This document describes a 5-year study assessing the impacts of control operations on a recreationally hunted feral swine population at Avon Park Air Force Range in Florida. Researchers monitored the population from 2008-2012 using a passive tracking index during dry and wet seasons, and hunter take rates during the dry season. All indices showed substantial declines in the feral swine population after control operations began in 2009, leveling off for the final two study years. Control efforts were limited by military missions and hunting seasons, preventing further reduction of the population. The tracking index also monitored coyotes and detected black bears and panthers on the base.
This study evaluated and compared the vegetative composition of four tern nesting colonies in Southern New England. Data was collected on plant species and coverage at Common Tern, Roseate Tern, and random nest and sampling sites. In total, 34 plant species were documented, with 16 being relatively common across multiple sites. The sites varied in plant species richness and prevalence of invasive species. Management of vegetation is important for maintaining suitable open nesting habitat for terns but requires a combination of removal, soil, and habitat construction methods.
Aquatic Insect Similarity Connecting Natural Wetland Habitat and Ricefield fo...Innspub Net
This study describes the relationship of the natural wetland habitats in Agusan Marsh to nearby rice fields and its implication to ecological rice production. Aquatic insects play multiple roles in the ecosystem such as predators, prey to other animals and decomposers which help in maintaining ecological balance. Results revealed that the diversity of odonates was highest in the sedge-dominated swamp among natural habitats which corresponds to the adjoining ricefields. The pattern of clustering of odonates show 3 groups; the rice-sago and rice-sedges sub-cluster, the rice-bangkal, rice-Terminalia, rice-fern, bangkal, sago and sedges sub-cluster, and the Terminalia forest as the outlier. The diversity of semi-aquatic and aquatic bugs was highest in the Bangkal forest while the lowest was in the fern-dominated swamp. The pattern of clustering shows 2 sub-clusters and the outlier Bangkal forest. On aquatic beetles, highest diversity was in the Terminalia forest. The sub-cluster consists of Terminalia habitat and rice-fern, while the other sub-cluster includes rice-Bangkal and rice-Terminalia. The resulting patterns of similarity in diversity and distribution of species in natural habitats and nearby ricefields indicate that ricefields are important temporary habitats for some aquatic insect species and serve as stepping stones for the movement of the insects.
Analyses of Community Attributes of Meiofauna Under A Pollution Regime in the...Agriculture Journal IJOEAR
— Meiofauna community attributes of the lower Bonny estuary was investigated under a pollution regime. This is germane because of the strategic importance of meiofauna in the food chain and the sustenance of any aquatic ecosystem. Five stations representing diverse land based activities such as refuse dump site, fuel depot, fishing-landing areas, residential housing and a station located upstream (control) were chosen for investigation. The meiofauna samples were collected from sediments in three replicate spots per station per sampling months at low tide. Sediment samples were stirred through (63-212nm) meshes to separate meiofauna and organic debris. Meiofauna samples were processed by first washing Meiofauna through a sieve of fine mesh size made of silk material, in order to wash off formation and excess silt or mud. Meiofauna samples were sorted out and identified using standard keys. Result from field studies indicate that meiofauna population abundance and diversity varied slightly between stations and seasons. There was however, no established trend. Meiofauna were generally more abundant in the rainy season than in the dry season. In general, pollution indicator meiofaunal taxa were more in all stations except the control station; the increasing impact of pollutants in the river led to a rise in the relative abundance of r-strategist genera like Chromadora. We conclude that the application of meiofaunal indices can be a useful tool for assessing the environmental quality of estuarine ecosystems.
This document summarizes a project to control invasive plant species on 565+ acres of conservation land owned by the University of Central Florida (UCF). The project aims to initially control four priority invasive species through contractual work. The land has a high potential for natural regeneration due to a diverse native seed bank and variety of habitat types. Long-term management will include continued invasive species control funded by UCF's Environmental Initiative to benefit listed species on the property and increase connectivity to other conservation lands.
Long-range migration in insects : Mechanisms and monitoring techniquesmanjeet singh
It contains the detail about orientation and mechanism of flight of Aphid, Locust, Dragonfly and Monarch butterfly and migration monitoring techniques also described.
Seasonal changes on termite foraging behaviour under different habitats in ru...Alexander Decker
1. The study examined the effect of wet and dry seasons on termite foraging behavior and abundance in cropland, forest, and grassland habitats in Rufiji District, Tanzania over one year.
2. Termite abundance was significantly different between the three habitats, with the highest abundance found in croplands. However, termite species richness was not significantly different between habitats.
3. Termite abundance was higher during the wet season than the dry season across all habitat types due to increased food and moisture availability. Species richness was also generally higher in the wet season.
This document provides an environmental assessment of the flora and fauna at Gondwana Sanctuary, a 40-hectare property in Tyagarah, Australia that is applying to convert to a community title development. A survey found no threatened plant species but identified potential for habitat development. Fauna surveys identified three vulnerable mammal species, nine regionally significant species, and that 36-50% of species in a nearby nature reserve are now present at Gondwana. The assessment concludes that while no species are currently threatened at Gondwana, continued habitat development has benefits for vulnerable and significant local species.
This document provides an overview of alien species and evolutionary ecology. It defines key terms and outlines the history of the field. It discusses the large number and impacts of alien species introductions in North America, particularly their ecological and economic costs. It explores how the introduction of alien species can lead to rapid evolution in both alien and native species through processes like dispersal adaptation, establishment in new environments, and interactions with climate change. The document also notes conservation challenges around defining alien species in a changing climate.
This study evaluated and compared the vegetative composition of four tern nesting colonies in Southern New England. Data was collected on plant species and coverage at Common Tern, Roseate Tern, and random nest and sampling sites. In total, 34 plant species were documented, with 16 being relatively common across multiple sites. The sites varied in plant species richness and prevalence of invasive species. Management of vegetation is important for maintaining suitable open nesting habitat for terns but requires a combination of removal, soil, and habitat construction methods.
Aquatic Insect Similarity Connecting Natural Wetland Habitat and Ricefield fo...Innspub Net
This study describes the relationship of the natural wetland habitats in Agusan Marsh to nearby rice fields and its implication to ecological rice production. Aquatic insects play multiple roles in the ecosystem such as predators, prey to other animals and decomposers which help in maintaining ecological balance. Results revealed that the diversity of odonates was highest in the sedge-dominated swamp among natural habitats which corresponds to the adjoining ricefields. The pattern of clustering of odonates show 3 groups; the rice-sago and rice-sedges sub-cluster, the rice-bangkal, rice-Terminalia, rice-fern, bangkal, sago and sedges sub-cluster, and the Terminalia forest as the outlier. The diversity of semi-aquatic and aquatic bugs was highest in the Bangkal forest while the lowest was in the fern-dominated swamp. The pattern of clustering shows 2 sub-clusters and the outlier Bangkal forest. On aquatic beetles, highest diversity was in the Terminalia forest. The sub-cluster consists of Terminalia habitat and rice-fern, while the other sub-cluster includes rice-Bangkal and rice-Terminalia. The resulting patterns of similarity in diversity and distribution of species in natural habitats and nearby ricefields indicate that ricefields are important temporary habitats for some aquatic insect species and serve as stepping stones for the movement of the insects.
Analyses of Community Attributes of Meiofauna Under A Pollution Regime in the...Agriculture Journal IJOEAR
— Meiofauna community attributes of the lower Bonny estuary was investigated under a pollution regime. This is germane because of the strategic importance of meiofauna in the food chain and the sustenance of any aquatic ecosystem. Five stations representing diverse land based activities such as refuse dump site, fuel depot, fishing-landing areas, residential housing and a station located upstream (control) were chosen for investigation. The meiofauna samples were collected from sediments in three replicate spots per station per sampling months at low tide. Sediment samples were stirred through (63-212nm) meshes to separate meiofauna and organic debris. Meiofauna samples were processed by first washing Meiofauna through a sieve of fine mesh size made of silk material, in order to wash off formation and excess silt or mud. Meiofauna samples were sorted out and identified using standard keys. Result from field studies indicate that meiofauna population abundance and diversity varied slightly between stations and seasons. There was however, no established trend. Meiofauna were generally more abundant in the rainy season than in the dry season. In general, pollution indicator meiofaunal taxa were more in all stations except the control station; the increasing impact of pollutants in the river led to a rise in the relative abundance of r-strategist genera like Chromadora. We conclude that the application of meiofaunal indices can be a useful tool for assessing the environmental quality of estuarine ecosystems.
