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LIST OF PUBLICATIONS
1. Books
1970 Gorillas. Arthur Barker, London.
1974 Horses, Asses and Zebras in the Wild. David and Charles, Newton Abbot.
1981 Ancestors for the Pigs: taxonomy and phylogeny of the genus Sus. (Technical Bulletin No. 3, Department of
Prehistory, Research School of Pacific Studies, ANU), VII + 96pp.
1989 A Theory of Primate and Human Evolution. Oxford University Press. (2nd ed. 1991).
2001 Primate Taxonomy. Washington: Smithsonian Institution Press.
2004 (with David W.Cameron) Bones, Stones and Molecules. Amsterdam etc.: Elsevier Academic Press.
2008 Extended Family: Long Lost Cousins. A personal look at the history of primatology. Arlington, VA:
Conservation International.
2011 (with Peter Grubb) Ungulate Taxonomy. Johns Hopkins University Press, Baltimore.
2. Edited Books
1989 Skeptical. (Edited by Don Laycock, David Vernon, Colin Groves and Simon Brown.) Australian Skeptics.
3. Papers
1963 Multivariate Analysis of the Skulls of Asiatic Wild Asses. Ann. Mag. N.H. 6:329-336.
1965 Skull Changes Due to Captivity in Certain Equidae. Z.f. Saugetierk. 31:44-6.
1965 Description of a New Subspecies of Rhinoceros, from Borneo. Saugetierkundl. Mitt. 13:128-131.
1966 (with J.R. Napier) Catalogue of the skeletal material of Gorilla in the British Isles. J. Zool. 148:153-161.
1966 (with F. Ziccardi and A. Toschi). Sull'asino Selvatico Africano. Rich Zool. Appl. Caccia Suppl. 5(1):1-30.
1967 Ecology and Taxonomy of the Gorilla. Nature 213:890-3.
1967 The Nomenclature of the Eastern Lowland Gorilla. Nature 215:1172.
1967 The Rhinoceroses of Southeast Asia. Saugetierkundl. Mitt. 15:221-237.
1967 (with D.L. Harrison). The Taxonomy of the Gazelles of Arabia. J. Zool. 152:381-7.
1967 Geographic Variation in the Black Rhinoceros. Z.f. Saugetierk. 32:267-276.
1967 Geographic Variation in the Hoolock or White-browed Gibbon. Folia Primat. 7:276-283.
1967 (with V. Mazak). Taxonomic Problems of Asiatic Wild Asses; with the description of a new subspecies. Z.f.
Saugetierk. 32:321-355.
1968 Simia leucophaea F. Cuvier, 1807 (Mammalia): Proposed validation under the plenary powers. Bull. Zool.
Nomen. 25:36-7.
1968 The Classification of the Gibbons (Primates, Pongidae). Z.f. Saugetierk. 33:239-246.
1968 A New Subspecies of Gibbon from Northern Thailand. Proc. Biol. Soc. Wash. 81:625-8.
1969 Hylobates lar (Linneaus, 1771), H. entelloides (I. Geoffroy St. Hilaire, 1842) and H. Hoolock (Harlan, 1834),
(Mammalia Pongidae): proposal to place these names on the official list of specific names in Zoology. Bull.
Zool. Nomen. 25:162-4.
1969 (with J. R. Napier). Dentition and diet in Australopithecines. Proc. VIII Congr. Anthrop. Ethnol. Sci.
Tokyo/Kyoto 3:273-6.
1969 The Taxonomy of the Anoa. Beaufortia (Amsterdam) 17:1-12.
1969 The Smaller Gazelles of the Genus Gazella de Blainville, 1816. Z.f. Saugetierk. 34:38-60.
1970 The Forgotten Leaf-eaters, and the Phylogeny of the Colobinae. In J.R. Napier and P.H. Napier (eds) The Old
World Monkeys. Academic Press.
1970 (with R.W. Thorington). Annotated classification of the Cercopithecoidea. In Napier and Napier, op. cit.
1970 Taxonomic and Individual Variation in Gibbons. Symp. Zool. Soc. London, No. 26:127-134.
1970 Population Systematics of the Gorilla. J. Zool. 161:287-300.
1971 Request for a declaration modifying Article 1, so as to exclude names proposed for domestic animals from
Zoological Nomenclature. Bull. Zool. Nomencl. 27:269-272.
1971 Pongo Pygmaeus. Mammalian Species No. 4:1-6.
1971 Distribution and Place of Origin of the Gorilla. Man, N.S. 6:44-51.
1971 Geographic and individual variation in Bornean Gibbons, with remarks on the systematics of the subgenus
Hylobates. Folia Primat. 14:139-153.
1971 Species Characters in Rhinoceros Horns. Z.f. Saugetierk. 36:238-252.
1971 Systematics of the Genus Nycticebus. Proc. 3rd Int. Congr. Primat., Zurich 1970, 1:44-53.
1972 Ceratotherium Simum. Mammalian Species No. 8:1-6.
1972 (with J. Sabater Pi). The importance of higher Primates in the diet of the Fang of Rio Muni. Man, 7:239-243.
1972 Systematics and Phylogeny of Gibbons. In D. Rumbaugh (ed) Gibbon and Siamang, 1:1-89. Karger, Basel.
1972 (with F. Kurt) Dicerorhinus Sumatrensis. Mammalian Species No. 21:1-6.
1972 Phylogeny and Classification of Primates. In R.N. T-W-Fiennes (ed) Pathology of Simian Primates, 1:11-57.
Karger, Basel.
1973 Notes on the Ecology and Behaviour of the Angola Colobus (Colobus angolensis) in N.E. Tanzania. Folia
Primat. 20:12-26.
1973 (with N. K. Humphrey). Asymmetry in Gorilla Skulls: evidence of lateralised brain function? Nature 244:53-4.
2
1974 A note on the systematic position of the Muntjac (Artiodactyla, Cervidae). Z.f. Saugetierk. 39:369-372.
1974 (with P. Grubb). A New Duiker from Rwanda. Rev. Zool. Afr. 88:189-196.
1974 (with P. Andrews and J.F.M. Horne). The Tana Colobus and Mangabey. Oryx 12:565-575.
1974 New Evidence on the Evolution of the Apes and Man. Vest. Ustr. Ust. Geol. 49:53-6.
1974 Taxonomy and Phylogeny of Prosimians. In R.D. Martin, G.A. Doyle and A.C. Walker (eds) Prosimian
Biology, pp. 449-473. Duckworth.
1975 Taxonomic notes on the White Rhinoceros Ceratotherium simum (Burchell, 1817). Saugetierkundl. Mitt.
23:200-212.
1975 (with P. Andrews and J.F.M. Horne). Ecology of the Tana River Floodplain, Kenya. J. E. Afr. N.H. Soc. Nat.
Mus. 151:1-31.
1975 Notes on the Gazelles. 1. Gazella rufifrons and the Zoogeography of Central African Bovidae. Z.f. Saugetierk.
40:308-319.
1975 Comment on the proposed conservation of Pan Oken, 1816 and Panthera Oken, 1816. Bull.Zool.Nomen. 32:68.
1975 (with V. Mazak). An approach to the taxonomy of the Hominidae: Gracile Villafranchian Hominids of Africa.
Casopis Pro Min. Geol. 20:225-247.
1976 (with P. Andrews). Gibbons and Brachiation. In D.Rumbaugh (ed) Gibbon and Siamang 4:167-218. Karger.
1976 The Apes. Dyn 3:29-36.
1976 River of Monkeys. Hemisphere, 20(9):28-32.
1976 The Origin of the Mammalian Fauna of Sulawesi (Celebes). Z.f. Saugetierkunde 41:201-216.
1976 (with H. Hemmer and P. Grubb). Notes on the Sand Cat, Felis margerita Loche, 1858. Z.f. Saugetierkunde
41:286-303.
1976 Taxonomy of Moschus (Mammalia: Artiodactyla) of the Indian region. J. Bombay N. H. Soc. 72:662-676.
1977 (with Alan G. Goodall). The Conservation of Eastern Gorillas. In H.S.H. Prince Rainier III of Monaco and
Geoffrey H. Bourne (ed), Primate Conservation, pp. 599-637. Academic Press, New York.
1977 Hylobates lar (Linnaeus, 1771), H. entelloides (I. Geoffroy St. Hilaire, 1842) and H. Hoolock (Harlan, 1834),
(Mammalia, Hylobatidae): proposal to place these names on the Official List of Specific Names in Zoology.
Bull. Zool. Nomencl. 34:75-9.
1977 Sima syndactyla Raffles, 1821 (Mammalia: Hylobatidae); proposal to give this name precedence over Sima
gibbon, C. Miller, 1779. Bull. Zool. Nomencl. 34:104-5.
1977 Comments on request for a declaration modifying Article I so as to exclude names proposed for domesticated
animals from Zoological Nomenclature. Bull. Zool. Nomencl. 34:137-9.
1978 Phylogenetic and population systematics of the Mangabeys (Primates: Cercopithecoidea). Primates 19:1-34.
1978 (with L.C.Rookmaaker). The taxonomic status of the Dwarf Blue Sheep (Artiodactyla: Bovidae).
Saugetierkundl. Mitt. 26:177-183.
1978 (with C. Smeenk). On the Type material of Cervus nippon Temminck, 1836; with a revision of Sika Deer from
the main Japanese islands. Zool. Meded. Leiden 53:11-28.
1978 The extinct Cape Rhinoceros, Diceros bicornis bicornis (Linnaeus, 1758). Saugetierkundl. Mitt. 26:117-28.
1978 (with V. Mazak and P.H.J. van Bree). On a skin and skull of the Bali Tiger, and a list of preserved specimens of
Panthera tigris balica (Schwarz, 1912). Z.f. Saugetierkunde 43:108-113.
1978 Comments on the application concerning Simia syndactyla, Raffles 1821 (Mammalia: Hylobatidae). Bull. Zool.
Nomencl. 35:76.
1978 Reply to Van Gelder. Bull. Zool Nomencl. 36:9-10.
1979 (with Kenhelm W. Stott, Jr). Systematic relationships of Gorillas from Kahuzi, Tshiaberimu and Kayonza.
Folia Primatol. 32:161-179.
1979 Comment on Rightmire, 'Implications of Border Cave skeletal remains for late Pleistocene human evolution.'
Current Anthrop. 20:29.
1979 A View of Race. Hemisphere 23:36-8.
1979 What does it mean to be conscious? Behav. Brain. Sci. 1:575-6.
1980 Notes on the Systematics of Babyrousa (Artiodactyla, Suidae). Zoologische Mededelingen. Leiden 55:29-46.
1980 (with C. Guerin). Le Rhinoceros sondaicus annamiticus (Mammalia, Perissodactyla) d'indochine: distinction
taxinomique et anatomique; relations phyletiques. Geobios 13:199-208.
1980 Speciation in Macaca: the view from Sulawesi. In Donald D. Lindburg (ed) The Macaques: studies in ecology,
behaviour and evolution, pp. 84-124. Van Nostrand Reinhold Company, New York.
1980 The Chinese Mountain Cat (Felis bieti). Carnivore, 3:35-41.
1980 A further note on Moschus. J. Bombay N.H. Soc. 77:130-3.
1981 (with D.P. Willoughby). Studies on the taxonomy and phylogeny of the genus Equus. 1. Subgeneric
classification of the recent species. Mammalia 45:321:354.
1981 Promise unfulfilled: a rejoinder to Peter Wilson. Canberra Anthrop. 4:101-9.
1981 Notes on the Gazelles. 2. Subspecies and clines in the Springbok (Antidorcas). Z.f.Saugetierkunde 46:189-197.
1981 Comment on Shipman et al., 'Butchering of Giant Geladas at an Acheulian Site', Current Anthrop. 22:265.
1981 Systematic relationships in the Bovini (Artiodactyla, Bovidae). Z.f. Zool. Syst.Evolutionsforschung 19:264-278.
3
1981 Notes on the Gazelles. 3. The Dorcas Gazelles of North Africa. Annali del Museo Civico di Storia Naturale di
Genova 83:455-471.
