, a population establishing themselves in a new area will undergo rapid exponential growth. 5/11/2013 ‘J’ Curves 14
  • 15. • Can you think of any examples where ‘J’ curve population growth would be extremely desirable. IB/ESS • Initially exponential growth will occur but eventually the increase in numbers will not be supported by the environment. Author-Guru • This type of population growth is rarely seen in nature. 5/11/2013 ‘J’ Curves 15
  • 16. • It is very natural to ask the linked questions - does humanity have a carrying capacity and, if so, what is it - and when will we reach or overshoot this IB/ESS • Furthermore, experience with other species tells us that, ultimately, resource limitations and/or habitat degradation will force the human population curves to approach an upper limit - the carrying capacity, often symbolized as " K" by ecologists. Author-Guru • ‘As we have seen, the human population growth curve is currently following an exponential curve or a "J-shape”. Common sense tells us that such growth cannot continue otherwise within a few hundred years every square foot of the Earth's surface would be taken up by a human. 5/11/2013 Is there a Carrying Capacity for Homo sapiens? 16
  • 17. Activity IB/ESS Author-Guru 2.6.3 – Describe the role of density-dependent and densityindependent factors, and internal and external factors, in the regulation of populations. 5/11/2013 • Complete the activity – The new zoos 17
  • 18. • These factors exert their effect irrespective of the size of the population when the catastrophe struck. IB/ESS Author-Guru • The following factors are classed as densityindependent factors: • Drought • Freezes • Hurricanes • Floods • Forest Fires 5/11/2013 Density Independent Factors 18
  • 19. The drought ended in 1978, but even with ample food once again available the finch population recovered only slowly. IB/ESS The decline (from 1400 to 200 individuals) occurred because of a severe drought that reduced the quantity of seeds on which this species feeds. Author-Guru This graph shows the decline in the population of one of Darwin's finches (Geospiza fortis) on Daphne Major, a tiny (100-acre) member of the Galapagos Islands. 5/11/2013 Density Independent Factors 19
  • 20. • Many rodent populations (e.g., lemmings in the Arctic) also go through such boom-and-bust cycles. IB/ESS • Read the information about the gypsy moth. Author-Guru • Intraspecific Competition competition between members of the same species. 5/11/2013 Density Dependant Factors 20
  • 21. • This occurs when two species share overlapping ecological niches, they may be forced into competition for the resource(s) of that niche. IB/ESS • This can include food, nesting sites, sunlight. Author-Guru • Interspecific Competition – this is competition between different species for different resources. 5/11/2013 Density Dependant Factors 21
  • 22. Specifications Author-Guru IB/ESS 5/11/2013 • 2.6.4 – Describe the principles associated with survivorship curves including K- and r -strategists. 22
  • 23. • Ragweed is well adapted to exploiting it’s environment in a hurry – before competitors can become established! IB/ESS Author-Guru • “I once ploughed up an old field and allowed it to lie fallow. In the first season it grew a large crop of ragweed.” 5/11/2013 R-Strategists 23
  • 24. R-Strategists • Can you think of any other animals that may be r-strategists? IB/ESS Author-Guru • We say that they have a high value of ‘r’ • They are called r-strategists 5/11/2013 • Ragweed’s approach to continued survival is through rapid reproduction. 24
  • 25. 2. 3. 4. 5. 6. 7. IB/ESS 1. In general, r-strategists share a number of features: Usually found in disturbed and/or transitory habitats Have short life spans Begin breeding early in life Have short gestation times Produce large numbers of offspring Take little care of their offspring (infant mortality large) Have efficient means of dispersal to new habitats Author-Guru • 5/11/2013 R-Strategists 25
  • 26. • K-strategists have a stable population that is close to K. • There is nothing to be gained from a high r. • The species will benefit the most by a close adaptation to the conditions of the environment. IB/ESS Author-Guru • When a habitat become filled with a diverse collection of creatures competing with one another for resources, the advantage shifts to KStrategists 5/11/2013 K-Strategists 26