This document summarizes a project to control invasive plant species on 565+ acres of conservation land owned by the University of Central Florida (UCF). The project aims to initially control four priority invasive species through contractual work. The land has a high potential for natural regeneration due to a diverse native seed bank and variety of habitat types. Long-term management will include continued invasive species control funded by UCF's Environmental Initiative to benefit listed species on the property and increase connectivity to other conservation lands.
Long-range migration in insects : Mechanisms and monitoring techniquesmanjeet singh
It contains the detail about orientation and mechanism of flight of Aphid, Locust, Dragonfly and Monarch butterfly and migration monitoring techniques also described.
Seasonal changes on termite foraging behaviour under different habitats in ru...Alexander Decker
1. The study examined the effect of wet and dry seasons on termite foraging behavior and abundance in cropland, forest, and grassland habitats in Rufiji District, Tanzania over one year.
2. Termite abundance was significantly different between the three habitats, with the highest abundance found in croplands. However, termite species richness was not significantly different between habitats.
3. Termite abundance was higher during the wet season than the dry season across all habitat types due to increased food and moisture availability. Species richness was also generally higher in the wet season.
This document provides an environmental assessment of the flora and fauna at Gondwana Sanctuary, a 40-hectare property in Tyagarah, Australia that is applying to convert to a community title development. A survey found no threatened plant species but identified potential for habitat development. Fauna surveys identified three vulnerable mammal species, nine regionally significant species, and that 36-50% of species in a nearby nature reserve are now present at Gondwana. The assessment concludes that while no species are currently threatened at Gondwana, continued habitat development has benefits for vulnerable and significant local species.
This document provides an overview of alien species and evolutionary ecology. It defines key terms and outlines the history of the field. It discusses the large number and impacts of alien species introductions in North America, particularly their ecological and economic costs. It explores how the introduction of alien species can lead to rapid evolution in both alien and native species through processes like dispersal adaptation, establishment in new environments, and interactions with climate change. The document also notes conservation challenges around defining alien species in a changing climate.
Beetles in Selected Barangays of Catarman, Northern Samar, Philippinesijtsrd
This initial research effort was conducted to identify the beetle species in five selected barangays of Catarman, Northern Samar, and describe the environmental factors that affect the presence of beetles in the study area. It used the descriptive method of research, with the purposive sampling technique as the sampling procedure utilized.Due to limitations of the researcher, a total of only seventeen 17 beetle species, belonging to eleven 11 families, were found present in different sampling sites. The most abundant of the beetle species individual was Harmonia axyridis, constituting 76.14 of the total number of beetles in the study area, while the least abundant were Alaus sp., and Diphucephala sp., each with only one 1 individual present, accounting for a relative abundance of only 0.46 . The air and soil temperature range between 27oC to 29oC. Grasses, trees, shrubs, and herbs were the common vegetation present in the study sites where beetles were found. This implies that environmental factors were conducive for the beetles to thrive in the area. The presence of different beetle species indicates the study area has a diverse beetle fauna, albeit it is only a small fraction of the total beetle species listed in the country. It is imperative that conservation efforts should be strengthened by the LGU in order to conserve not only the beetle population in the study area, but other organisms, both flora and fauna, as well. Jehosaphat C. Jazmin | Abel Alejandro U. Flores, Jr. ""Beetles in Selected Barangays of Catarman, Northern Samar, Philippines"" Published in International Journal of Trend in Scientific Research and Development (ijtsrd), ISSN: 2456-6470, Volume-3 | Issue-3 , April 2019, URL: https://www.ijtsrd.com/papers/ijtsrd22937.pdf
Paper URL: https://www.ijtsrd.com/other-scientific-research-area/other/22937/beetles-in-selected-barangays-of-catarman-northern-samar-philippines/jehosaphat-c-jazmin
Biodiversity declines with increasing distance from the equator, such that tropical regions near the equator harbor more species than temperate or polar regions further from the equator. This pattern is demonstrated by countries like Colombia, located near the equator, having over 1400 bird species, compared to places like New York at 41°N only having 105 bird species. Tropical forests at the equator can have ten times more vascular plant species than forests of equal area in temperate regions. This distribution of species can be studied under latitudinal gradients and species-area relationships.
1. There are various patterns of distribution for different species based on factors like habitat, geography, and climate. No two species have an identical distribution.
2. The causes of distribution patterns vary depending on the taxonomic level from broad families to specific genera.
3. Distribution patterns also vary depending on the spatial scale examined from global to local levels. Key factors that influence patterns include geological history, climate, food and habitat availability, competition, and human activities.
The document summarizes a study examining the vegetation that has established over four years on a fauna overpass in Brisbane, Australia. Researchers surveyed plant species present on the overpass and compared it to adjacent forest vegetation. They found 45 plant species on the overpass, with over half planted but many self-sown from local topsoil. Survival of planted species was high at 95%. Vegetation similarity to forest was moderate for groundcover but high for tree species. The developing vegetation structure is providing habitat for birds and other wildlife using the overpass to cross the road.
Koi_2015_Bulldozers vs. butterflies-conservation concernsSandy Koi
This document discusses the threats facing endangered butterflies and other invertebrates in South Florida's pine rocklands habitat from urban development and habitat loss. It describes conflicts between conservation and development, including events that destroyed colonies of the endangered Atala Hairstreak butterfly. Ongoing development threatens isolated colonies of other endangered butterflies like the Bartram's Scrub Hairstreak and Florida Leafwing that depend on the unique pine rocklands habitat and host plants. Global biodiversity loss is occurring rapidly as habitats are destroyed and wildlife is fragmented into isolated populations.
Psammophilus dorsalis agamas were studied across urban and rural areas in India to understand how they persist in urban environments. The study found:
1) The agamas' diet in both areas was mainly composed of ants, indicating flexibility.
2) Dietary overlap was high between urban and rural populations (80.3%), and between males and females in each area (80-91%).
3) Surprisingly, urban agamas had higher body mass indices than rural ones, despite consuming a greater diversity and volume of prey in rural areas.
4) Males moved more in rural areas, likely due to higher energy expenditure, but all agamas managed to hunt enough in fragmented urban habitats.
Diversity and abundance of terrestrial mammals in the northern periphery of ...Innspub Net
The Tanoé-Ehy Marsh Forest (TEMF), an unprotected forest that is about to be turn into a voluntary nature reserve is a forest block in south-eastern Côte d’Ivoire known as being of great importance for the conservation of biodiversity. But in the rainy season, that forest is largely over flooded and terrestrial mammals are likely to move to the periphery and make them vulnerable to anthropogenic threats. Our objectives are to determine the diversity, the relative abundance of terrestrial mammals and their spatial distribution in the northern periphery of the TEMF during the rainy season. We collected data by conducting recce surveys after interviews with local people. In total, we obtained 33 species among which ten primates’ species were identified. According to the recce survey, the African Civet (Civettictis civetta) and the Bushbuck (Tragelaphus scriptus) are the most common and abundant species in the study area with 12.7% and 12.0% of relative abundance respectively. In addition, six of the listed species are on the IUCN Red List, including Piliocolobus waldronae, a critically endangered species, Cercocebus lunulatus and Cercopithecus roloway endangered species, Colobus vellerosus, Phataginus tetradactyla and Phataginus tricuspis, three vulnerable species. Thus, the consideration of the periphery for the sustainable management of the TEMF is proving to be an imperative.
Koi & Daniels_2015_New Atala Life history_Florida EntomologistSandy Koi
This document summarizes a study on the life history of the Florida hairstreak butterfly (Eumaeus atala). Key findings include:
- The butterfly was once common in southeast Florida but became rare/extinct due to habitat loss and overharvesting of its host plant. A small population was discovered in 1979.