1981 Comment on Johnson, 'Bonobos: generalised hominid prototypes or specialised insular dwarfs?' Current
Anthrop. 22:366.
1981 Survey of the Tana river Colobus and Mangabey. National Geographic Society Research Reports 13:235-242.
1982 Asian Rhinoceroses - down but not out. Malayan Naturalist 36:11-22.
1982 (with P. Grubb). The species of Muntjac (genus Muntiacus) in Borneo: unrecognised sympatry in tropical deer.
Zoologische Mededelingen, Leiden 56:203-216.
1982 (with P. R. Groves). The Lions of India. Hemisphere 26(4):206-209.
1982 The skulls of Asian Rhinoceroses: wild and captive. Zoo Biology 1:251-261.
1982 The Systematics of Tree Kangaroos (Dendrolagus; Marsupialia, Macropodidae). Aust. Mammal. 5:157-186.
1982 A note on geographic variation in the Indian Blackbuck (Antilope cervicapra Linnaeus, 1758). Rec. Zool. Surv.
India 79:489-503.
1982 Bos gaurus H. Smith, 1827 (Mammalia, Artiodactyla): proposed conservation. Bull. Zool. Nom. 39:279-280.
1982 Antilope depressicornis H. Smith, 1827, and Anoa quarlesi Ouwens, 1910 (Mammalia, Artiodactyla): proposed
conservation. bull. Zool. Nom. 39:281-2.
1983 Phylogeny of the living species of Rhinoceros. Z. Zool. Syst. u. Evolutionsforschung 21:293-313.
1983 Notes on the Gazelles. 4. The Arabian Gazelles collected by Hemprich and Ehrenberg. Z.f. Saugetierkunde
48:371-381.
1983 Geographic variation in the Barasingha or Swamp Deer (Cervus duvauceli). J. Bombay N.H. Soc. 79:620-629.
1983 (with P.H. Napier). Simia fascicularis Raffles, 1821 (Mammalia, Primates): Request for the suppression under
the Plenary Powers of Simia aygula Linnaeus, 1758, senior synonym. Bull. Zool. Nom. 40:117-8.
1983 (with S. Chakraborty). The Calcutta collection of Asian Rhinoceros. Rec. Zool. Surv. India 80:251-263.
1983 (with P. Grubb). Notes on the Taxonomy of the Deer (Mammalia, Cervidae) of the Philippines. Zool. Anz. Jena
210:119-144.
1983 (with T. Flannery). The Mammals of West Irian. Hemisphere, 27:378-84.
1984 A new subspecies of Sable antelope, Hippotragus niger (Harris, 1838). Rev. Zool Afr. 97:821-8.
1984 (with P. M. Monappa). Elephants at Work. Hemisphere 28(5):285-291.
1984 Of Mice and Men and Pigs in the Indo-Australian Archipelago. Canberra Anthropology, 7:1-19.
1984 Cranial and dental characteristics in the systematics of Old World Felidae. Carnivore, 5 (for 1982):35-46.
1984 A new look at the taxonomy and phylogeny of the gibbons. In H.Preuschoft et al.(ed) The Lesser Apes 542-561.
1984 Pigs East of the Wallace Line. J. Société des Océanistes, 39:105-119.
1984 (with W.A. Laurie and E.M. Lang) Rhinoceros unicornis. Mammalian species No. 211:1-6.
1985 Mammal Faunas and the Palaeogeograhy of the Indo-Australian Region. Courier Forschungsinstitut
Senckenberg No. 69 pp. 267-273.
1985 (with J. Sabater Pi). From Ape's Nest to Human Fix-point. Man 20:22-447.
1985 (with D.M. Lay). A new species of the genus Gazelles from the Arabian Peninsula. Mammalia 49:27-36.
1985 An Introduction to the Gazelles. Chinkara, 1, 1:4-10.
1985 (with V. Weitzel). The Nomenclature and Taxonomy of the Colobine Monkeys of Java. Int J.Primat.6:399-409.
1985 The Case of the Pygmy Gorilla: a cautionary tale for cryptozoology. Cryptozoology 4:37-44.
1985 Plio-Pleistocene Mammals in Island Southeast Asia. Mod. Quaternary Res. S.E. Asia 9:43-54.
1985 (with L.B. Holthuis). The Nomenclature of the Orang Utan. Zoo. Meded. Leiden 59:411-7.
1986 (with Feng Zuojian). The status of Musk Deer from Anhui Province, China. Acta Theriol. Sin 6:101-6.
1986 The Yahoo, the Yowie, and Reports of Australian Hairy Bipeds. Cryptozoology 5:47-54.
1986 Systematics of the Great Apes. Comparative Primate Biology 1:187-217.
1986 The taxonomy, distribution and adaptations of Recent Equids. In R.H. Meadow and H.P. Uerpmann (eds),
Equids in the Ancient World, pp. 11-65.
1986 (with P.Grubb) Relationships of Living Deer. In C.Wemmer (ed), Biology and Management of the Cervidae, 21-
59.
1986 On the Highland Cuscuses of New Guinea currently referred to Phalanger vestitus. In M. Archer, (ed) Possums
and Opossums: Studies in Evolution 559-567.
1986 On the Cuscuses of the Phalanger orientalis group from Indonesian Territory. In M. Archer, (ed) Possums and
Opossums: Studies in Evolution 569-579.
1987 Monkey Business with Red Apes. J. Hum. Evol. 16:537-542.
1987 (with J.F. Eisenberg and K. MacKinnon). Tapire. In Grzimeks Enzyklopadie, 4:598-608.
1987 Obituary: John Russell Napier (1917-1987). J.Hum.Evol. 16:533-5.
1988 (with M. Lyons and A. Kovarch). A survey of beliefs on creation and evolution; or, did Julius Caesar Really
Exist? The Skeptic 8, 4:19-24.
1988 (with J. McCarron-Benson, A. Meek and B. Puza). Creation Week in Canberra. The Skeptic 8, 4:6-9, 40.
1988 (with R.H. Eaglen). Systematics of the Lemuridae (Primates, Strepsirhini) J. Hum. Evol. 17:513-538.
1988 The Evolutionary Ecology of the Hominoidea. An.Psicol. 39:87-98.
4
1988 (with G.G. Scott & K.C. Richardson). Osteological differences of the skulls of Lasiorhinus latifrons Owen,
1845 and Vombatus ursinus Shaw, 1800 (Marsupialia: Vombatidae). Aust. J. Zool. 36:599-609.
1989 Feral Mammals of the Mediterranean Islands: documents of early domestication. pp. 46-58 in J. Clutton-Brock,
(ed) The Walking Larder. London: Unwin Hyman.
1989 [Articles in Skeptical (ed. Laycock, Vernon, Groves and Brown): 'Introduction: Why be Skeptical?'; 'Aliens
Among Us'; 'Channelling' (with W. Grey)' 'Clairvoyance'; 'Creationism'; 'Dolphins'; 'The Mystic East'; 'Near
Death Experiences'; 'The New Age'; 'Noah's Ark'; 'Prophecy'; 'Racism'; 'Reincarnation' (with W. Grey); 'Shamans
and Sorcerers'; 'Unknown Animals'; 'Velikovsky'; 'Von Daniken'.]
1989 (with G.G. Scott and K.C. Richardson). Skull morphometrics of Lasiorhinus latifrons (Owen, 1845)
(Marsupialia: Vombatidae). Rec.S.Aust.Mus. 23:1-5.
1989 A catalogue of the genus Gazella. In A. Dixon & D. Jones eds., Conservation and Biology of Desert Antelopes,
193-8.
1989 Hominidae. Pp.910-916 in Fauna of Australia: Vol. 1B, Mammalia. (D.W. Alton & B.J. Richardson, eds.).
AGPS, Canberra.
Equidae. Pp.1039-1043 in ibid. (with *J. Giles). Suidae. Pp.1044-1049. (with *J.F. Bishop). Cervidae.
Pp.1054-1060. Bovidae. Pp.1061-1066.
1989 (with T. Flannery, K. Aplin and M. Adams.) Revision of the New Guinean genus Mallomys (Muridae:
Rodentia), with descriptions of two new species from subalpine habitats. Rec. Aust. Mus., 41:83-105.
1989 The origin of human species: what we have learned about human evolution in the 130 years since On the origin
of species. The Medical Journal of Australia 151:677-688.
1989 Brief Communication: Weights of Wild Orang Utans. Amer.J.Phys.Anthrop. 81:1-3.
1989 Natural selection and intelligent ancestors. Mankind, 19:76-82.
1989 The gazelles of the Arabian peninsula. In A.H. Abu-Zinada, P.D. Goriup and I.A. Nader, eds., Wildlife
Conservation and Development in Saudi Arabia, 237-48.
1990 (with P.Grubb). Muntiacidae. In G.A. Bubenik & A.B. Bubenik (eds) Horns, Pronghorns and Antlers, 132-68.
1990 (with Wang Yingxiang). The gibbons of the subgenus Nomascus (Primates, Mammalia). Zool.Res. 11:147-54.
1990 The centrifugal pattern of speciation in Meganesian rainforest mammals. Mem.Qd.Mus. 28:325-8.
1990 (with T. Flannery). Revision of the families and genera of bandicoots. In J.H. Seebeck, R.L. Wallis, P.R.
Brown and C.M. Kemper, eds., Bandicoots and Bilbies, 1-11.
1990 Out of Africa - here we go again. In L. Freedman, ed., Is our Future limited by our Past?: Proceedings of the
Third Conference of the Australasian Society for Human Biology, pp. 13-25.
1990 (with R. Markham). Weights of wild Orang Utans. Amer. J. Phys. Anthrop. 81:1-3.
1990 The taxonomy of buffaloes. Buffalo Bulletin, 9:65-8.
1990 "Primates" in Wheldon Owen's Encyclopaedia of Mammals, Wheldon Owen, Sydney.
1990 (with L. Christidis). A Method for Identifying Night Monkeys (Aotus) by Simple Karyotyping and External
Appearance. Australian Primatology, 5:11-12.
1991 (with J.D. Paterson). Testing hominoid phylogeny with the PHYLIP program. J.Hum.Evol. 20:167-83.
1991 (with I. Tattersall). Geographical variation in the Fork-marked Lemur, Phaner furcifer (Primates,
Cheirogaleidae). Folia primatol. 56:39-49.
1991 Genes, Genitals and Genius:The Evolutionary Ecology of Race. In P.O'Higgins, ed., Human Biology: An
Integrative Science: Proc. Fourth Conference Australasian Society for Human Biology, pp. 419-432.
1992 How old are subspecies? A tiger's eye view of human evolution. Archaeology in Oceania, 27:153-160.
1992 Gorillas - where and what? Gorilla, 15,2:2,3.
1992 (with C. Westwood and B.T. Shea). Unfinished Business: Mahalanobis and a Clockwork Orang. Journal of
Human Evolution, 22:327-340.
1993 Bad Medicine for Wildlife. The Skeptic, 13(1):12-14, 17.
1993 Catalogue of Primates in the Collection of the Museum of Victoria, Melbourne (with S. Bricknell). Australian
Primatology 7(4):20-27.
1993 Catalogue of Primates in the Macleay Museum, Sydney University. Australian Primatology 7(3):8-9.
1993 (with D.W.Cameron). Pliocene Hominins: hunters and/or scavengers? Perspectives in Human Biology 3:23-31.
1993 (with M.Karami). A mammal species new for Iran: Gazella gazella Pallas, 1766 (Artiodactyla: Bovidae).
J.Sci.I.R.Iran, 4:81-89.
