Nitrogen is essential for plant growth but is often limited in availability. Plants have evolved symbiotic relationships with nitrogen-fixing bacteria to access atmospheric nitrogen. These relationships include root and stem nodule symbioses with rhizobia and Frankia bacteria that live inside root nodules. Cyanobacteria also engage in nitrogen-fixing symbioses, living inside specialized plant tissues. Associative bacteria in the rhizosphere can also increase nitrogen availability through nitrogen fixation. These symbioses allowed plants to colonize land and overcome nitrogen limitations.