- The objective of the study was to document life history parameters like longevity, reproduction, and development to help conservation efforts.
- A captive breeding population was established and studied under controlled conditions. Wild larvae and pupae were also collected from natural areas for replenishment.
- Preliminary results found longevity was greater than previously recorded and reproductive behavior supports increased genetic exchange.
This document discusses transboundary ecosystems and species that occur in British Columbia and neighboring jurisdictions. It notes that 96% of BC's known species are transboundary, sharing range with other provinces, states or territories. Many of these transboundary species and entire ecosystems, like the antelope-brush ecosystem and Garry oak woodlands, are at high risk of disappearing from BC. The document argues that transboundary species should be protected for several reasons, including to maintain healthy ecosystems, preserve genetic diversity, support sustainable resource use, adapt to climate change, and fulfill legal/ethical obligations across borders. It concludes that current protections for transboundary species in BC are inadequate and recommendations are needed.
1. The document discusses biological invasions, defining invasive species as non-native species that harm the environment, economy, or human health.
2. It provides examples of invasive plant, animal, and microbe species like cogon grass, feral pigs, European green crabs, and water hyacinth.
3. The stages of invasion are described as introduction, colonization, establishment, and dispersal, with human transport being a major pathway of introduction.
This document discusses the importance of wildlife corridors and their status in Nepal. It notes that wildlife corridors are important as they allow the movement and migration of wildlife, provide habitat, and enable gene exchange between populations. The document reports that Nepal has identified seven corridors in the Terai Arc Landscape, and that major threats include land use change, infrastructure development, grazing, and logging. It concludes that corridor conservation has been initiated by governments and organizations, and recommends that Nepal establish policies and incentives to support connectivity conservation.
Agent Orange was a herbicide and defoliant used by the U.S. military during the Vietnam War. It was composed of equal parts of two herbicides, 2,4-D and 2,4,5-T, and was contaminated with dioxin. Agent Orange was sprayed over jungles to remove leaves and vegetation that provided cover for enemy forces. While effective as a defoliant, Agent Orange caused widespread health and environmental impacts due to its toxic dioxin contamination, including increased rates of cancer and birth defects among those exposed. The document discusses the chemical composition and effects of Agent Orange.
Cassava variety tolerance to spider mite attack in relation to leaf cyanide l...Alexander Decker
This study evaluated how cyanide content in cassava leaves influences damage from the cassava green mite, Mononychellus progresivus. Nine cassava varieties from different regions were planted and infested with M. progresivus. Mite populations and leaf damage were monitored over 55 days. Varieties differed in time taken to reach peak mite densities and damage levels. Higher cyanide content in leaves was positively correlated with greater biomass loss and mite population increase. This information can help breeders develop varieties tolerant to the cassava green mite that are also safe for human consumption.
This article lists nectar sources that have been observed and documented to be used by the atala butterfly in Florida. It provides the scientific names and families of over 30 plant species that the atala butterfly has been seen nectaring on. Maintaining populations of both the atala butterfly's larval host plant (coontie) and suitable nectar plants for adults is important for the conservation of this vulnerable species, as development continues to reduce its wild habitat in South Florida. Additional observations of atala butterfly colonies can be found online.
Understanding properties of food webs, such as their topology or stability, and the rules underlying food web structure, has been a key issue in ecology for now more than half a century. Because obtaining data on food webs has long been a hard task by itself, this research field has progressed slowly, and its dynamical aspects have seldom been empirically considered. However, technical advances, like next generation sequencing or the possibility of retrieving past ecosystems in sediment cores, have paved the way for massive data and the analysis of time series on food webs, while new models allow better predictions about food web dynamics. Making use of such existing data sets, this working group aimed at assessing the effects of biological invasions on food web topology, the fluxes of energy and nutrients throughout the network, and its ultimate effects on biodiversity. The working group has provided an integrative view on this topic, simultaneously tackling empirical, theoretical and applied aspects of biological invasions in food webs. Obvious applications will arise both from the numerous transports of invasive species and from the reshuffling of natural communities that is expected under global change scenarios. The working group comprised theoreticians and empiricists, biological invasion specialists as well as food web and host-parasite network experts, and benefited from existing experience in the field of ecoinformatics and massive data management in ecology.
Article 'Response of moths (Lepidoptera: Heterocera) to livestock grazing in Mongolian rangelands' by Khishigdelger Enkhtura, Martin Pfeiffera, Ariuntsetseg Lkhagvaa, Bazartseren Boldgiva
This document summarizes a study on biodiversity change along a gradient of human impact within the Ferlo Biosphere Reserve in northern Senegal. The study analyzed woody vegetation across 110 plots in the core area, buffer zone, and transition area. A total of 49 species across 16 families were identified. Species abundance and diversity indices revealed that the buffer zone and transition area, which experience greater human activity, had higher diversity and stand organization than the core area. The four most abundant species comprised 68% of total individuals. The study found biodiversity varied along the human impact gradient within the reserve.
1) Veterans groups filed a lawsuit alleging that a federally mandated study on Agent Orange was canceled due to White House pressure, costing $43 million.
2) Workers at chemical plants that produced Agent Orange in the 1960s reported health issues like skin diseases and liver damage from exposure.
3) Memos from the 1960s/70s show that Agent Orange spraying in Vietnam provided little military benefit and had negative environmental and psychological impacts by destroying civilian crops, but the defoliation program continued.
Distribution of ground dwelling spider genera among berseem crop at Okara dis...Innspub Net
The present study was designed to record the distribution of ground dwelling spider genera among berseem crop at Okara district. Sampling was made from Trifolium crop on fortnight basis through pitfall traps. Equal number of traps were placed in three rows e.g. along the boundary, middle of the field and centre of the field. Each trap was filled with mixture solution of alcohol and glycerin (70:30%) along with few drops of kerosene oil. After 5 days interval sample traps was collected and spider specimens were washed with distilled water and permanently stored in labeled glass vials, and brought into the Pest Control Laboratory, Departmentof Zoology, Wildlife and Fisheries, University of Agriculture, Faisalabad. Thereafter, each spider specimen was identified according to the taxonomic material and internet source. Identified data was analyzed statistically to quantify their spatial distribution. Overall maximum spatial distribution of spider population was documented in middle transect than boundary and centre of the berseem crop. It was also observed that temperature, humidity, vegetation and prey availability were the major factors that effecting the spider population. Population variations were recorded during the months of February, March and April in 2015, due to rise of temperature, decrease of humidity and availability of prey. It was concluded that despite to cosmopolitan nature, spiders have some correlation with suitable local conditions or habitat. More over, spiders are cost effective, functionally significant and play a key role in regulating decomposer population. Get the full articles at: http://www.innspub.net/jbes/distribution-of-ground-dwelling-spider-genera-among-berseem-crop-at-okara-district-pakistan/
Beetles in Selected Barangays of Catarman, Northern Samar, Philippinesijtsrd
This initial research effort was conducted to identify the beetle species in five selected barangays of Catarman, Northern Samar, and describe the environmental factors that affect the presence of beetles in the study area. It used the descriptive method of research, with the purposive sampling technique as the sampling procedure utilized.Due to limitations of the researcher, a total of only seventeen 17 beetle species, belonging to eleven 11 families, were found present in different sampling sites. The most abundant of the beetle species individual was Harmonia axyridis, constituting 76.14 of the total number of beetles in the study area, while the least abundant were Alaus sp., and Diphucephala sp., each with only one 1 individual present, accounting for a relative abundance of only 0.46 . The air and soil temperature range between 27oC to 29oC. Grasses, trees, shrubs, and herbs were the common vegetation present in the study sites where beetles were found. This implies that environmental factors were conducive for the beetles to thrive in the area. The presence of different beetle species indicates the study area has a diverse beetle fauna, albeit it is only a small fraction of the total beetle species listed in the country. It is imperative that conservation efforts should be strengthened by the LGU in order to conserve not only the beetle population in the study area, but other organisms, both flora and fauna, as well. Jehosaphat C. Jazmin | Abel Alejandro U. Flores, Jr. ""Beetles in Selected Barangays of Catarman, Northern Samar, Philippines"" Published in International Journal of Trend in Scientific Research and Development (ijtsrd), ISSN: 2456-6470, Volume-3 | Issue-3 , April 2019, URL: https://www.ijtsrd.com/papers/ijtsrd22937.pdf
Paper URL: https://www.ijtsrd.com/other-scientific-research-area/other/22937/beetles-in-selected-barangays-of-catarman-northern-samar-philippines/jehosaphat-c-jazmin
Biodiversity declines with increasing distance from the equator, such that tropical regions near the equator harbor more species than temperate or polar regions further from the equator. This pattern is demonstrated by countries like Colombia, located near the equator, having over 1400 bird species, compared to places like New York at 41°N only having 105 bird species. Tropical forests at the equator can have ten times more vascular plant species than forests of equal area in temperate regions. This distribution of species can be studied under latitudinal gradients and species-area relationships.