1993 The Chinkara (Gazella bennetti) in Iran, with the description of two new subspecies. J.Sci.I.R.Iran 4:166-178.
1993 Speciation in Living Hominoid Primates. In W. H. Kimbel and L. B. Martin (eds.), Species, Species Concepts
and Primate Evolution. New York: Plenum Press.
1993 Testing rhinoceros subspecies by multivariate analysis. Pp. 92-100 in O.A. Ryder (ed.) Rhinoceros Biology and
Conservation. Zoological society of San Diego.
1993 Orders Monotremata, Dasyuromorphia, Peramelemorphia, Notoryctemorphia, Diprotodontia and Primates. Pp.
13-14, 29-62 and 243-278 in D.E. Wilson and D.A.M. Reeder (eds.), Mammal Species of the World: a
taxonomic and geographic reference. Smithsonian Institution Press.
1993 (with A. Greth, D. Williamson, G. Schwede, M. Vassart). Bilkis gazelle in Yemen - status and taxonomic
relationships. Oryx, 27:239-244.
5
1993 (with R. Angst and C. Westwood). The status of Colobus polykomos dollmani Schwarz. International Journal
of Primatology, 14:573-586.
1993 Why do Langurs kill each others' babies? Australian Primatology, 7:2-7.
1993 Human origins; The Neanderthals; and Dating the Past. Pp. 33-52, 68 and 74-75, The Illustrated History of
Humankind, Vol. 1, The First Humans. Harper San Francisco.
1994 The origin of modern humans. Interdisciplinary Science Reviews, 19:23-34.
1994 Morphology, habitat and taxonomy. Pp 39-59 in Lee Boyd & Katherine A.Houpt (eds.), Przewalski’s Horse: the
History and Biology of an Enadangered Species. Albany, NY: SUNY Press.
1994 (with A.K.Kes Hillman-Smith). Diceros bicornis. Mammalian Species, no.455, 8pp.
1995 Domesticated and Commensal Mammals of Austronesia and their Histories. Pp. 152-163 in The Austronesians:
Historical and Comparative Perspectives (P. Bellwood, J.J. Fox and D. Tryon, eds.). Canberra: Department of
Anthropology, ANU.
1995 (with John W.H.Trueman). Lemurid systematics revisited. Journal of Human Evolution, 28:427-437.
1995 (with H.Mendelssohn and Y.Yom-Tov). Gazella gazella. Mammalian Species, no.490, 7pp.
1995 (with H.Mendelssohn and Y.Yom-Tov). Gazella dorcas. Mammalian Species, no.491, 6pp.
1995 On the nomenclature of domestic animals. Bull.Zool.Nomencl., 52:137-141.
1995 (with C.R.Westwood). Skulls of the Blaauwbok, Hippotragus leucophaeus. Z.f.Säugetierkunde, 60:314-8.
1995 (with Wang Yingxiang and Peter Grubb). Taxonomy of Musk-deer, genus Moschus (Moschidae, Mammalia).
Acta Theriologica Sinica, 15:181-197.
1995 Microtaxonomy and its implications for captive breeding. Pp 24-28 in U.Ganslosser, J.K.Hodges &
W.Kaumanns (eds.), Research and Captive Propagation. Fürth: Filander Verlag.
1996 (with J.Shoshani, E.L.Simons & G.F.Gunnell). Primate phylogeny: morphological vs molecular results.
Molecular Phylogenetics and Evolution, 5:102-154.
1996 Taxonomic diversity in Arabian gazelles: the state of the art. Pp 8-39 in A.Greth, C.Magin & M.Ancrenaz
(eds.), Conservation of Arabian Gazelles. National Commission for Wildlife Conservation and Development,
Riyadh, Saudi Arabia.
1996 Australopithecus afarensis Johanson, 1978 (Mammalia, Primates): proposed conservation of the specific name.
Bulletin of Zoological Nomenclature, 53:24-7.
1996 (with A.Gentry & J.Clutton-Brock). Proposed conservation of usage of 15 mammal specific names based on
wild species which are antedated by or contemporary with those based on domestic anaimals. Bulletin of
Zoological Nomenclature, 53:28-37.
1996 (with P.M.Goonan & R.D.Smith). Karyotype polymorphism in the Slender Loris (Loris tardigradus). Folia
primatologica, 65:100-9.
1996 The nomenclature of the Tanzanian Mangabey and the Siberut Macaque. Australian Primatology, 10, 4:2-5.
1996 Great Apes: the conflict of gene-pools, conservation and personhood. Perspectives in Human Biology 2:31-36.
1996 Hovering on the brink: nearly but not quite getting to Australia. Perspectives in Human Biology, 2:83-7.
1996 From Ussher to Slusher; from Archbish to Gish; or, not in a million years... Archaeol. Oceania 31:145- 151.
1996 The taxonomy of the Asian Wild Buffalo from the Asian mainland. Zeitschrift für Säugetierkunde 61:327-338.
1997 Major biogeographic regions of the world. Section of article “Biosphere”. Pp 1160-1167 (vol.14) in The New
Encyclopaedia Britannica: Macropaedia, 15th edition.
1997 (with G.B.Schaller, G.Amato & K.Khounbouline). Rediscovery of the wild pig Sus bucculentus. Nature
386:335.
1997 Taxonomy of wild pigs (Sus) of the Philippines. Zoological Journal of the Linnaean Society, 120:163-191.
1997 Taxonomy and phylogeny of Primates. In A.Blancher, J.Klein & W.W.Socha (eds.), Molecular Biology and
Evolution of Blood Group and MHC Antigens in Primates. Berlin: Springer Verlag, Berlin Heidelberg and New
York, pp.3-23.
1997 The taxonomy of Arabian Gazelles. In K.Habibi, A.H.Abuzinada & I.A.Nader (eds.), The Gazelles of Arabia,
Riyadh: National Commission for Wildlife Conservation and Development, Publication No.29, Eng.Ser., 24-51.
1997 Thinking about evolutionary change: the polarity of our ancestors. In G.A.Clark and C.M.Willermet (eds),
Conceptual Issues in Modern Human Origins. Aldine de Gruyter, N.Y., 1997, pp 319-326.
1997 Species concept in palaeoanthropology. Perspectives in Human Biology, 3:13-20.
1997 Leopard-cats, Prionailurus bengalensis (Carnivora: Felidae) from Indonesia and the Philippines, with the
description of two new subspecies. Zeitschrift für Säugetierkunde, 62:330-338.
1998 The proximal ulna from Klasies River. Journal of Human Evolution, 34:119-121.
1998 Systematics of Tarsiers and Lorises. Primates, 39:13-27.
1998 (with M.Goodman, C.A.Porter, J.Czelusniak, S.L.Page, H.Schneider, J.Shoshani and G.Gunnell). Toward a
phylogenetic classification of Primates based on DNA evidence complemented by fossil evidence. Molecular
Phylogenetics and Evolution, 9:585-598.
1998 (with H.Mendelssohn and B.Shalmon). A new subspecies of Gazella gazella from the southern Negev. Isreal
Journal of Zoology, 43:209-215.
1998 Obituary: Prudence Hero Napier (1916-1997). International Journal of Primatology, 19:203-205.
6
1999 Die Nashörner - Stammesgeschichte und Verwandtschaft. In Anonymous (ed.), Die Nashörner: Begegnung mit
urzeitlichen Kolossen, 14-32. Furth: Filander Verlag.
1999 (with T.F.Flannery). A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new
species and subspecies. Mammalia, 62:367-396.
1999 The advantages and disadvantages of being domesticated (A Keynote Address). Perspectives in Human
Biology, 4:1-12.
1999 Australopithecus garhi: a new-found link? Reports of the National Center for Science Education, 19, 3:10-13.
1999 Nomenclature of African Plio-Pleistocene hominins. Journal of Human Evolution, 37:869-872.
1999 (with A.Thorne). The terminal Pleistocene and early Holocene populations of northern Africa. Homo, 50,
3:249-262.
2000 (with P.Grubb, J.P.Dudley & J.Shoshani). Living African elephants belong to two species: Loxodonta africana
(Blumenbach, 1797) and Loxodonta cyclotis (Matschie, 1900). Elephant, 2, 4:1-4.
2000 (with P.Grubb), Do Loxodonta cyclotis and L.africana interbreed? Elephant, 2, 4:4-7.
2000 What are the elephants of West Africa? Elephant, 2, 4:7-8.
2000 (with P.Grubb). Are there Pygmy Elephants? Elephant, 2, 4:8-10.
2000 Phylogenetic relationships within recent Antilopini (Bovidae). In E.S.Vrba and G.B.Schaller (eds.), Antelopes,
Deer, and Relatives, 223-233. New Haven & London: Yale University Press.
2000 (with G.B.Schaller). The phylogeny and biogeography of the newly discovered Annamite artiodactyls. In
E.S.Vrba and G.B.Schaller (eds.), Antelopes, Deer, and Relatives, 261-282. New Haven & London: Yale
University Press.
2000 Phylogeny of the Cercopithecoidea. In P.Whitehead and C.J.Jolly, eds., Old World Monkeys, 77-98.
Cambridge: Cambridge University Press.
2000 (with O.A.Ryder). Systematics and phylogeny of the horse. In A.T.Bowling and A.Ruvinsky (eds.), The
Genetics of the Horse, 1-24. Wallingford (Oxon., U.K.) & New York: CABI Publishing.
2000 (with A.Thorne). The affinities of the Klasies River Mouth remains. In J.S.Chisholm (ed.), Towards
Consilience: Perspectives in Human Biology, 5:43-53.
2000 (with M.Chech, A.Thorne & E.Trinkaus). A new reconstruction of the Shanidar 5 cranium. Paléorient 25:143-
146.
2000 The genus Cheirogaleus: unrecognised biodiversity in Dwarf Lemurs. International Journal of Primatology,
21:943-962.
2000 La spéciation chez les primates. Primatologie, 3:213-236.
2001 Mammals in Sulawesi: where did they come from and when, and what happened to them when they got there?
Pp 333-342 in I.Metcalfe, J.M.B.Smith, M.Morwood & I.Davidson (eds.), Faunal and Floral Migrations and
Evolution in SE Asia-Australasia. Lisse/Abingdon/Exton (PA)/Tokyo: A.A.Balkema.
2001 Time and taxonomy. Ludus Vitalis, 9:91-96.
2001 Why taxonomic stability is a bad idea, or Why are there so few species of Primates (or are there?). Evolutionary
Anthropology, 10:192-198.
2001 (with A.Braun, P.Grubb, Yang Q. & Xia L.). Catalogue of the Musée Heude collection of mammal skulls. Acta
Zootaxonomia Sinica, 26:608-660.
2002 (with A.Meder). A model of gorilla life history. Australasian Primatology, 15:2-15.
2002 (with A.Braun & P.Grubb). Rediscovery of the type specimen of Bubalus mindorensis Heude, 1888.
Mammalian Biology, 67:246-249.
2002 (with P.Grubb). Appendix C – Revision and classification of the Cephalophinae. Pp 703-728 in V.J.Wilson,
Duikers of Africa: Masters of the African Forest Floor. Bulawayo: Chipangali Wildlife Trust.
2002 Taxonomy of living Equidae. Pp 94-107 in P.D.Moehlman (ed.), Equids: Zebras, Asses and Horses. Gland,
Switzerland: IUCN.
2002 (with A.Kitchener). New insights into the taxonomy of Macaca pagensis of the Mentawai Islands, Sumatra.
Mammalia, 66:533-542.
2003 The tarsiers of Sulawesi. Pp 179-195 in P.C.Wright, E.L.Simons & S.Gursky (eds.), Tarsiers: Past, Present, and
Future. New Brunswick: Rutgers U.P.
2003. (with A.B.Taylor). Patterns of mandibular variation in Pan and Gorilla. Journal of Human Evolution, 44:529-
561.
2003 (with J.Harding). Morphology, morphometrics and taxonomy, in Field and Laboratory Methods in Primatology,
Joanna M. Setchell and Deborah J. Curtis (eds), Cambridge University Press, Cambridge, pp 140-157.