  • 27. IB/ESS K-strategists share these qualities: Found in a stable habitat Long life spans Begin breeding later in life Long gestation times Produce small numbers of offspring Take good care of their young – infant mortality low 7. Have evolved to become increasingly efficient at exploiting an ever-narrower slice of their environment. Author-Guru • 1. 2. 3. 4. 5. 6. 5/11/2013 K-Strategists 27
  • 28. Survivorship Curves Author-Guru IB/ESS 5/11/2013 • The graph shows 4 representative survivorship curves. 28
  • 29. Survivorship Curves IB/ESS Author-Guru • Curve B – typical of populations in which factors such as starvation and disease inhibit the effects of aging and infant mortality is high. 5/11/2013 • Curve A – characteristic of organisms that have low mortality until late in life when aging takes its toll. • Curve C – a theoretical curve for an organism whereby the chance of death is equal at all stages • Curve D – typical of organisms that produce huge numbers of offspring accompanied by high rates of mortality. 29
  • 30. Survivorship Curves • The Californian side-blotted lizard Author-Guru IB/ESS • R-strategists usually have D survivorship curves. 5/11/2013 • K-strategists usually have survivorship curves somewhere between A and C. 30
  • 31. • 2.6.7 – Describe factors affecting the nature of climax communities. IB/ESS • 2.6.6 – Explain the changes in energy flows, gross and net productivity, diversity and mineral cycling in different stages of succession. Author-Guru • 2.6.5 – Describe the concept and processes of succession in a named habitat. 5/11/2013 Specifications 31
  • 32. Succession – An intro • Succession takes places as a result of complex interactions of biotic and abiotic factors. • Early communities modify the physical environment causing it to change. • This in turn alters the biotic community which further alters the physical environment and so on. IB/ESS Author-Guru • A forest following a disturbance such as a fire. 5/11/2013 • The gradual process by which the species population of a community changes is called ecological succession. 32
  • 33. Succession – What happens? IB/ESS Author-Guru • A succession (or sere) proceeds in seral stages, until the formation of a climax community is reached. 5/11/2013 • Each successive community makes the environment more favourable for the establishment of new species. 33
  • 34. Primary Succession IB/ESS • You will be studying glacial moraines in detail as well as the succession occurring on bare rock. Author-Guru • Can you think of examples where this would occur? 5/11/2013 • Refers to colonization of regions where there is no pre-existing community. 34
  • 35. Succession IB/ESS Author-Guru • Study the information on glacial moraines and answer the following questions: 5/11/2013 • Community changes on a glacial moraines 35
  • 36. 1. 2. Explain what is meant by a climax community. Explain each of the following changes which occur during succession: Species diversity increases Gross production increases Stability of the ecosystem increases Give two reasons why farmland in the UK does not reach a climax community. a) b) c) 3. IB/ESS During succession there is a change in species composition of a community. There are also changes in species diversity, stability of the ecosystem, and in gross and net production until a climax community is reached. Author-Guru • 5/11/2013 Questions – Glacial Moraines 36
  • 37. IB/ESS • Secondary Succession – occurs on sites that have previously supported a community of some sort. • Examples? Author-Guru • Primary Succession – occurs on newly formed habitats that have not previously supported a community. • Examples? 5/11/2013 Primary and Secondary Succession 37
  • 38. Lichens, bryophytes and annual herbs After 100-200 years Slower growing broadleaf species e.g. oak Complex Community Fast growing trees e.g. Ash Author-Guru IB/ESS Bare Rock Mosses, Grasses and small shrubs 5/11/2013 Primary Succession – Bare Rock 38 Example for a Northern Hemisphere lithosere: a succession on bare rock
  • 39. • As these species begin to grow well, they produce shade. Their own seedlings grow more poorly than shade-adapted plants. • Plants that grow well under full sun are replaced by plants that germinate and grow better in deeper shade. IB/ESS • Examples = lichens, grasses, herbs Author-Guru • These are usually fast growing plants that photosynthesize well in full sunlight. • We call these pioneer species making up the pioneer community 5/11/2013 In Summary - the 1st Invaders! 39