1. There are various patterns of distribution for different species based on factors like habitat, geography, and climate. No two species have an identical distribution.
2. The causes of distribution patterns vary depending on the taxonomic level from broad families to specific genera.
3. Distribution patterns also vary depending on the spatial scale examined from global to local levels. Key factors that influence patterns include geological history, climate, food and habitat availability, competition, and human activities.
The document summarizes a study examining the vegetation that has established over four years on a fauna overpass in Brisbane, Australia. Researchers surveyed plant species present on the overpass and compared it to adjacent forest vegetation. They found 45 plant species on the overpass, with over half planted but many self-sown from local topsoil. Survival of planted species was high at 95%. Vegetation similarity to forest was moderate for groundcover but high for tree species. The developing vegetation structure is providing habitat for birds and other wildlife using the overpass to cross the road.
Koi_2015_Bulldozers vs. butterflies-conservation concernsSandy Koi
This document discusses the threats facing endangered butterflies and other invertebrates in South Florida's pine rocklands habitat from urban development and habitat loss. It describes conflicts between conservation and development, including events that destroyed colonies of the endangered Atala Hairstreak butterfly. Ongoing development threatens isolated colonies of other endangered butterflies like the Bartram's Scrub Hairstreak and Florida Leafwing that depend on the unique pine rocklands habitat and host plants. Global biodiversity loss is occurring rapidly as habitats are destroyed and wildlife is fragmented into isolated populations.
Psammophilus dorsalis agamas were studied across urban and rural areas in India to understand how they persist in urban environments. The study found:
1) The agamas' diet in both areas was mainly composed of ants, indicating flexibility.
2) Dietary overlap was high between urban and rural populations (80.3%), and between males and females in each area (80-91%).
3) Surprisingly, urban agamas had higher body mass indices than rural ones, despite consuming a greater diversity and volume of prey in rural areas.
4) Males moved more in rural areas, likely due to higher energy expenditure, but all agamas managed to hunt enough in fragmented urban habitats.
Diversity and abundance of terrestrial mammals in the northern periphery of ...Innspub Net
The Tanoé-Ehy Marsh Forest (TEMF), an unprotected forest that is about to be turn into a voluntary nature reserve is a forest block in south-eastern Côte d’Ivoire known as being of great importance for the conservation of biodiversity. But in the rainy season, that forest is largely over flooded and terrestrial mammals are likely to move to the periphery and make them vulnerable to anthropogenic threats. Our objectives are to determine the diversity, the relative abundance of terrestrial mammals and their spatial distribution in the northern periphery of the TEMF during the rainy season. We collected data by conducting recce surveys after interviews with local people. In total, we obtained 33 species among which ten primates’ species were identified. According to the recce survey, the African Civet (Civettictis civetta) and the Bushbuck (Tragelaphus scriptus) are the most common and abundant species in the study area with 12.7% and 12.0% of relative abundance respectively. In addition, six of the listed species are on the IUCN Red List, including Piliocolobus waldronae, a critically endangered species, Cercocebus lunulatus and Cercopithecus roloway endangered species, Colobus vellerosus, Phataginus tetradactyla and Phataginus tricuspis, three vulnerable species. Thus, the consideration of the periphery for the sustainable management of the TEMF is proving to be an imperative.
Koi & Daniels_2015_New Atala Life history_Florida EntomologistSandy Koi
This document summarizes a study on the life history of the Florida hairstreak butterfly (Eumaeus atala). Key findings include:
- The butterfly was once common in southeast Florida but became rare/extinct due to habitat loss and overharvesting of its host plant. A small population was discovered in 1979.
- The objective of the study was to document life history parameters like longevity, reproduction, and development to help conservation efforts.
- A captive breeding population was established and studied under controlled conditions. Wild larvae and pupae were also collected from natural areas for replenishment.
- Preliminary results found longevity was greater than previously recorded and reproductive behavior supports increased genetic exchange.
This document discusses transboundary ecosystems and species that occur in British Columbia and neighboring jurisdictions. It notes that 96% of BC's known species are transboundary, sharing range with other provinces, states or territories. Many of these transboundary species and entire ecosystems, like the antelope-brush ecosystem and Garry oak woodlands, are at high risk of disappearing from BC. The document argues that transboundary species should be protected for several reasons, including to maintain healthy ecosystems, preserve genetic diversity, support sustainable resource use, adapt to climate change, and fulfill legal/ethical obligations across borders. It concludes that current protections for transboundary species in BC are inadequate and recommendations are needed.
1. The document discusses biological invasions, defining invasive species as non-native species that harm the environment, economy, or human health.
2. It provides examples of invasive plant, animal, and microbe species like cogon grass, feral pigs, European green crabs, and water hyacinth.
3. The stages of invasion are described as introduction, colonization, establishment, and dispersal, with human transport being a major pathway of introduction.
This document discusses the importance of wildlife corridors and their status in Nepal. It notes that wildlife corridors are important as they allow the movement and migration of wildlife, provide habitat, and enable gene exchange between populations. The document reports that Nepal has identified seven corridors in the Terai Arc Landscape, and that major threats include land use change, infrastructure development, grazing, and logging. It concludes that corridor conservation has been initiated by governments and organizations, and recommends that Nepal establish policies and incentives to support connectivity conservation.
Agent Orange was a herbicide and defoliant used by the U.S. military during the Vietnam War. It was composed of equal parts of two herbicides, 2,4-D and 2,4,5-T, and was contaminated with dioxin. Agent Orange was sprayed over jungles to remove leaves and vegetation that provided cover for enemy forces. While effective as a defoliant, Agent Orange caused widespread health and environmental impacts due to its toxic dioxin contamination, including increased rates of cancer and birth defects among those exposed. The document discusses the chemical composition and effects of Agent Orange.
Cassava variety tolerance to spider mite attack in relation to leaf cyanide l...Alexander Decker
This study evaluated how cyanide content in cassava leaves influences damage from the cassava green mite, Mononychellus progresivus. Nine cassava varieties from different regions were planted and infested with M. progresivus. Mite populations and leaf damage were monitored over 55 days. Varieties differed in time taken to reach peak mite densities and damage levels. Higher cyanide content in leaves was positively correlated with greater biomass loss and mite population increase. This information can help breeders develop varieties tolerant to the cassava green mite that are also safe for human consumption.
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This document summarizes a study examining factors that influenced changes in wild turkey populations in California from 1972 to 2013. The study used breeding bird survey data, land cover data, and data on turkey translocations and hunting to build statistical models analyzing the effects of various predictors on population change. The models found that hunting increased as populations increased, indicating hunting was not limiting populations. Land cover effects varied by scale, with urban land cover positively correlated with population change at smaller distances. Forest, grassland and agriculture were negatively correlated at an intermediate distance. Overall, wild turkey populations in California increased 10% annually during the study period. The results suggest increasing urban land cover, which provides more food, has helped wild turkey populations grow.