2003 (with D.Bulbeck, D.Rayner and P.Raghavan). The Contribution of South Asia to the Peopling of Australasia and
the relevance of Basel’s Naturhistorisch Museum anthropological collection to the project aims, Bulletin der
Schweizerischen Gesellschaft fur Anthropologie / Bulletin de la Societe Suisse d’Anthropologie, 2: pp 49-70
2003 (with R.Stumpf, J.Polk, J.Oates, W.Jungers, C.Heesy and J.Fleagle). Patterns of Diversity in Gorilla Cranial
Morphology, in Gorilla Biology: A Multidisciplinary perspective, Andrea Taylor and Michele Goldsmith (eds),
Cambridge University Press, Cambridge, pp 35-61.
2003 A History of Gorilla Taxonomy, in Gorilla Biology: A Multidisciplinary perspective, Andrea Taylor and
Michele Goldsmith (eds), Cambridge University Press, Cambridge, pp 15-33.
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2003 Taxonomy of ungulates of the Indian subcontinent. Journal of Bombay natural History Society, 100:341-362.
2004 (with D.Cameron) Bones, Stones and Molecules: “Out of Africa” and Human Origins, pp 395. Elsevier Inc.
Academic Press, Burlington, MA.
2004 Charles Oxnard and the aye-aye: morpometrics, cladistics, and two very special primates. In Shaping Primate
Evolution: Form, Function, and Behavior, Fred Anapol, Rebecca Z. German, Nina G. Jablonski (eds),
Cambridge University Press, Cambridge, UK, pp 368-377.
2004 Taxonomy and biogeography of primates in Vietnam and surrounding regions. In Conservation of Primates in
Vietnam, Tilo Nadler, Ulrike Streicher and Ha Thang Long (eds), Haki Publishing, Hanoi, pp 15-20.
2004 (with H.Schulze) Asian Lorises, Taxonomic problems caused by Illegal Trade. In Conservation of Primates in
Vietnam, Tilo Nadler, Ulrike Streicher and Ha Thang Long (eds), Haki Publishing, Hanoi, pp 33-36.
2004 (with C. Bell) New investigations on the taxonomy of the zebras genus Equus, subgenus Hippotigris.
Mammalian Biology, 69: 182-196.
2004 (with A. Gentry & J. Clutton-Brock). The naming of wild animal species and their domestic derivatives. Journal
of Archaeological Science, 31: 645-651.
2004 (with R.L.Pan). Cranial variation among the Asian colobines, In Shaping Primate Evolution: Form, Function,
and Behavior, Fred Anapol, Rebecca Z. German, Nina G. Jablonski (eds) 1st edition, Cambridge University
Press, Cambridge, UK, pp 45-65.
2004 (with Pan, R. and C.E.Oxnard). Relationships between the fossil Colobine Mesopithecus pentelicus and extant
Cercopithecoids, based on dental metrics. American Journal of Primatology, 62:287-299.
2004 (with E.Meijaard) A taxonomic revision of the Tragulus mouse-deer (Artiodactyla). Zoological Journal of the
Linnean Society, 140: 63-102.
2004 (with E.Meijaard). The Biogeographical Evolution and Phylogeny of the genus Presbytis. Primate Report, 68:
71-90.
2004 (with C.Pitra, J. Fickel & E.Meijaard). Evolution and phylogeny of old world deer. Molecular Phylogenetics
and Evolution, 33: 880-895.
2004 (with E.Meijaard). Morphometrical relationships between South-east Asian deer (Cervidae, tribe Cervini):
evolutionary and biogeographic implications. Journal of Zoology, London, 2004:179-196.
2004 (with T.Geissmann and C.Roos). The Tenasserim Lutung, Trachypithecus barbei (Blyth, 1847) (Primates:
Cercopithecidae): Description of a live specimen, and a reassessment of phylogenetic affinities, taxonomic
history, and distribution, Contributions to Zoology, 73: 271-282.
2004 The What, Why and How of Primate Taxonomy. International Journal of Primatology 25:1105-1126.
2005 The taxonomy of Primates in the laboratory context. In The Laboratory Primate, Sonia Wolfe-Coote, ed.,
Elsevier Academic Press, 3-15.
2005 Domestic and wild mammals: naming and identity. In Animal Names, A. Minelli, G. Ortalli & G. Sangha,
eds, Venice, Istituto Veneto di Scienze, Lettere ed Arti, 151-157.
2005 Geographic variation within Eastern Chimpanzees (Pan troglodytes cf schweinfurthii Giglioli, 1872).
Australasian Primatology, 17, 2: 19-46.
2005 (with E.Meijaard). Interspecific variation in Moschiola, the Indian Chevrotain. Raffles Bulletin of Zoology,
12:413-421.
2005 The genus Cervus in eastern Eurasia. European Journal of Wildlife Research, 52:14-22 and DOI
10.1007/s10344-005-0011-5.
2005 Orders Monotremata, Dasyuromorphia, Peramelemorphia, Notoryctemorphia, Diprotodontia and Primates in
Mammal Species of the World: A Taxonomic and Geographic Reference. Don E Wilson & DeeAnn M. Reader
(eds) 3rd edition, Johns Hopkins University Press, Baltimore, pp 1-2, 22, 23-37, 38-42, 43-70, 111-184.
2005 (with K.Helgen). Biodiversity in Sri Lanka and the Western Ghats. Science, 308:199.
2005 (with A.Mootnick). A New Generic Name for the Hoolock Gibbon (Hylobatidae). International Journal of
Primatology, 26:971-975.
2006 (with A B. Rylands, R.A.Mittermeier, L. Cortes-Ortiz &J J.H Hines). Taxonomy and distributions of
Mesoamerican Primates. In New Perspectives in the Study of Mesoamerican Primates, A.Estrada, P. A. Garber,
M.S.M Pavelka & L. Luecke, eds, Springer, 29-79.
2006 (Articles) Biogeography; Cladistics; Fossey; Orangutan; Primate Taxonomy; Yeti. In Encyclopaedia of
Anthropology, H. James Birx, ed., Sage Publications (Thousand Oaks, etc).
2006 (with E.Meijaard). The geography of mammals and rivers in mainland Southeast Asia. In Primate Biogeogaphy:
Progress and Prospects, S.M.Lehman & J.G.Fleagle, eds, Springer, 305-329.
2006 (with S.Merker) Tarsius lariang: A New Primate Species from Western Central Sulawesi. International Journal
of Primatology, 27: 465-485.
2006. Taxonomy and Biogeography of the Primates of Western Uganda. In Primates of Western Uganda, N.E.Newton-
Fisher, H.Notman, J.D.Paterson & V.Reynolds, eds, Springer, 3-20.
2006. From Calpe to Catanduanes: Babewynes, apes, marmesettes and othere dyverse bestes. In The Barbary
Macaque: Biology, Management and Conservation, J.K.Hodges & J.Cortes, eds., Nottingham University Press,
1-15.
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2006 Humans among Primates. In J.R.Merrick, M.Archer, G.M.Hickey & M.S.Y.Lee (eds.), Evolution and
Biogeography of Australasian Vertebrates. Oatlands, NSW: Australian Scientific Publishing.
2006 (with D. Argue, D. Donlon and R. Wright) Homo floresiensis: Microcephalic, pygmoid, Australopithecus, or
Homo? Journal of Human Evolution, 51:360-374.
2006 (with J.H.Mazak). A taxonomic revision of the tigers (Panthera tigris) of Southeast Asia. Mammalian Biology
71:268-287.
2007 (with D.Zinner, C.Roos, J-L.Fauusser & Y.Rumpler). Disputed taxonomy classification of sportive lemurs
(Lepilemur) in NW Madagascar. Lemur News, 12:53-56.
2007. The endemic Uganda Mangabey, Lophocebus ugandae, and other members of the albigena-group (Lophocebus).
Primate Conservation 22: 123-128.
2007 The taxonomic diversity of the Colobinae of Africa. Journal of Anthropological Sciences, 85:7-34.
2007 (with K. M. Helgen). Craniodental characters in the taxonomy of Propithecus. International Journal of
Primatology, 28: 1363-1383.
2007 Species concepts and speciation: facts and fantasies. In W. Henke & I.Tattersall (eds.), Handbook of
Palaeoanthropology, three: 1861-1879. Berlin etc: Springer.
2007 (with C.Smeenk). The nomenclature of the African wild ass. Zool. Med. Leiden 81:121-135.
2007 Current views on the taxonomy and zoogeography of the genus Sus. In U.Albarella, K.Dobney, A.Ervynck &
P.Rowley-Conwy (eds), Pigs and Humans: 10,000 Years of Interaction, 15-29. Oxford: Oxford University Press.
2007 (with S.Mukherjee). Geographic variation in jungle cat (Felis chaus Schreber, 1777) (Mammalia, Carnivora,
Felidae) body size: is competition responsible? Biological Journal of the Linnean Society, 92:163–172.
2007 The Homo floresiensis controversy. Hayati Journal of Biosciences, 123-126.
2008 Walking with Hobbits. Australasian Science, March 2008: 16-19.
2008 Extended Family: Long Lost Cousins. A personal look at the history of primatology. Arlington, Virginia:
Conservation International.
2008 (with M.Shekelle & D.Brandon Jones). Taxonomic history of the tarsiers, evidence for the origins of Buffon's
tarsier, and the fate of Tarsius spectrum Pallas, 1778. In Primates of the Oriental Night (M Shekelle et al., eds.),
LIPI Press, 1-12.
2008 (with M. Shekelle, S. Gursky, I.Neri-Arboleda & A.Nietsch). A method for multivariate analysis and
classification of tarsier tail tuft.In Primates of the Oriental Night (M Shekelle et al., eds.), LIPI Press, 71-84.
2008 (with I.Maryanto). Craniometry of slow lorises (genus Nycticebus) of insular Southeast Asia.In Primates of the
Oriental Night (M Shekelle et al., eds.), LIPI Press, 115-122.
2008 (with J.F.Oates & P.D.Jenkins). The type locality of Pan troglodytes vellerosus (Gray, 1862), and implications
for the nomenclature of West African chimpanzees. Primates (published online: DOI 10.1007/s10329-008-0116-
z).
2008 Meiri, S, Meijaard, E, Wich, S et al 2008, 'Mammals of Borneo- small size on a large island', Journal of
Biogeography, vol. 35, no. 6, pp. 1087-1094.
2008 (with R.A.Mittermeier, J.U.Ganzhorn, W.R.Konstant, K.Glander, I.Tattersall, A.B.Rylands, A.Hapke,
J.Ratsimbazafy, M.I.Mayor, E.E.Louis Jr., Y.Rumpler, C.Schwitzer & R.M.Rasoloarison). Lemur diversity in
Madagascar. International Journal of Primatology 29:1607-1656.
2009 (with M.Shekelle, S.Merker & J.Supriatna). Tarsius tumpara: a new tarsier species from Siau Island, North
Sulawesi. Primate Conservation 23:55-64.
2009 (with C.Rajapaksha & K.Manamendra-Arachchi) The taxonomy of the endemic golden palm civet of Sri Lanka.
Zoological Journal of the Linnean Society 155,: 238–251.
2009 The A-Line Przewalski horse. News Biosphere Reserve "Askania Nova" 11:97-103.
2009 (with M.C.Westaway) The mark of ancient Java is on none of them. Archaeology in Oceania, 44:84-95.
2009 (with J.Jayewardene). The wild buffalo of Sri Lanka. Taprobanica, 1:55-62.
Editorial. Taprobanica 1:78-79.
2009 with Rajapaksha, C & Manemandraarachchi, K The taxonomy of the endemic golden palm civet of Sri Lanka.
Zoological Journal of the Linnean Society, 155:238-251.