  • 40. Secondary Succession • Humans may deflect the natural course of succession in these circumstances (e.g. by mowing or farming). • This leads to the development of a different climax community than would otherwise develop naturally. IB/ESS Author-Guru • These events do not involve loss of the soil. • Secondary succession therefore occurs more rapidly than primary succession. 5/11/2013 • This type of succession takes place after a land clearance (e.g. from fire or landslide). 40
  • 41. 1-2 3-5 Young broad leaved woodland 31-150 Mature woodland: mainly oak Scrub: shrubs and small trees IB/ESS Time to develop: Years Grasses and low growing perennials Author-Guru Primary Bare Earth Open pioneer community (annual grasses) 5/11/2013 Secondary Succession – Cleared Land 16-30 41 150+ = climax community
  • 42. Succession Continues • Animal species have a profound affect on the plant species occurring within a habitat. • Changing conditions in the present community allows for new species to become established (the future community). • Succession continues until the climax community is reached. IB/ESS Author-Guru • Decomposers will join the community as well as animal species. 5/11/2013 • As the plant community changes, the soil will also undergo changes (abiotic factors will change). 42
  • 43. Wetland Succession 5/11/2013 • Wetland areas present a special case of ecological succession. Open water Plant invasion Siltation and Infilling Author-Guru IB/ESS • Wetlands are constantly changing: • Wetland ecosystem may develop in a variety of ways: 43
  • 44. Wetland Succession • In special circumstances, a an acid peat bog may develop. (may take 5000+ years). IB/ESS Author-Guru • In non-acidic, poorly drained areas, a swamp will eventually develop into a fen. 5/11/2013 • In well drained areas, pasture or heath may develop as a result of succession from fresh water to dry land. 44
  • 45. IB/ESS • This is called productivity Author-Guru • Think back to the work on food webs/chains • It is often useful to know how much energy is passing through a trophic level over a period of time. 5/11/2013 Productivity • Productivity is a measure of the amount of energy incorporated into the organisms in a trophic level, in an area, over a certain period of time. 45
  • 46. IB/ESS • 2.5.6 – Define and Calculate the values of gross and net productivity from given data Author-Guru • 2.5.5 – Define the terms gross productivity, net productivity, primary productivity, secondary productivity, gross primary productivity and net primary productivity. 5/11/2013 Specifications 46
  • 47. Productivity • The rate at which producers convert light energy into chemical energy is called primary productivity. IB/ESS Author-Guru • It is therefore measured in units of kilojoules per square metre per year (kJm-2year-1) 5/11/2013 • The area is normally one square metre and the time is usually one year. 47
  • 48. Gross Productivity • The producers use some of this energy during respiration and energy needs which is eventually lost to the environment as heat. • The remaining energy is available to the herbivores and is known as net primary productivity (NPP) IB/ESS • It is related to the total amount of chemical energy incorporated into the producers. Author-Guru • This is sometimes shown as GPP – Gross Primary Productivity 5/11/2013 • Gross Productivity (GP) – is the total gain in energy or biomass per unit time. 48
  • 49. Recap of Definitions! • Gross Productivity (GP) = the total gain in energy/biomass per unit time. • Gross Primary Productivity (GPP) = the total gain in energy of the producers. • Net Productivity (NP) = the gain in energy/biomass per unit time remaining after allowing for respiration (R) loses. • Net Primary Productivity (NPP) = the gain in energy/biomass per unit time remaining after allowing for respiration loses which is passed onto the herbivores. IB/ESS • Secondary Productivity = The rate at which primary material is synthesised into animal tissue per unit area in a given time. Author-Guru • Primary Productivity = The rate at which energy/biomass is formed through photosynthesis 5/11/2013 • Productivity = production per unit time 49