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Feral Swine Project Report
1. University of Nebraska - Lincoln
DigitalCommons@University of Nebraska - Lincoln
USDA National Wildlife Research Center - Staff
Publications
Wildlife Damage Management, Internet Center for
2-28-2014
Impacts from Control Operations on a
Recreationally Hunted Feral Swine Population at a
Large Military Installation in Florida
Richard Engeman
Troy Hershberger
Steve Orzell
Rodney Felix
Gary Killian
See next page for additional authors
Follow this and additional works at: http://digitalcommons.unl.edu/icwdm_usdanwrc
Part of the Life Sciences Commons
This Article is brought to you for free and open access by the Wildlife Damage Management, Internet Center for at DigitalCommons@University of
Nebraska - Lincoln. It has been accepted for inclusion in USDA National Wildlife Research Center - Staff Publications by an authorized administrator
of DigitalCommons@University of Nebraska - Lincoln.
Engeman, Richard; Hershberger, Troy; Orzell, Steve; Felix, Rodney; Killian, Gary; Woolard, John; Cornman, Jon; Romano, David;
Huddleston, Chet; Zimmerman, Pat; Barre, Chris; Tillman, Eric; and Avery, Michael, "Impacts from Control Operations on a
Recreationally Hunted Feral Swine Population at a Large Military Installation in Florida" (2014). USDA National Wildlife Research
Center - Staff Publications. Paper 1425.
http://digitalcommons.unl.edu/icwdm_usdanwrc/1425
2. Authors
Richard Engeman, Troy Hershberger, Steve Orzell, Rodney Felix, Gary Killian, John Woolard, Jon Cornman,
David Romano, Chet Huddleston, Pat Zimmerman, Chris Barre, Eric Tillman, and Michael Avery
This article is available at DigitalCommons@University of Nebraska - Lincoln: http://digitalcommons.unl.edu/icwdm_usdanwrc/
1425
3. 1 23
Environmental Science and Pollution
Research
ISSN 0944-1344
Environ Sci Pollut Res
DOI 10.1007/s11356-014-2727-9
Impacts from control operations on
a recreationally hunted feral swine
population at a large military installation
in Florida
Richard Engeman, Troy Hershberger,
Steve Orzell, Rodney Felix, Gary Killian,
John Woolard, Jon Cornman, David
Romano, Chet Huddleston, et al.
4. 1 23
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5. RESEARCH ARTICLE
Impacts from control operations on a recreationally hunted feral
swine population at a large military installation in Florida
Richard Engeman & Troy Hershberger & Steve Orzell & Rodney Felix & Gary Killian &
John Woolard & Jon Cornman & David Romano & Chet Huddleston & Pat Zimmerman &
Chris Barre & Eric Tillman & Michael Avery
Received: 17 December 2013 /Accepted: 28 February 2014
# Springer-Verlag Berlin Heidelberg (outside the USA) 2014
Abstract Feral swine were targeted for control at Avon Park
Air Force Range in south-central Florida to avert damage to
sensitive wetland habitats on the 40,000-ha base. We conduct-
ed a 5-year study to assess impacts from control to this
population that had been recreationally hunted for many years.
Control was initiated in early 2009. The feral swine popula-
tion was monitored from 2008 to 2012 using a passive track-
ing index (PTI) during the dry and wet seasons and using
recreational hunter take rates from the dry season. All three
indices showed substantial feral swine declines after
implementing control, with indices leveling for the final two
study years. Military missions and recreational hunting sea-
sons impacted temporal and spatial consistency of control
application, thereby limiting further impacts of control efforts
on the feral swine population. The PTI was also able to
monitor coyotes, another invasive species on the base, and
detect Florida black bear and Florida panther, species of
particular concern.
Keywords Animal damage . Conservation . Coyote . Feral
hog . Hunter take . Invasive species . Passive track index .
Population monitoring . Sus scrofa
Introduction
Feral swine (Sus scrofa) are highly destructive exotic animals
that degrade habitats and archeological sites, prey on native
species, and compete with native species (Choquenot et al.
1996; Engeman et al. 2013a; Seward et al. 2004; Taft 1999;
U.S. Department of Agriculture 1999). They also harbor a
number of diseases transmittable to wildlife, livestock, or
humans (e.g., Conger et al. 1999; Corn et al. 2005; Romero
and Meade 1999; Taft 1999). Swine were one of the first
invasive exotic species to establish in Florida after being
introduced to Florida by DeSoto in 1539 (Towne and
Wentworth 1950). Feral swine possesses the highest repro-
ductive potential of any large mammal in North America
(Wood and Barrett 1979; Hellgren 1999) and, with subsequent
introductions, the range in the USA continues to expand (Corn
et al. 2005; Gipson et al. 1997). Feral swine currently inhabit
many areas in such large numbers that they adversely impact
wild land and agricultural ecosystems. They have been impli-
cated by some as the single greatest vertebrate modifier of
natural plant communities (Bratton 1977; Wood and Barrett
1979). Feral swine-rooting behavior, which can overturn or
dislodge soils and native vegetation, has been shown to alter
plant populations and ultimately change plant species compo-
sition (Bratton 1977).
Habitat damage by feral swine is most pronounced in wet
environments (e.g., Choquenot et al. 1996). In Florida, many
unique wetland habitats are rapidly disappearing (Florida
Responsible editor: Philippe Garrigues
R. Engeman (*) :G. Killian
National Wildlife Research Center, USDA/APHIS/WS, 4101
LaPorte Ave, Fort Collins, CO 80521-2154, USA
e-mail: richard.m.engeman@aphis.usda.gov
T. Hershberger :S. Orzell
Avon Park Air Force Range, 29 South Blvd., Avon Park AFR,
FL 33825, USA
R. Felix :E. Tillman :M. Avery
National Wildlife Research Center, USDA/APHIS/WS, Florida Field
Station, 2820 E University Blvd, Gainesville, FL 32641, USA
J. Woolard :J. Cornman :D. Romano :C. Huddleston :
P. Zimmerman :C. Barre
USDA/APHIS/WS-Florida, 2820 E University Blvd, Gainesville,
FL 32641, USA
Present Address:
R. Felix
Eglin AFB Natural Resources-Wildlife, 107 SR 85 North, Niceville,
FL 32578, USA
Environ Sci Pollut Res
DOI 10.1007/s11356-014-2727-9
Author's personal copy
6. Natural Areas Inventory 2010). For example, only 1 % of the
original extent of seepage slopes in Florida remains (Florida
Natural Areas Inventory 2010). Avon Park Air Force Range
(APAFR) contains globally rare wetlands, plants, and plant
communities, some of which are endemic to peninsular Flor-
ida (i.e., cutthroat grass and Panicum abscissum-dominated
wetlands) (Bridges and Orzell 1999; Orzell 1997), and feral
swine rooting consistently damages some of these ecological-
ly sensitive habitats. Moreover, APAFR also holds dozens of
archeological sites eligible for inclusion in the National Reg-
ister of Historic Places, with 42 % having had some level of
feral swine disturbance (Engeman et al. 2013a). Considerable
recreational hunting takes place on the base and is popular
with the public, but feral swine damage to those plant com-
munities prompted further efforts to reduce the feral swine
population through implementation of a control program.
Thus, monitoring population changes and trends is a key
performance metric for evaluating the need for and efficacy
of management actions (Engeman et al. 2013b), control oper-
ations in this case. We carried out and report here on a
multiyear investigation to evaluate the impacts of control on
feral swine population trends at this large base.
Methods
Avon Park Air Force Range
APAFR is a 42,430-ha military installation in south-central
Florida (27°35′N, 81°16′W), that was established during
World War II for air-to-ground training and related military
missions. It contains 23,600 ha of intact natural habitats
(Orzell 1997) and experiences a seasonal subtropical climate,
conventionally divided into a winter dry and summer wet
season (Chen and Gerber 1990; Slocum et al 2010). Over
33,000 ha are open to the public on a regular basis for various
recreational activities, including hunting. The Air Force is
federally mandated to manage the natural resources of APAFR
in support of both the military readiness mission and the
preservation of the area’s natural and cultural heritage. This
latter component is significant because APAFR has numerous
US-federally listed species, many state and globally imperiled
plants, and is recognized as a significant conservation area
(Orzell 1997; Stein et al 2008).