2010 (with P.Fernando & J.Robovsky). The sixth rhino: a taxonomic reassessment of the critically endangered
Northern White Rhinoceros. PLoS One 5, 4:e9703 (1-15).
2010 (with Helen J Chatterjee, Simon YW Ho & Ian Barnes). Estimating the phylogeny and divergence times of
primates using a supermatrix approach. BMC Evolutionary Biology 2009, 9:259 doi:10.1186/1471-2148-9-259.
2010 Leslie, D, Groves, C & Abramov, A 2010, 'Procapra przewalskii (Artiiodactyla: Bovidae)', Mammalian Species
(Online), 42, no. 860: 124-137.
2010 (with Cécile Callou & Jacques Cuisin) The Rediscovery of Buffon’s Tarsier. Int J Primatol 31:1055–1070.
2011 (with Myron Shekelle). The Genera and Species of Tarsiidae. Int J Primatol 31:1071–1082.
2011 The taxonomy and phylogeny of Ailurus. In Red Panda: Biology and Conservation of the First Panda
(A.R.Glatston, Ed.). Academic Press, 101-124.
2011 Tocheri, M, Solhan, C, Orr, C et al., Ecological divergence and medial cuneiform morphology in gorillas.
Journal of Human Evolution, 60:174-184.
9
2011 Groves, C & Leslie, D, 'Rhinoceros sondaicus (Perissodactyla: Rhinocerotidae)', Mammalian Species (Online),
43, no. 887: 190-208.
2011 Groves, C & Grubb, P. Ungulate Taxonomy. Johns Hopkins University Press, Baltimore.
2011 (articles in) Wood, B. (ed.), Wiley-Blackwell Encyclopedia of Human Evolution, Wiley-Blackwell Publishing
Inc, Chichester, West Sussex, UK..
2012 Speciation in hominin evolution. In African Genesis: Perspectives in Hominin Evolution (S.C.Reynolds &
A.Gallagher, eds.). Cambridge University Press, 45-62.
2012 Species concept in Primates. American Journal of Primatology, 1-5.
2012 Moschidae. In Handbook of Living Mammals (D.E.Wilson & R.Mittermeier, eds.), Lynx Edicions, in
combination with Conservation International and IUCN, 2.
2012 (with D.M.Leslie). Bovidae. In Handbook of Living Mammals (D.E.Wilson & R.Mittermeier, eds.), Lynx
Edicions, in combination with Conservation International and IUCN, 2.
2013 (with S.Gippoliti). “Taxonomic inflation” in the historical context of mammalogy and conservation. Hystrix:
Italian Journal of Zoology Online-first: http://www.italian-journal-of-mammalogy.it/article/view/8685/pdf.
2013 The nature of species: a rejoinder to Zachos et al. Mammalian Biology 78:7-9.

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Colin Groves CV

  • 1. LIST OF PUBLICATIONS 1. Books 1970 Gorillas. Arthur Barker, London. 1974 Horses, Asses and Zebras in the Wild. David and Charles, Newton Abbot. 1981 Ancestors for the Pigs: taxonomy and phylogeny of the genus Sus. (Technical Bulletin No. 3, Department of Prehistory, Research School of Pacific Studies, ANU), VII + 96pp. 1989 A Theory of Primate and Human Evolution. Oxford University Press. (2nd ed. 1991). 2001 Primate Taxonomy. Washington: Smithsonian Institution Press. 2004 (with David W.Cameron) Bones, Stones and Molecules. Amsterdam etc.: Elsevier Academic Press. 2008 Extended Family: Long Lost Cousins. A personal look at the history of primatology. Arlington, VA: Conservation International. 2011 (with Peter Grubb) Ungulate Taxonomy. Johns Hopkins University Press, Baltimore. 2. Edited Books 1989 Skeptical. (Edited by Don Laycock, David Vernon, Colin Groves and Simon Brown.) Australian Skeptics. 3. Papers 1963 Multivariate Analysis of the Skulls of Asiatic Wild Asses. Ann. Mag. N.H. 6:329-336. 1965 Skull Changes Due to Captivity in Certain Equidae. Z.f. Saugetierk. 31:44-6. 1965 Description of a New Subspecies of Rhinoceros, from Borneo. Saugetierkundl. Mitt. 13:128-131. 1966 (with J.R. Napier) Catalogue of the skeletal material of Gorilla in the British Isles. J. Zool. 148:153-161. 1966 (with F. Ziccardi and A. Toschi). Sull'asino Selvatico Africano. Rich Zool. Appl. Caccia Suppl. 5(1):1-30. 1967 Ecology and Taxonomy of the Gorilla. Nature 213:890-3. 1967 The Nomenclature of the Eastern Lowland Gorilla. Nature 215:1172. 1967 The Rhinoceroses of Southeast Asia. Saugetierkundl. Mitt. 15:221-237. 1967 (with D.L. Harrison). The Taxonomy of the Gazelles of Arabia. J. Zool. 152:381-7. 1967 Geographic Variation in the Black Rhinoceros. Z.f. Saugetierk. 32:267-276. 1967 Geographic Variation in the Hoolock or White-browed Gibbon. Folia Primat. 7:276-283. 1967 (with V. Mazak). Taxonomic Problems of Asiatic Wild Asses; with the description of a new subspecies. Z.f. Saugetierk. 32:321-355. 1968 Simia leucophaea F. Cuvier, 1807 (Mammalia): Proposed validation under the plenary powers. Bull. Zool. Nomen. 25:36-7. 1968 The Classification of the Gibbons (Primates, Pongidae). Z.f. Saugetierk. 33:239-246. 1968 A New Subspecies of Gibbon from Northern Thailand. Proc. Biol. Soc. Wash. 81:625-8. 1969 Hylobates lar (Linneaus, 1771), H. entelloides (I. Geoffroy St. Hilaire, 1842) and H. Hoolock (Harlan, 1834), (Mammalia Pongidae): proposal to place these names on the official list of specific names in Zoology. Bull. Zool. Nomen. 25:162-4. 1969 (with J. R. Napier). Dentition and diet in Australopithecines. Proc. VIII Congr. Anthrop. Ethnol. Sci. Tokyo/Kyoto 3:273-6. 1969 The Taxonomy of the Anoa. Beaufortia (Amsterdam) 17:1-12. 1969 The Smaller Gazelles of the Genus Gazella de Blainville, 1816. Z.f. Saugetierk. 34:38-60. 1970 The Forgotten Leaf-eaters, and the Phylogeny of the Colobinae. In J.R. Napier and P.H. Napier (eds) The Old World Monkeys. Academic Press. 1970 (with R.W. Thorington). Annotated classification of the Cercopithecoidea. In Napier and Napier, op. cit. 1970 Taxonomic and Individual Variation in Gibbons. Symp. Zool. Soc. London, No. 26:127-134. 1970 Population Systematics of the Gorilla. J. Zool. 161:287-300. 1971 Request for a declaration modifying Article 1, so as to exclude names proposed for domestic animals from Zoological Nomenclature. Bull. Zool. Nomencl. 27:269-272. 1971 Pongo Pygmaeus. Mammalian Species No. 4:1-6. 1971 Distribution and Place of Origin of the Gorilla. Man, N.S. 6:44-51. 1971 Geographic and individual variation in Bornean Gibbons, with remarks on the systematics of the subgenus Hylobates. Folia Primat. 14:139-153. 1971 Species Characters in Rhinoceros Horns. Z.f. Saugetierk. 36:238-252. 1971 Systematics of the Genus Nycticebus. Proc. 3rd Int. Congr. Primat., Zurich 1970, 1:44-53. 1972 Ceratotherium Simum. Mammalian Species No. 8:1-6. 1972 (with J. Sabater Pi). The importance of higher Primates in the diet of the Fang of Rio Muni. Man, 7:239-243. 1972 Systematics and Phylogeny of Gibbons. In D. Rumbaugh (ed) Gibbon and Siamang, 1:1-89. Karger, Basel. 1972 (with F. Kurt) Dicerorhinus Sumatrensis. Mammalian Species No. 21:1-6. 1972 Phylogeny and Classification of Primates. In R.N. T-W-Fiennes (ed) Pathology of Simian Primates, 1:11-57. Karger, Basel. 1973 Notes on the Ecology and Behaviour of the Angola Colobus (Colobus angolensis) in N.E. Tanzania. Folia Primat. 20:12-26. 1973 (with N. K. Humphrey). Asymmetry in Gorilla Skulls: evidence of lateralised brain function? Nature 244:53-4.
  • 2. 2 1974 A note on the systematic position of the Muntjac (Artiodactyla, Cervidae). Z.f. Saugetierk. 39:369-372. 1974 (with P. Grubb). A New Duiker from Rwanda. Rev. Zool. Afr. 88:189-196. 1974 (with P. Andrews and J.F.M. Horne). The Tana Colobus and Mangabey. Oryx 12:565-575. 1974 New Evidence on the Evolution of the Apes and Man. Vest. Ustr. Ust. Geol. 49:53-6. 1974 Taxonomy and Phylogeny of Prosimians. In R.D. Martin, G.A. Doyle and A.C. Walker (eds) Prosimian Biology, pp. 449-473. Duckworth. 1975 Taxonomic notes on the White Rhinoceros Ceratotherium simum (Burchell, 1817). Saugetierkundl. Mitt. 23:200-212. 1975 (with P. Andrews and J.F.M. Horne). Ecology of the Tana River Floodplain, Kenya. J. E. Afr. N.H. Soc. Nat. Mus. 151:1-31. 1975 Notes on the Gazelles. 1. Gazella rufifrons and the Zoogeography of Central African Bovidae. Z.f. Saugetierk. 40:308-319. 1975 Comment on the proposed conservation of Pan Oken, 1816 and Panthera Oken, 1816. Bull.Zool.Nomen. 32:68. 1975 (with V. Mazak). An approach to the taxonomy of the Hominidae: Gracile Villafranchian Hominids of Africa. Casopis Pro Min. Geol. 20:225-247. 1976 (with P. Andrews). Gibbons and Brachiation. In D.Rumbaugh (ed) Gibbon and Siamang 4:167-218. Karger. 1976 The Apes. Dyn 3:29-36. 1976 River of Monkeys. Hemisphere, 20(9):28-32. 1976 The Origin of the Mammalian Fauna of Sulawesi (Celebes). Z.f. Saugetierkunde 41:201-216. 1976 (with H. Hemmer and P. Grubb). Notes on the Sand Cat, Felis margerita Loche, 1858. Z.f. Saugetierkunde 41:286-303. 1976 Taxonomy of Moschus (Mammalia: Artiodactyla) of the Indian region. J. Bombay N. H. Soc. 72:662-676. 1977 (with Alan G. Goodall). The Conservation of Eastern Gorillas. In H.S.H. Prince Rainier III of Monaco and Geoffrey H. Bourne (ed), Primate Conservation, pp. 599-637. Academic Press, New York. 1977 Hylobates lar (Linnaeus, 1771), H. entelloides (I. Geoffroy St. Hilaire, 1842) and H. Hoolock (Harlan, 1834), (Mammalia, Hylobatidae): proposal to place these names on the Official List of Specific Names in Zoology. Bull. Zool. Nomencl. 34:75-9. 1977 Sima syndactyla Raffles, 1821 (Mammalia: Hylobatidae); proposal to give this name precedence over Sima gibbon, C. Miller, 1779. Bull. Zool. Nomencl. 34:104-5. 1977 Comments on request for a declaration modifying Article I so as to exclude names proposed for domesticated animals from Zoological Nomenclature. Bull. Zool. Nomencl. 34:137-9. 1978 Phylogenetic and population systematics of the Mangabeys (Primates: Cercopithecoidea). Primates 19:1-34. 1978 (with L.C.Rookmaaker). The taxonomic status of the Dwarf Blue Sheep (Artiodactyla: Bovidae). Saugetierkundl. Mitt. 26:177-183. 1978 (with C. Smeenk). On the Type material of Cervus nippon Temminck, 1836; with a revision of Sika Deer from the main Japanese islands. Zool. Meded. Leiden 53:11-28. 1978 The extinct Cape Rhinoceros, Diceros bicornis bicornis (Linnaeus, 1758). Saugetierkundl. Mitt. 26:117-28. 1978 (with V. Mazak and P.H.J. van Bree). On a skin and skull of the Bali Tiger, and a list of preserved specimens of Panthera tigris balica (Schwarz, 1912). Z.f. Saugetierkunde 43:108-113. 1978 Comments on the application concerning Simia syndactyla, Raffles 1821 (Mammalia: Hylobatidae). Bull. Zool. Nomencl. 35:76. 1978 Reply to Van Gelder. Bull. Zool Nomencl. 36:9-10. 1979 (with Kenhelm W. Stott, Jr). Systematic relationships of Gorillas from Kahuzi, Tshiaberimu and Kayonza. Folia Primatol. 32:161-179. 1979 Comment on Rightmire, 'Implications of Border Cave skeletal remains for late Pleistocene human evolution.' Current Anthrop. 20:29. 1979 A View of Race. Hemisphere 23:36-8. 1979 What does it mean to be conscious? Behav. Brain. Sci. 1:575-6. 1980 Notes on the Systematics of Babyrousa (Artiodactyla, Suidae). Zoologische Mededelingen. Leiden 55:29-46. 1980 (with C. Guerin). Le Rhinoceros sondaicus annamiticus (Mammalia, Perissodactyla) d'indochine: distinction taxinomique et anatomique; relations phyletiques. Geobios 13:199-208. 1980 Speciation in Macaca: the view from Sulawesi. In Donald D. Lindburg (ed) The Macaques: studies in ecology, behaviour and evolution, pp. 84-124. Van Nostrand Reinhold Company, New York. 1980 The Chinese Mountain Cat (Felis bieti). Carnivore, 3:35-41. 1980 A further note on Moschus. J. Bombay N.H. Soc. 77:130-3. 1981 (with D.P. Willoughby). Studies on the taxonomy and phylogeny of the genus Equus. 1. Subgeneric classification of the recent species. Mammalia 45:321:354. 1981 Promise unfulfilled: a rejoinder to Peter Wilson. Canberra Anthrop. 4:101-9. 1981 Notes on the Gazelles. 2. Subspecies and clines in the Springbok (Antidorcas). Z.f.Saugetierkunde 46:189-197. 1981 Comment on Shipman et al., 'Butchering of Giant Geladas at an Acheulian Site', Current Anthrop. 22:265. 1981 Systematic relationships in the Bovini (Artiodactyla, Bovidae). Z.f. Zool. Syst.Evolutionsforschung 19:264-278.