  • 50. IB/ESS • The rate at which plants can convert light energy into chemical energy is affected by many factors: • Sunlight • Water • Temperature • Amount of nutrients Author-Guru • Primary productivity varies greatly in different environments. 5/11/2013 Environmental Productivity 50
  • 51. Environmental Productivity • In the oceans however, the most productive areas are in cold regions due to the up-welling of water bringing plant nutrients with it. IB/ESS Author-Guru • This is due to good light levels and high temperatures in the tropics. 5/11/2013 • In natural ecosystems primary productivity tends to be highest in tropical regions. 51
  • 52. IB/ESS • We can calculate NPP for both producers and consumers as: NPP = GPP – energy used in respiration Author-Guru • We can calculate GPP as follows: GPP = NPP + R 5/11/2013 Important Calculations • In addition, the equation for consumers only is: GP = food eaten – faecal losses 52
  • 53. Calculating Productivity Values • What is the GPP of an ecosystem if the NPP is 1660 kJm2yr-1 and the energy lost during respiration is 573 kJm-2yr1? • What is the NPP if the GPP is 2700 kJm-2yr-1 and the energy used in respiration is 1850 kJm-2yr-1? IB/ESS Author-Guru • What is the % energy from sunlight that is fixed as GPP if the total energy from the sun in 3 x 106 and the gross primary productivity = 2.8 x 104? 5/11/2013 • Some easy ones to start you off! 53
  • 54. IB/ESS Author-Guru Now for some slightly harder ones! 5/11/2013 Calculating Productivity Values 54
  • 55. Therefore, for photoautotrophs, photosynthetic efficiency is determined as: Photosynthetic Efficiency = Net production ÷ Light Energy Absorbed IB/ESS Ecological Efficiency is the net production of new biomass at each trophic level as a percentage of the total energy flowing through that trophic level Author-Guru Energy flow diagrams illustrate energy flow through communities and include values for respiratory losses and energy flow through the decomposers Information from energy flow diagrams can be used to calculate ecological efficiencies 5/11/2013 Energy Flow Diagrams 55
  • 56. 5/11/2013 IB/ESS Author-Guru Use information from the energy flow diagram to: • Explain the meaning of the term Gross Primary Production • Explain the meaning of the term Net Primary Production • Calculate the Photosynthetic Efficiency of the phytoplankton 56
  • 57. 5/11/2013 IB/ESS Author-Guru Gross Primary Production is the total energy fixed by photoautotrophs during photosynthesis Net Primary Production is the energy stored as biomass (gross production – energy lost as heat in respiration) Photosynthetic Efficiency = 57 3.7 x 104 ------------ x 100 172 x 104 = 2.15%
  • 59. 114 x 103 kJ m-2year-1 NPP = 69.7 x 103 kJ m-2year-1 NPP = 200 kJ m-2year-1 Author-Guru GPP = 24 x106 kJ m-2year-1 IB/ESS 5/11/2013 Photosynthetic efficiency = 1.3% NPP = 800 kJ m-2year-1 59
  • 60. • This is because climax communities are better adapted to an efficient rate of utilisation of their resources. • They become stable. IB/ESS • As ecosystems become more diverse, the overall GPP is also going to increase. Author-Guru • The NPP and GPP of any ecosystem is going to fluctuate. This is especially the case during each seral stage. 5/11/2013 Finally Back To Succession! 60
  • 61. The Early Stages • This is due to low respiration rates of the initial producers and therefore a lot of energy available to be passed on. • This allows the system to grow and biomass to accumulate. IB/ESS • Net Productivity = High Author-Guru • This is due to the initial conditions and the relatively low density of producers. 5/11/2013 • Gross Productivity = Low 61
  • 62. The Later Stages • Net Productivity = Low • Increased rates of respiration and other energy sapping activities by consumers means that NP will begin approaching zero. IB/ESS Author-Guru • This is due to an increase in the consumer community who can synthesise a lot of energy from the food they eat. 5/11/2013 • Gross Productivity = High 62
  • 63. • Ultimately, the climate will be responsible for affecting the nature of the climax community unless human or other factors maintain an equilibrium at a sub-climax community. IB/ESS • Climax communities are more stable that the seral stages that preceded them. Author-Guru • Succession comes to an end with the establishment of a mature, relatively stable community – the climax 5/11/2013 The Climax 63