The geologic backbone of APAFR is the Bombing Range
Ridge (BRR), a Plio-Pleistocene sand ridge running north-
south across the base. The topographic relief of this unique
geologic feature accounts for the presence and location of
many of the sensitive wetland plant communities. The
APAFR landscape varies from a low-lying elevation of
9.1 m in long-hydroperiod marshes along the Kissimmee
River to pine savanna-grasslands with embedded short-
hydroperiod marshes, to herbaceous seepage slopes and
forested seepage bays on the side slopes of the Bombing
Range Ridge, to xeric uplands on the ridge top. The ridge acts
as an unconsolidated aquifer, storing water which percolates
slowly downward until it meets an impervious layer on the
side slopes causing it to seep out on the side slopes to create
seepage slopes. These seepage slopes are found north-south
along the length of the ridge through the base, while wet
pinelands and prairies (i.e., wet prairies and flatwoods) char-
acterize much of the low-lying flatland adjacent to the BRR.
Many of the plant community types at APAFR are globally
imperiled and are dominated by endemic plant associations
found only in peninsular Florida (Bridges and Orzell 1999;
Orzell 1997).
The wet season lasts, on average, 133 days from May 21 to
October 1, while the dry season, despite being almost twice as
long, from October 2 to May 20, has only half the rainfall
(Slocum et al 2010), thereby drying soil moisture and lower-
ing water levels especially in seasonal low-elevation wetlands.
Avariety of seasonal wetland plant communities found on the
base are highly sensitive to ground disturbance (e.g., seepage
slopes, wet flatwoods, wet prairies, and peaty marshes) and
harbor many rare and endemic plants (Bridges and Orzell
1999; Florida Natural Areas Inventory 2010; Harper et al.
1998; Orzell 1997). The climate’s seasonality coupled with
the pronounced elevation gradient from wet to xeric plant
communities and the resultant fluctuations in soil moisture
causes wetland plant communities to be seasonally vulnerable
to feral swine rooting. The impact of feral swine on these
natural resources is an ecological management issue, and this
concern led to implementation of control activities to further
constrain and reduce feral swine populations beyond the ben-
efits obtained from recreational hunting (USDA 2009).
Passive tracking index methodology
We applied passive tracking index (PTI) methodology similar
in principle to the methods successfully used by Engeman
et al. (2007) to monitor feral swine in another large natural
area in Florida with the methodology customized for the
logistics and landscape at APAFR. Data collection for an
index is most efficient if observation stations can be placed
to intercept predicted daily activity of the animals (Engeman
2005; Engeman et al. 2013b). Roads and tracks through native
terrain offer many species of animals convenient travel routes
making them prime locations to place observation stations.
This often has proven to be the case for monitoring feral swine
in various places in the world including Florida (e.g., Elledge
2011; Engeman et al. 2001, 2007; Jiang et al. 2006;
Theuerkauf and Rouys 2008). Tracking plots were 0.8 km in
length and located randomly along roads throughout APAFR
(Fig. 1) as an efficient design for sampling on a large scale
(Pearson and Ruggiero 2003; Engeman et al. 2007).
Environ Sci Pollut Res
Author's personal copy
7. The surfaces of the plots were smoothed for reading tracks,
and the number of track intrusions by feral swine and other
wildlife were counted and recorded at each plot the following
day (Engeman et al. 2001, 2007). Track plot counts obtained
for analysis were “passive” because they did not involve the
use of attractants or drives to bring feral swine to the plots.
Daily counts of tracking plot intrusions were produced when
feral swine and other animals intercepted the plots during their
normal daily activity. To prepare the track plots, we used an
all-terrain vehicle (ATV) to tow a drag device or mechanical
rake to smooth the track plot surface (Fig. 2). Feral swine and
other animal tracks were easily detected in the freshly
smoothed, typically sandy, soil.
We used the same plot locations, recorded by GPS, on each
sampling occasion (e.g., Ryan and Heywood 2003; Engeman
2005). Tracking data were collected from 40 permanent plot
locations during two seasons—dry season (late fall/early
winter, December/January) and wet season (late spring/
early summer, late May/early July). It is appropriate to only
compare index values across years using data obtained from
the same season (Engeman 2005; Engeman et al. 2013b). We
collected the initial track data in December 2008 prior to
implementation of feral swine control operations, and subse-
quently twice per year from those same plots for wet
season and dry season from 2009 through 2012. Feral
swine control operations concluded just prior to the final
dry season observations in 2012.
Much of the perimeter of the base did not pose a barrier to
feral swine movement, i.e., there were no natural barriers or
wildlife-proof fences that would deter immigration. Of partic-
ular interest for feral swine management at APAFR is a private
hunting club on its north border where feral swine are
sporadically released to be hunted. Such releases represent
an artificially increased prospect for immigration to the base
beyond what would occur naturally among the feral swine
living in the vicinity. We used seven tracking plots near the
northern border to see if their index values were higher relative
to trends observed for the base as a whole, which might
indicate an immigration pulse from the neighboring property
Fig. 1 Location of tracking plots
for calculating a passive tracking
index to monitor feral swine and
sympatric species at Avon Park
Air Force Range in south-central
Florida. Shaded areas indicate
impact areas, ranges with
restricted access during military
missions and are closed to all
public access
Fig. 2 Preparation of tracking plots using a drag device (left photo) and
mechanical rake (right photo) to smooth the track plot surface
Environ Sci Pollut Res
Author's personal copy
8. prior to their diffusion throughout the base. This subset of
north perimeter plots seems a relatively small number, but
considering they each were 0.8 km in length, additional plots
could not be included from the larger set or created and still be
considered as representing the perimeter area. Further, subsets
of tracking plots have been successfully used in other appli-
cations to look at immigration and contrast with results from a
larger overall area (e.g., Engeman et al. 2003a, b).
One of the benefits from using passive tracking plots to
intercept the daily activities of animals without an attractant is
it offers the opportunity to simultaneously monitor a variety of
animal species (e.g., Engeman 2005). Thus, we took advan-
tage of the possibility for monitoring other invasive species,
and even rare species (although for very uncommon species,
the method serves primarily as an efficient means to detect
presence in a large area, rather than to monitor population
trends). While our focus was to monitor feral swine, we felt it
important to document the broader utility of the PTI method
and to obtain and report information on other invasive or rare
species that would be of broad management interest in the
general geographic area. In particular, coyotes (Canis latrans)
are invasive in Florida (Schmitz and Brown 1994) and have
been increasing their populations and expanding ranges in
Florida (Coates et al. 2002; Wooding and Hardisky 1990).
Coyotes could pose a threat to livestock grazing on the base,
game species such as deer (Odocoileus virginianus) and wild
turkeys (Meleagris gallopavo), threatened and endangered
species, and other species of concern (e.g., Mastro et al.
2012). Besides feral swine, we were most interested in ancil-
lary monitoring of coyotes as potentially significant invasive
predators, as well as any rare species detected by the tracking
plots.
We calculated PTI for feral swine for dry season
during 2008–2012 and for wet season during 2009–
2012 by applying the indexing paradigm of Engeman
(2005), specifically applied to tracking plots for feral swine
(Engeman et al. 2001, 2007). The mean measurement across
plots was calculated for each day, and the index values were
the means of the daily means (Engeman 2005):
PTI ¼
1
d
Xd
j¼1
1
sj
Xsj
i¼1
xij
where xij represents the number of feral swine intrusions at the
ith tracking plot on the jth day, d is the number of days of
observation, and sj is the number of plots contributing data on
the jth day. SAS PROC VARCOMP, using restricted maxi-
mum likelihood estimation (REML) (SAS Institute 2004),
was used to calculate the variance components (Searle et al.
1992) needed in the variance estimation formula (Engeman
2005). Independence among plots or among days is not
required for these calculations (Engeman 2005). Variance
estimates were incorporated in Z tests for assessing differences
between PTI values (Engeman 2005).