  • 3. 3 1981 Notes on the Gazelles. 3. The Dorcas Gazelles of North Africa. Annali del Museo Civico di Storia Naturale di Genova 83:455-471. 1981 Comment on Johnson, 'Bonobos: generalised hominid prototypes or specialised insular dwarfs?' Current Anthrop. 22:366. 1981 Survey of the Tana river Colobus and Mangabey. National Geographic Society Research Reports 13:235-242. 1982 Asian Rhinoceroses - down but not out. Malayan Naturalist 36:11-22. 1982 (with P. Grubb). The species of Muntjac (genus Muntiacus) in Borneo: unrecognised sympatry in tropical deer. Zoologische Mededelingen, Leiden 56:203-216. 1982 (with P. R. Groves). The Lions of India. Hemisphere 26(4):206-209. 1982 The skulls of Asian Rhinoceroses: wild and captive. Zoo Biology 1:251-261. 1982 The Systematics of Tree Kangaroos (Dendrolagus; Marsupialia, Macropodidae). Aust. Mammal. 5:157-186. 1982 A note on geographic variation in the Indian Blackbuck (Antilope cervicapra Linnaeus, 1758). Rec. Zool. Surv. India 79:489-503. 1982 Bos gaurus H. Smith, 1827 (Mammalia, Artiodactyla): proposed conservation. Bull. Zool. Nom. 39:279-280. 1982 Antilope depressicornis H. Smith, 1827, and Anoa quarlesi Ouwens, 1910 (Mammalia, Artiodactyla): proposed conservation. bull. Zool. Nom. 39:281-2. 1983 Phylogeny of the living species of Rhinoceros. Z. Zool. Syst. u. Evolutionsforschung 21:293-313. 1983 Notes on the Gazelles. 4. The Arabian Gazelles collected by Hemprich and Ehrenberg. Z.f. Saugetierkunde 48:371-381. 1983 Geographic variation in the Barasingha or Swamp Deer (Cervus duvauceli). J. Bombay N.H. Soc. 79:620-629. 1983 (with P.H. Napier). Simia fascicularis Raffles, 1821 (Mammalia, Primates): Request for the suppression under the Plenary Powers of Simia aygula Linnaeus, 1758, senior synonym. Bull. Zool. Nom. 40:117-8. 1983 (with S. Chakraborty). The Calcutta collection of Asian Rhinoceros. Rec. Zool. Surv. India 80:251-263. 1983 (with P. Grubb). Notes on the Taxonomy of the Deer (Mammalia, Cervidae) of the Philippines. Zool. Anz. Jena 210:119-144. 1983 (with T. Flannery). The Mammals of West Irian. Hemisphere, 27:378-84. 1984 A new subspecies of Sable antelope, Hippotragus niger (Harris, 1838). Rev. Zool Afr. 97:821-8. 1984 (with P. M. Monappa). Elephants at Work. Hemisphere 28(5):285-291. 1984 Of Mice and Men and Pigs in the Indo-Australian Archipelago. Canberra Anthropology, 7:1-19. 1984 Cranial and dental characteristics in the systematics of Old World Felidae. Carnivore, 5 (for 1982):35-46. 1984 A new look at the taxonomy and phylogeny of the gibbons. In H.Preuschoft et al.(ed) The Lesser Apes 542-561. 1984 Pigs East of the Wallace Line. J. Société des Océanistes, 39:105-119. 1984 (with W.A. Laurie and E.M. Lang) Rhinoceros unicornis. Mammalian species No. 211:1-6. 1985 Mammal Faunas and the Palaeogeograhy of the Indo-Australian Region. Courier Forschungsinstitut Senckenberg No. 69 pp. 267-273. 1985 (with J. Sabater Pi). From Ape's Nest to Human Fix-point. Man 20:22-447. 1985 (with D.M. Lay). A new species of the genus Gazelles from the Arabian Peninsula. Mammalia 49:27-36. 1985 An Introduction to the Gazelles. Chinkara, 1, 1:4-10. 1985 (with V. Weitzel). The Nomenclature and Taxonomy of the Colobine Monkeys of Java. Int J.Primat.6:399-409. 1985 The Case of the Pygmy Gorilla: a cautionary tale for cryptozoology. Cryptozoology 4:37-44. 1985 Plio-Pleistocene Mammals in Island Southeast Asia. Mod. Quaternary Res. S.E. Asia 9:43-54. 1985 (with L.B. Holthuis). The Nomenclature of the Orang Utan. Zoo. Meded. Leiden 59:411-7. 1986 (with Feng Zuojian). The status of Musk Deer from Anhui Province, China. Acta Theriol. Sin 6:101-6. 1986 The Yahoo, the Yowie, and Reports of Australian Hairy Bipeds. Cryptozoology 5:47-54. 1986 Systematics of the Great Apes. Comparative Primate Biology 1:187-217. 1986 The taxonomy, distribution and adaptations of Recent Equids. In R.H. Meadow and H.P. Uerpmann (eds), Equids in the Ancient World, pp. 11-65. 1986 (with P.Grubb) Relationships of Living Deer. In C.Wemmer (ed), Biology and Management of the Cervidae, 21- 59. 1986 On the Highland Cuscuses of New Guinea currently referred to Phalanger vestitus. In M. Archer, (ed) Possums and Opossums: Studies in Evolution 559-567. 1986 On the Cuscuses of the Phalanger orientalis group from Indonesian Territory. In M. Archer, (ed) Possums and Opossums: Studies in Evolution 569-579. 1987 Monkey Business with Red Apes. J. Hum. Evol. 16:537-542. 1987 (with J.F. Eisenberg and K. MacKinnon). Tapire. In Grzimeks Enzyklopadie, 4:598-608. 1987 Obituary: John Russell Napier (1917-1987). J.Hum.Evol. 16:533-5. 1988 (with M. Lyons and A. Kovarch). A survey of beliefs on creation and evolution; or, did Julius Caesar Really Exist? The Skeptic 8, 4:19-24. 1988 (with J. McCarron-Benson, A. Meek and B. Puza). Creation Week in Canberra. The Skeptic 8, 4:6-9, 40. 1988 (with R.H. Eaglen). Systematics of the Lemuridae (Primates, Strepsirhini) J. Hum. Evol. 17:513-538. 1988 The Evolutionary Ecology of the Hominoidea. An.Psicol. 39:87-98.
  • 4. 4 1988 (with G.G. Scott & K.C. Richardson). Osteological differences of the skulls of Lasiorhinus latifrons Owen, 1845 and Vombatus ursinus Shaw, 1800 (Marsupialia: Vombatidae). Aust. J. Zool. 36:599-609. 1989 Feral Mammals of the Mediterranean Islands: documents of early domestication. pp. 46-58 in J. Clutton-Brock, (ed) The Walking Larder. London: Unwin Hyman. 1989 [Articles in Skeptical (ed. Laycock, Vernon, Groves and Brown): 'Introduction: Why be Skeptical?'; 'Aliens Among Us'; 'Channelling' (with W. Grey)' 'Clairvoyance'; 'Creationism'; 'Dolphins'; 'The Mystic East'; 'Near Death Experiences'; 'The New Age'; 'Noah's Ark'; 'Prophecy'; 'Racism'; 'Reincarnation' (with W. Grey); 'Shamans and Sorcerers'; 'Unknown Animals'; 'Velikovsky'; 'Von Daniken'.] 1989 (with G.G. Scott and K.C. Richardson). Skull morphometrics of Lasiorhinus latifrons (Owen, 1845) (Marsupialia: Vombatidae). Rec.S.Aust.Mus. 23:1-5. 1989 A catalogue of the genus Gazella. In A. Dixon & D. Jones eds., Conservation and Biology of Desert Antelopes, 193-8. 1989 Hominidae. Pp.910-916 in Fauna of Australia: Vol. 1B, Mammalia. (D.W. Alton & B.J. Richardson, eds.). AGPS, Canberra. Equidae. Pp.1039-1043 in ibid. (with *J. Giles). Suidae. Pp.1044-1049. (with *J.F. Bishop). Cervidae. Pp.1054-1060. Bovidae. Pp.1061-1066. 1989 (with T. Flannery, K. Aplin and M. Adams.) Revision of the New Guinean genus Mallomys (Muridae: Rodentia), with descriptions of two new species from subalpine habitats. Rec. Aust. Mus., 41:83-105. 1989 The origin of human species: what we have learned about human evolution in the 130 years since On the origin of species. The Medical Journal of Australia 151:677-688. 1989 Brief Communication: Weights of Wild Orang Utans. Amer.J.Phys.Anthrop. 81:1-3. 1989 Natural selection and intelligent ancestors. Mankind, 19:76-82. 1989 The gazelles of the Arabian peninsula. In A.H. Abu-Zinada, P.D. Goriup and I.A. Nader, eds., Wildlife Conservation and Development in Saudi Arabia, 237-48. 1990 (with P.Grubb). Muntiacidae. In G.A. Bubenik & A.B. Bubenik (eds) Horns, Pronghorns and Antlers, 132-68. 1990 (with Wang Yingxiang). The gibbons of the subgenus Nomascus (Primates, Mammalia). Zool.Res. 11:147-54. 1990 The centrifugal pattern of speciation in Meganesian rainforest mammals. Mem.Qd.Mus. 28:325-8. 1990 (with T. Flannery). Revision of the families and genera of bandicoots. In J.H. Seebeck, R.L. Wallis, P.R. Brown and C.M. Kemper, eds., Bandicoots and Bilbies, 1-11. 1990 Out of Africa - here we go again. In L. Freedman, ed., Is our Future limited by our Past?: Proceedings of the Third Conference of the Australasian Society for Human Biology, pp. 13-25. 1990 (with R. Markham). Weights of wild Orang Utans. Amer. J. Phys. Anthrop. 81:1-3. 1990 The taxonomy of buffaloes. Buffalo Bulletin, 9:65-8. 1990 "Primates" in Wheldon Owen's Encyclopaedia of Mammals, Wheldon Owen, Sydney. 1990 (with L. Christidis). A Method for Identifying Night Monkeys (Aotus) by Simple Karyotyping and External Appearance. Australian Primatology, 5:11-12. 1991 (with J.D. Paterson). Testing hominoid phylogeny with the PHYLIP program. J.Hum.Evol. 20:167-83. 1991 (with I. Tattersall). Geographical variation in the Fork-marked Lemur, Phaner furcifer (Primates, Cheirogaleidae). Folia primatol. 56:39-49. 1991 Genes, Genitals and Genius:The Evolutionary Ecology of Race. In P.O'Higgins, ed., Human Biology: An Integrative Science: Proc. Fourth Conference Australasian Society for Human Biology, pp. 419-432. 1992 How old are subspecies? A tiger's eye view of human evolution. Archaeology in Oceania, 27:153-160. 1992 Gorillas - where and what? Gorilla, 15,2:2,3. 1992 (with C. Westwood and B.T. Shea). Unfinished Business: Mahalanobis and a Clockwork Orang. Journal of Human Evolution, 22:327-340. 1993 Bad Medicine for Wildlife. The Skeptic, 13(1):12-14, 17. 1993 Catalogue of Primates in the Collection of the Museum of Victoria, Melbourne (with S. Bricknell). Australian Primatology 7(4):20-27. 1993 Catalogue of Primates in the Macleay Museum, Sydney University. Australian Primatology 7(3):8-9. 1993 (with D.W.Cameron). Pliocene Hominins: hunters and/or scavengers? Perspectives in Human Biology 3:23-31. 1993 (with M.Karami). A mammal species new for Iran: Gazella gazella Pallas, 1766 (Artiodactyla: Bovidae). J.Sci.I.R.Iran, 4:81-89. 1993 The Chinkara (Gazella bennetti) in Iran, with the description of two new subspecies. J.Sci.I.R.Iran 4:166-178. 1993 Speciation in Living Hominoid Primates. In W. H. Kimbel and L. B. Martin (eds.), Species, Species Concepts and Primate Evolution. New York: Plenum Press. 1993 Testing rhinoceros subspecies by multivariate analysis. Pp. 92-100 in O.A. Ryder (ed.) Rhinoceros Biology and Conservation. Zoological society of San Diego. 1993 Orders Monotremata, Dasyuromorphia, Peramelemorphia, Notoryctemorphia, Diprotodontia and Primates. Pp. 13-14, 29-62 and 243-278 in D.E. Wilson and D.A.M. Reeder (eds.), Mammal Species of the World: a taxonomic and geographic reference. Smithsonian Institution Press. 1993 (with A. Greth, D. Williamson, G. Schwede, M. Vassart). Bilkis gazelle in Yemen - status and taxonomic relationships. Oryx, 27:239-244.
  • 5. 5 1993 (with R. Angst and C. Westwood). The status of Colobus polykomos dollmani Schwarz. International Journal of Primatology, 14:573-586. 1993 Why do Langurs kill each others' babies? Australian Primatology, 7:2-7. 1993 Human origins; The Neanderthals; and Dating the Past. Pp. 33-52, 68 and 74-75, The Illustrated History of Humankind, Vol. 1, The First Humans. Harper San Francisco. 1994 The origin of modern humans. Interdisciplinary Science Reviews, 19:23-34. 1994 Morphology, habitat and taxonomy. Pp 39-59 in Lee Boyd & Katherine A.Houpt (eds.), Przewalski’s Horse: the History and Biology of an Enadangered Species. Albany, NY: SUNY Press. 1994 (with A.K.Kes Hillman-Smith). Diceros bicornis. Mammalian Species, no.455, 8pp. 1995 Domesticated and Commensal Mammals of Austronesia and their Histories. Pp. 152-163 in The Austronesians: Historical and Comparative Perspectives (P. Bellwood, J.J. Fox and D. Tryon, eds.). Canberra: Department of Anthropology, ANU. 1995 (with John W.H.Trueman). Lemurid systematics revisited. Journal of Human Evolution, 28:427-437. 1995 (with H.Mendelssohn and Y.Yom-Tov). Gazella gazella. Mammalian Species, no.490, 7pp. 1995 (with H.Mendelssohn and Y.Yom-Tov). Gazella dorcas. Mammalian Species, no.491, 6pp. 1995 On the nomenclature of domestic animals. Bull.Zool.Nomencl., 52:137-141. 1995 (with C.R.Westwood). Skulls of the Blaauwbok, Hippotragus leucophaeus. Z.f.Säugetierkunde, 60:314-8. 1995 (with Wang Yingxiang and Peter Grubb). Taxonomy of Musk-deer, genus Moschus (Moschidae, Mammalia). Acta Theriologica Sinica, 15:181-197. 1995 Microtaxonomy and its implications for captive breeding. Pp 24-28 in U.Ganslosser, J.K.Hodges & W.Kaumanns (eds.), Research and Captive Propagation. Fürth: Filander Verlag. 1996 (with J.Shoshani, E.L.Simons & G.F.Gunnell). Primate phylogeny: morphological vs molecular results. Molecular Phylogenetics and Evolution, 5:102-154. 1996 Taxonomic diversity in Arabian gazelles: the state of the art. Pp 8-39 in A.Greth, C.Magin & M.Ancrenaz (eds.), Conservation of Arabian Gazelles. National Commission for Wildlife Conservation and Development, Riyadh, Saudi Arabia. 1996 Australopithecus afarensis Johanson, 1978 (Mammalia, Primates): proposed conservation of the specific name. Bulletin of Zoological Nomenclature, 53:24-7. 1996 (with A.Gentry & J.Clutton-Brock). Proposed conservation of usage of 15 mammal specific names based on wild species which are antedated by or contemporary with those based on domestic anaimals. Bulletin of Zoological Nomenclature, 53:28-37. 1996 (with P.M.Goonan & R.D.Smith). Karyotype polymorphism in the Slender Loris (Loris tardigradus). Folia primatologica, 65:100-9. 1996 The nomenclature of the Tanzanian Mangabey and the Siberut Macaque. Australian Primatology, 10, 4:2-5. 1996 Great Apes: the conflict of gene-pools, conservation and personhood. Perspectives in Human Biology 2:31-36. 1996 Hovering on the brink: nearly but not quite getting to Australia. Perspectives in Human Biology, 2:83-7. 1996 From Ussher to Slusher; from Archbish to Gish; or, not in a million years... Archaeol. Oceania 31:145- 151. 1996 The taxonomy of the Asian Wild Buffalo from the Asian mainland. Zeitschrift für Säugetierkunde 61:327-338. 1997 Major biogeographic regions of the world. Section of article “Biosphere”. Pp 1160-1167 (vol.14) in The New Encyclopaedia Britannica: Macropaedia, 15th edition. 1997 (with G.B.Schaller, G.Amato & K.Khounbouline). Rediscovery of the wild pig Sus bucculentus. Nature 386:335. 1997 Taxonomy of wild pigs (Sus) of the Philippines. Zoological Journal of the Linnaean Society, 120:163-191. 1997 Taxonomy and phylogeny of Primates. In A.Blancher, J.Klein & W.W.Socha (eds.), Molecular Biology and Evolution of Blood Group and MHC Antigens in Primates. Berlin: Springer Verlag, Berlin Heidelberg and New York, pp.3-23. 1997 The taxonomy of Arabian Gazelles. In K.Habibi, A.H.Abuzinada & I.A.Nader (eds.), The Gazelles of Arabia, Riyadh: National Commission for Wildlife Conservation and Development, Publication No.29, Eng.Ser., 24-51. 1997 Thinking about evolutionary change: the polarity of our ancestors. In G.A.Clark and C.M.Willermet (eds), Conceptual Issues in Modern Human Origins. Aldine de Gruyter, N.Y., 1997, pp 319-326. 1997 Species concept in palaeoanthropology. Perspectives in Human Biology, 3:13-20. 1997 Leopard-cats, Prionailurus bengalensis (Carnivora: Felidae) from Indonesia and the Philippines, with the description of two new subspecies. Zeitschrift für Säugetierkunde, 62:330-338. 1998 The proximal ulna from Klasies River. Journal of Human Evolution, 34:119-121. 1998 Systematics of Tarsiers and Lorises. Primates, 39:13-27. 1998 (with M.Goodman, C.A.Porter, J.Czelusniak, S.L.Page, H.Schneider, J.Shoshani and G.Gunnell). Toward a phylogenetic classification of Primates based on DNA evidence complemented by fossil evidence. Molecular Phylogenetics and Evolution, 9:585-598. 1998 (with H.Mendelssohn and B.Shalmon). A new subspecies of Gazella gazella from the southern Negev. Isreal Journal of Zoology, 43:209-215. 1998 Obituary: Prudence Hero Napier (1916-1997). International Journal of Primatology, 19:203-205.