Hunter take rates
Hunter take (catch per effort) is a widely applied method for
assessing relative abundance of wildlife, including wild swine
(Boitani et al. 1994; Fernandez-Llario et al. 2003). Catch per
effort is often formulated as pigs per hunter-day or pigs per
100 hunter-days. Large sampling areas and sample sizes help
reduce bias in hunt indices (Siren et al. 2004). Fortunately,
APAFR is a large controlled area with a history of consider-
able public hunting in designated areas. The bombing ranges
are off limits to the public.
Recreational hunters entering APAFR must check in at the
gate, and then again upon leaving when take is recorded.
Thus, reliable data were available on hunter take and the
corresponding number of hunter-days for each hunting season
each year. A take-per-effort index was calculated for each
season and standardized as take per 100 hunter-days.
Recreational hunting at APAFR is carried out during var-
ious hunting seasons through the year. Many are short special
seasons that can be designated at any time during the year with
no temporal regularity from year to year. Three hunting sea-
sons, one each for three hunting methods, archery, muzzle
loader guns, and standard guns (rifles), occur in the same
timeframes each year. This temporal regularity across years
is crucial, because, as with the PTI data, index values for a
particular measurement method are appropriately examined
across years when measurements are made in the same
timeframe each year in the same manner (Engeman et al.
2013b). Archery and muzzle loader gun seasons are short
and involve far fewer hunters specializing in these firearms
than partake in the lengthy (7–8 weeks) standard gun season.
Consequently, we used only the data from the annual standard
gun season to calculate a hunter take index each year because
only that season was at the same time each year and involved a
sufficient number of hunter-days each year (∼5,000–
6,500 hunter-days) to be considered reliable for indicating
population trends (see Siren et al. 2004).
Feral swine removal during control operations
The initial PTI data collection began in the dry season
(December) in 2008 and served as the precontrol baseline for
the dry season time frame. There was no corresponding
baseline for the wet season time periods. Operational
removal of feral swine (in contrast with recreational hunting)
was initiated in January 2009 and continued through Septem-
ber 2012. Feral swine were removed by agreement with U.S.
Department of Agriculture/Wildlife Services (WS), the federal
agency responsible for managing conflicts with wildlife (U.S.
Environ Sci Pollut Res
Author's personal copy
9. Department of Agriculture/Animal and Plant Health Inspec-
tion Service, U.S. Department of Agriculture/Forest Service
and Department of Interior/Bureau of Land Management
1997), using only approved and humane methods to euthanize
animals, which conform to guidelines in the 2013 Re-
port of the American Veterinary Medical Association
Panel on Euthanasia (2013) and set forth as agency
policy in USDA/APHIS/WS Directive 2.505.
Swine were primarily removed by capture in pen traps and
sharpshooting. After identification of the most favorable lo-
cations to carry out control activities, baiting with soured corn
was initiated to condition the feral swine to feeding at bait
sites. Once feral swine were consistently feeding at a bait site,
traps were set up and the feral swine conditioned to feeding in
and around the open trap. After feral swine consistently were
entering the pen trap to feed, the trap would be set. These
custom-designed traps were collapsible for portability, but
exhibiting extreme durability, and able to capture groups of
feral swine, including the largest individuals (Engeman et al.
2010). Feral swine were also removed by sharpshooting op-
portunistically, especially in bombing range areas, which are
unavailable to public access. Feral swine trapping was not
permitted in public hunting areas during hunting seasons,
although trapping was allowed on bombing ranges during
these times if the ranges were not active.
Results
Feral swine
The monitoring results using the PTI and hunter take are
summarized in Table 1, with the PTI summarized for the base
as a whole, and also for the tracking plots near the north
perimeter where pulse immigrations from feral swine releases
adjacent to the base were suspected to occur. Hunter take rates
are also given in Table 1 and are part of the dry season period
observations.
By the conclusion of 2010, the dry season and hunter
take observations represented two full years of control activ-
ities (all of 2009 and 2010) and the wet season observa-
tions represented 1.5 years of control activities (all of 2009
and half of 2010). By 2010, the dry season PTI and
hunter take indices had each shown successive sharp de-
creases (Table 1) from the 2008 precontrol values (Z>3.13,
p<0.0017, across both indices), and, similarly, the wet
season PTI values had declined sharply in 2010 from 2009,
the initial year for wet season measurements (Z=3.94,
p=0.0001). Since 2010, all PTI values remained on average
less than one intrusion observed per 0.8 km tracking plot per
day for both the dry season and wet season seasons.
During this time, both the wet season and dry season
PTI values for the north perimeter followed suit and
were always lower than the base-wide values, except in 2009
when the north perimeter values for wet season and dry
season both spiked well above the values for the base as a
whole. For the precontrol observation in 2008, the dry
season PTI for the north perimeter was 18 % less than that for
the base as a whole. However, for 2009, it was 48 % higher. In
all subsequent years, it was again lower than for the base as a
whole: 27, 68, and 18 % less than the base-wide PTI, respec-
tively, for years 2010–2012. This discrepancy in pattern be-
tween the dry season PTI values for the base as a whole
and the north perimeter plots suggests an immigration pulse
may have occurred in 2009. Although precontrol observations
were not available for the wet season, a similar pattern
emerged, with the base-wide PTI dropping precipitously from
2009 (after a half year of control activities) to 2010 (after
1.5 years of control activities). Again, only in 2009 was the
wet season PTI value from the north perimeter plots
higher than the value from the base as a whole. Relative to
the base-wide index values, the wet season index values
from the north perimeter tracking plots were 38 % higher in
2009, and then 14, 34, and 34 % lower, respectively, for the
years 2010–2012. Considering the feral swine habitat near the
north border is not as optimal as further south on the base
Table 1 Passive tracking index (PTI) and hunter take results for feral
swine at Avon Park Air Force Range for the dry season (late fall–early
winter) and wet season (late spring–early summer) observation
periods from 2008 to 2012. The 2008 dry season results form a
baseline prior to implementation of operational swine control
Year Feral swine passive track index (PTI) (study initiated in dry season 2008) Take/100 hunter-days (gun season)
Wet season Dry season
Whole base N perim Whole base N perim Whole base
2008* – – 2.38 1.94 3.15
2009 2.58 3.57 1.63 2.42 2.53
2010 0.43 0.37 0.92 0.67 1.59
2011 0.64 0.42 0.60 0.19 2.23
2012 0.73 0.48 0.91 0.75 2.18
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10. where most of the wetlands occur, and considering the north
perimeter PTI values were uniformly less than the base-wide
PTI value in every year for both seasons except 2009, the data
suggest a possible immigration pulse along the north border in
2009.
The PTI for the dry season period reached a minimum
in 2011, 75 % less than the precontrol value (Z=4.31,
p<0.0001). In 2012, the dry season PTI increased to a
value nearly identical (Z=0.0.02, p=0.98) to that in 2010 (still
62 % less than for precontrol). Feral swine control was
discontinued at the end of September 2012, meaning this final
dry season PTI was obtained a quarter year after the end
of control operations. While not detectable statistically, the
wet season PTI had also increased from 2011 to 2012
while control was ongoing (Z=0.32, p=0.75). The wet
season PTI and hunter take indices each reached their mini-
mums in 2010. For all three indices, there was a slight
rebound/leveling off in the last year or two of the study, with
the concluding index values still well lower than during the
first 2 years (2008 and 2009).
Coyotes
The PTI indexing results for coyotes are summarized in Table 2
where the tracking plot data suggest coyotes increased to a
somewhat level population on the base. Substantial differences
in PTI values between seasons for wild canids were expected
due to potential differences in activity between seasons and
illustrate why it would be inappropriate to compare index
values from dry and wet seasons (e.g., Allen et al.
2011; Allen et al. 2013; Engeman 2005). However, it is fitting
to examine the trends indicated among the two seasons
(Engeman 2005). The dry season index value for the
initial year (2008) was considerably less than all subsequent
years’ PTI values, with each of those being at least 59 %
higher than for 2008. From 2009 through 2012, the dry season
PTI fluctuated without showing an increasing or decreasing
trend. The same was true for all wet season values which were
obtained in this same span of years from 2009 to 2012.
Rare wildlife species
The track plot observations in June 2010 included one occur-
rence of Florida black bear (Ursus americanus floridanus)
tracks, while the track plot observations in June 2011 included
an observation of Florida panther (Puma concolor coryi)
tracks. The Florida panther is federally listed as an endangered
species, while the Florida black bear was recently removed
from state listing as threatened. However, the subpopulation in
which APAFR lies is at or below the “minimum subpopula-
tion objective” for state bear management, and APAFR is
particularly important for this subpopulation (Florida Fish
and Wildlife Conservation Commission 2012). While one
set of tracks on one plot during one sampling occasion does
not present an indexing opportunity for either species, these
observations along with previous occasional sightings of bears
and panthers at APAFR document at least temporary presence
and could provide information to consider in managing natu-
ral resources.
Discussion
While recreational hunting can impose a source of mortality
on a feral swine population, hunters typically do not target all
population segments equally, potentially limiting the severity
of population reduction (Braga et al. 2010; Festa-Bianchet
2007; Keuling et al. 2013; Plhal et al. 2011). In contrast,
operational control targets all demographics and typically is
cost-effective (Engeman et al. 2003a, b; 2004; 2007, 2010),
yet it requires consistent application to be effective. In Florida,
feral swine are ubiquitous and pose a constant threat from
immigration into an area in which their populations have been
reduced by control efforts. That, coupled with feral swine
having the highest reproductive potential of any large mam-
mal in North America (Wood and Barrett 1979; Hellgren
1999), means the beneficial effects of feral swine population
reduction can be quickly undone. Here, we have documented
feral swine population declines after implementing control
over a large area that had been receiving consistent hunting
pressure for many years. Although having an ecologically
similar nearby 40,000-ha site without feral swine control to
use for comparison was impossible, there could be no other
rational explanation for the parallel decreases among the three
population indices. That is, there were no significant disease
outbreaks, climatic events, or any other potentially population
decimating factors that occurred during the course of the
study. Importantly, we also were able to identify factors that
deterred maximal feral swine population reductions, in
Table 2 Passive tracking index (PTI) results for coyotes at Avon Park
Air Force Range for the dry season (late fall–early winter) and
wet season (late spring–early summer) observation periods from 2008 to
2012
Year Coyote passive track index (PTI)
Wet season Dry season
2008a
– 0.80
2009 0.70 1.29
2010 0.49 1.27
2011 0.56 1.48
2012 0.69 1.36
a
The study was initiated in early dry season 2008
Environ Sci Pollut Res
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11. particular anthropogenic induced immigration and limitations
on the consistency with which control could be applied.
Our study provides strong evidence that sporadic immigra-
tion pulses of feral swine can occur from the hunting club
property bordering APAFR on the north. As this study was
concluding, a feral swine-proof fence was being installed
along the base’s northeast border to deter some such immigra-
tion (Fig. 1). A feral swine-proof fence was also constructed
specifically to protect an archeological site of significant
importance on the base (Engeman et al. 2013a). The consid-
eration of fencing raises an interesting dilemma for managing
feral swine to protect sensitive wetland plant communities. In
some circumstances, it may be possible to use feral swine-
proof fencing to prevent immigration, while eradicating all
feral swine existing on the interior. Conducted iteratively, this
approach in conjunction with the necessary control operations
to eliminate feral swine inside fenced areas could result in
relatively permanent, large feral swine-free areas. However,
the upfront cost for fencing and concomitant control to re-
move feral swine inside the fences may represent the equiva-
lent of many years of control operations. Additionally, fences
constructed in sandy substrates in a region regularly subjected
to hurricanes might require considerable maintenance. More-
over, fence lines are problematic to maintain in fire-frequented
ecosystems such as in Florida, where fire lanes needed to
protect fences further disrupt ecosystem function at a land-
scape level comparable to hog rooting ecosystem damage
(Hutchison and Roberts 2009). This would be especially true
if fencing were used to protect specific, relatively small plant
community sites. A careful economic evaluation would be in
order for determining optimal strategies for keeping feral
swine populations from affecting vulnerable resources.
While the index values for feral swine remained consider-
ably lower from 2010 through 2012 than in 2008 (for dry
season and hunter take) and in 2009 (all three indices), the
minimum values for all three indices occurred in 2010 or
2011, and increased from 2011 to 2012 for dry season and
successively from 2010 to 2012 for the wet season index
values. Various factors could account for these observations.
To achieve and maintain maximal efficacy, control should not
only be maintained over years, but it should also be consis-
tently applied within each year. Otherwise, the rapidity with
which feral swine can immigrate and/or reproduce can reduce
the efficacy and efficiency of the control program. In the day-
to-day efforts of removing feral swine at APAFR, a challenge
existed to manage trapping schedules within periods of active
military missions where access to substantial areas within
APAFR are prohibited or restricted due to safety and risk
management concerns (Fig. 1). Moreover, during hunting
seasons, trapping was restricted to areas of APAFR closed to
hunting, which primarily were the impact areas where military
operations are focused. This resulted in frequent periods when
and areas where control could not be exercised. Baiting and
trapping efforts require revisiting sites at timed intervals to
monitor activity, replenish baits, set/reset traps, and remove
trapped animals. If conflicts between the planned baiting/
trapping and range schedules or hunting seasons occurred,
the feral swine removal effort had to be modified or curtailed.
Military training activity at APAFR increased during the
study, as did the number of special hunting seasons each year.
A partial solution to constraints on feral swine trapping in-
volved the use of automated bucket feeders at potential trap
sites within impact areas to maintain attracting feral swine
without needing human presence. Automated bucket feeders
needed to be checked once a week compared to traditional
baiting which involves daily visits. This approach helped
alleviate some of the logistical access issues to impact areas,
but not for carrying out control within the much larger public
access portion of the base during hunting seasons where such
activities would have been in direct conflict with hunting
(Fig. 1).
Besides protecting endangered plant species and rare plant
communities, the reduction of feral swine populations at
APAFR may help protect the Florida panthers and Florida
black bears that reside on or pass through the base, as detected
by the PTI observations. Pseudorabies (PRV), a density-
dependent disease, is carried by feral swine in many areas
and is usually fatal in secondary species, including felids and
bears (Dow and McFerran 1963; Glass et al. 1994; Schultze
et al. 1986; Zanin et al. 1997). Moreover, 42 of 60 feral swine
(70 %) from APAFR tested from 2009 to 2011 were seropos-
itive for pseudorabies (M. Milleson, personal communica-
tion). While this percent does not translate directly to active
infection, it does indicate a high likelihood that active infec-
tion is present at any given time in the feral swine population
at APAFR. Feral swine are a common prey item for Florida
panthers (Maehr et al. 1990) and lethality to Florida panthers
from preying upon an infected feral swine has long been
suspected (Roelke et al. 1993) and demonstrated (Glass
et al. 1994). Similarly, bears are also highly susceptible to
PRV-induced mortality (Schultze et al. 1986; Zanin et al.
1997), and could face similar risks as the panthers.
There are many ecological and economically valid con-
cerns for reducing feral swine populations at APAFR includ-
ing the protection of sensitive plant communities,
archeological sites, native grass seed sources, livestock graz-
ing pastureland, and a variety of rare plant and animal species.
We saw in this study that a control program can reduce a feral
swine population that already had been annually subjected
recreational hunting pressure for many years. We suspect from
the leveling of the three population indices during the final
2 years of the study that the effects of control were not fully
realized due to the inconsistencies with which it was applied
each year. A limited degree of fencing is now in place on the
base as barriers to feral swine, but only the area encompassed
by fencing to protect a significant archeological site, if
Environ Sci Pollut Res
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12. diligently maintained, is fully protected (Engeman et al.
2013a). This fencing was installed at an approximate cost of
$18,000 USD (Engeman et al. 2013a) indicating the level of
financial commitment that would be needed to exclude feral
swine from larger areas of the base with fencing. The optimal
application of resources for protecting the natural treasures on
this large base will undoubtedly pose management resource
allocation dilemmas.
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