  • 6. 6 1999 Die Nashörner - Stammesgeschichte und Verwandtschaft. In Anonymous (ed.), Die Nashörner: Begegnung mit urzeitlichen Kolossen, 14-32. Furth: Filander Verlag. 1999 (with T.F.Flannery). A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies. Mammalia, 62:367-396. 1999 The advantages and disadvantages of being domesticated (A Keynote Address). Perspectives in Human Biology, 4:1-12. 1999 Australopithecus garhi: a new-found link? Reports of the National Center for Science Education, 19, 3:10-13. 1999 Nomenclature of African Plio-Pleistocene hominins. Journal of Human Evolution, 37:869-872. 1999 (with A.Thorne). The terminal Pleistocene and early Holocene populations of northern Africa. Homo, 50, 3:249-262. 2000 (with P.Grubb, J.P.Dudley & J.Shoshani). Living African elephants belong to two species: Loxodonta africana (Blumenbach, 1797) and Loxodonta cyclotis (Matschie, 1900). Elephant, 2, 4:1-4. 2000 (with P.Grubb), Do Loxodonta cyclotis and L.africana interbreed? Elephant, 2, 4:4-7. 2000 What are the elephants of West Africa? Elephant, 2, 4:7-8. 2000 (with P.Grubb). Are there Pygmy Elephants? Elephant, 2, 4:8-10. 2000 Phylogenetic relationships within recent Antilopini (Bovidae). In E.S.Vrba and G.B.Schaller (eds.), Antelopes, Deer, and Relatives, 223-233. New Haven & London: Yale University Press. 2000 (with G.B.Schaller). The phylogeny and biogeography of the newly discovered Annamite artiodactyls. In E.S.Vrba and G.B.Schaller (eds.), Antelopes, Deer, and Relatives, 261-282. New Haven & London: Yale University Press. 2000 Phylogeny of the Cercopithecoidea. In P.Whitehead and C.J.Jolly, eds., Old World Monkeys, 77-98. Cambridge: Cambridge University Press. 2000 (with O.A.Ryder). Systematics and phylogeny of the horse. In A.T.Bowling and A.Ruvinsky (eds.), The Genetics of the Horse, 1-24. Wallingford (Oxon., U.K.) & New York: CABI Publishing. 2000 (with A.Thorne). The affinities of the Klasies River Mouth remains. In J.S.Chisholm (ed.), Towards Consilience: Perspectives in Human Biology, 5:43-53. 2000 (with M.Chech, A.Thorne & E.Trinkaus). A new reconstruction of the Shanidar 5 cranium. Paléorient 25:143- 146. 2000 The genus Cheirogaleus: unrecognised biodiversity in Dwarf Lemurs. International Journal of Primatology, 21:943-962. 2000 La spéciation chez les primates. Primatologie, 3:213-236. 2001 Mammals in Sulawesi: where did they come from and when, and what happened to them when they got there? Pp 333-342 in I.Metcalfe, J.M.B.Smith, M.Morwood & I.Davidson (eds.), Faunal and Floral Migrations and Evolution in SE Asia-Australasia. Lisse/Abingdon/Exton (PA)/Tokyo: A.A.Balkema. 2001 Time and taxonomy. Ludus Vitalis, 9:91-96. 2001 Why taxonomic stability is a bad idea, or Why are there so few species of Primates (or are there?). Evolutionary Anthropology, 10:192-198. 2001 (with A.Braun, P.Grubb, Yang Q. & Xia L.). Catalogue of the Musée Heude collection of mammal skulls. Acta Zootaxonomia Sinica, 26:608-660. 2002 (with A.Meder). A model of gorilla life history. Australasian Primatology, 15:2-15. 2002 (with A.Braun & P.Grubb). Rediscovery of the type specimen of Bubalus mindorensis Heude, 1888. Mammalian Biology, 67:246-249. 2002 (with P.Grubb). Appendix C – Revision and classification of the Cephalophinae. Pp 703-728 in V.J.Wilson, Duikers of Africa: Masters of the African Forest Floor. Bulawayo: Chipangali Wildlife Trust. 2002 Taxonomy of living Equidae. Pp 94-107 in P.D.Moehlman (ed.), Equids: Zebras, Asses and Horses. Gland, Switzerland: IUCN. 2002 (with A.Kitchener). New insights into the taxonomy of Macaca pagensis of the Mentawai Islands, Sumatra. Mammalia, 66:533-542. 2003 The tarsiers of Sulawesi. Pp 179-195 in P.C.Wright, E.L.Simons & S.Gursky (eds.), Tarsiers: Past, Present, and Future. New Brunswick: Rutgers U.P. 2003. (with A.B.Taylor). Patterns of mandibular variation in Pan and Gorilla. Journal of Human Evolution, 44:529- 561. 2003 (with J.Harding). Morphology, morphometrics and taxonomy, in Field and Laboratory Methods in Primatology, Joanna M. Setchell and Deborah J. Curtis (eds), Cambridge University Press, Cambridge, pp 140-157. 2003 (with D.Bulbeck, D.Rayner and P.Raghavan). The Contribution of South Asia to the Peopling of Australasia and the relevance of Basel’s Naturhistorisch Museum anthropological collection to the project aims, Bulletin der Schweizerischen Gesellschaft fur Anthropologie / Bulletin de la Societe Suisse d’Anthropologie, 2: pp 49-70 2003 (with R.Stumpf, J.Polk, J.Oates, W.Jungers, C.Heesy and J.Fleagle). Patterns of Diversity in Gorilla Cranial Morphology, in Gorilla Biology: A Multidisciplinary perspective, Andrea Taylor and Michele Goldsmith (eds), Cambridge University Press, Cambridge, pp 35-61. 2003 A History of Gorilla Taxonomy, in Gorilla Biology: A Multidisciplinary perspective, Andrea Taylor and Michele Goldsmith (eds), Cambridge University Press, Cambridge, pp 15-33.
  • 7. 7 2003 Taxonomy of ungulates of the Indian subcontinent. Journal of Bombay natural History Society, 100:341-362. 2004 (with D.Cameron) Bones, Stones and Molecules: “Out of Africa” and Human Origins, pp 395. Elsevier Inc. Academic Press, Burlington, MA. 2004 Charles Oxnard and the aye-aye: morpometrics, cladistics, and two very special primates. In Shaping Primate Evolution: Form, Function, and Behavior, Fred Anapol, Rebecca Z. German, Nina G. Jablonski (eds), Cambridge University Press, Cambridge, UK, pp 368-377. 2004 Taxonomy and biogeography of primates in Vietnam and surrounding regions. In Conservation of Primates in Vietnam, Tilo Nadler, Ulrike Streicher and Ha Thang Long (eds), Haki Publishing, Hanoi, pp 15-20. 2004 (with H.Schulze) Asian Lorises, Taxonomic problems caused by Illegal Trade. In Conservation of Primates in Vietnam, Tilo Nadler, Ulrike Streicher and Ha Thang Long (eds), Haki Publishing, Hanoi, pp 33-36. 2004 (with C. Bell) New investigations on the taxonomy of the zebras genus Equus, subgenus Hippotigris. Mammalian Biology, 69: 182-196. 2004 (with A. Gentry & J. Clutton-Brock). The naming of wild animal species and their domestic derivatives. Journal of Archaeological Science, 31: 645-651. 2004 (with R.L.Pan). Cranial variation among the Asian colobines, In Shaping Primate Evolution: Form, Function, and Behavior, Fred Anapol, Rebecca Z. German, Nina G. Jablonski (eds) 1st edition, Cambridge University Press, Cambridge, UK, pp 45-65. 2004 (with Pan, R. and C.E.Oxnard). Relationships between the fossil Colobine Mesopithecus pentelicus and extant Cercopithecoids, based on dental metrics. American Journal of Primatology, 62:287-299. 2004 (with E.Meijaard) A taxonomic revision of the Tragulus mouse-deer (Artiodactyla). Zoological Journal of the Linnean Society, 140: 63-102. 2004 (with E.Meijaard). The Biogeographical Evolution and Phylogeny of the genus Presbytis. Primate Report, 68: 71-90. 2004 (with C.Pitra, J. Fickel & E.Meijaard). Evolution and phylogeny of old world deer. Molecular Phylogenetics and Evolution, 33: 880-895. 2004 (with E.Meijaard). Morphometrical relationships between South-east Asian deer (Cervidae, tribe Cervini): evolutionary and biogeographic implications. Journal of Zoology, London, 2004:179-196. 2004 (with T.Geissmann and C.Roos). The Tenasserim Lutung, Trachypithecus barbei (Blyth, 1847) (Primates: Cercopithecidae): Description of a live specimen, and a reassessment of phylogenetic affinities, taxonomic history, and distribution, Contributions to Zoology, 73: 271-282. 2004 The What, Why and How of Primate Taxonomy. International Journal of Primatology 25:1105-1126. 2005 The taxonomy of Primates in the laboratory context. In The Laboratory Primate, Sonia Wolfe-Coote, ed., Elsevier Academic Press, 3-15. 2005 Domestic and wild mammals: naming and identity. In Animal Names, A. Minelli, G. Ortalli & G. Sangha, eds, Venice, Istituto Veneto di Scienze, Lettere ed Arti, 151-157. 2005 Geographic variation within Eastern Chimpanzees (Pan troglodytes cf schweinfurthii Giglioli, 1872). Australasian Primatology, 17, 2: 19-46. 2005 (with E.Meijaard). Interspecific variation in Moschiola, the Indian Chevrotain. Raffles Bulletin of Zoology, 12:413-421. 2005 The genus Cervus in eastern Eurasia. European Journal of Wildlife Research, 52:14-22 and DOI 10.1007/s10344-005-0011-5. 2005 Orders Monotremata, Dasyuromorphia, Peramelemorphia, Notoryctemorphia, Diprotodontia and Primates in Mammal Species of the World: A Taxonomic and Geographic Reference. Don E Wilson & DeeAnn M. Reader (eds) 3rd edition, Johns Hopkins University Press, Baltimore, pp 1-2, 22, 23-37, 38-42, 43-70, 111-184. 2005 (with K.Helgen). Biodiversity in Sri Lanka and the Western Ghats. Science, 308:199. 2005 (with A.Mootnick). A New Generic Name for the Hoolock Gibbon (Hylobatidae). International Journal of Primatology, 26:971-975. 2006 (with A B. Rylands, R.A.Mittermeier, L. Cortes-Ortiz &J J.H Hines). Taxonomy and distributions of Mesoamerican Primates. In New Perspectives in the Study of Mesoamerican Primates, A.Estrada, P. A. Garber, M.S.M Pavelka & L. Luecke, eds, Springer, 29-79. 2006 (Articles) Biogeography; Cladistics; Fossey; Orangutan; Primate Taxonomy; Yeti. In Encyclopaedia of Anthropology, H. James Birx, ed., Sage Publications (Thousand Oaks, etc). 2006 (with E.Meijaard). The geography of mammals and rivers in mainland Southeast Asia. In Primate Biogeogaphy: Progress and Prospects, S.M.Lehman & J.G.Fleagle, eds, Springer, 305-329. 2006 (with S.Merker) Tarsius lariang: A New Primate Species from Western Central Sulawesi. International Journal of Primatology, 27: 465-485. 2006. Taxonomy and Biogeography of the Primates of Western Uganda. In Primates of Western Uganda, N.E.Newton- Fisher, H.Notman, J.D.Paterson & V.Reynolds, eds, Springer, 3-20. 2006. From Calpe to Catanduanes: Babewynes, apes, marmesettes and othere dyverse bestes. In The Barbary Macaque: Biology, Management and Conservation, J.K.Hodges & J.Cortes, eds., Nottingham University Press, 1-15.
  • 8. 8 2006 Humans among Primates. In J.R.Merrick, M.Archer, G.M.Hickey & M.S.Y.Lee (eds.), Evolution and Biogeography of Australasian Vertebrates. Oatlands, NSW: Australian Scientific Publishing. 2006 (with D. Argue, D. Donlon and R. Wright) Homo floresiensis: Microcephalic, pygmoid, Australopithecus, or Homo? Journal of Human Evolution, 51:360-374. 2006 (with J.H.Mazak). A taxonomic revision of the tigers (Panthera tigris) of Southeast Asia. Mammalian Biology 71:268-287. 2007 (with D.Zinner, C.Roos, J-L.Fauusser & Y.Rumpler). Disputed taxonomy classification of sportive lemurs (Lepilemur) in NW Madagascar. Lemur News, 12:53-56. 2007. The endemic Uganda Mangabey, Lophocebus ugandae, and other members of the albigena-group (Lophocebus). Primate Conservation 22: 123-128. 2007 The taxonomic diversity of the Colobinae of Africa. Journal of Anthropological Sciences, 85:7-34. 2007 (with K. M. Helgen). Craniodental characters in the taxonomy of Propithecus. International Journal of Primatology, 28: 1363-1383. 2007 Species concepts and speciation: facts and fantasies. In W. Henke & I.Tattersall (eds.), Handbook of Palaeoanthropology, three: 1861-1879. Berlin etc: Springer. 2007 (with C.Smeenk). The nomenclature of the African wild ass. Zool. Med. Leiden 81:121-135. 2007 Current views on the taxonomy and zoogeography of the genus Sus. In U.Albarella, K.Dobney, A.Ervynck & P.Rowley-Conwy (eds), Pigs and Humans: 10,000 Years of Interaction, 15-29. Oxford: Oxford University Press. 2007 (with S.Mukherjee). Geographic variation in jungle cat (Felis chaus Schreber, 1777) (Mammalia, Carnivora, Felidae) body size: is competition responsible? Biological Journal of the Linnean Society, 92:163–172. 2007 The Homo floresiensis controversy. Hayati Journal of Biosciences, 123-126. 2008 Walking with Hobbits. Australasian Science, March 2008: 16-19. 2008 Extended Family: Long Lost Cousins. A personal look at the history of primatology. Arlington, Virginia: Conservation International. 2008 (with M.Shekelle & D.Brandon Jones). Taxonomic history of the tarsiers, evidence for the origins of Buffon's tarsier, and the fate of Tarsius spectrum Pallas, 1778. In Primates of the Oriental Night (M Shekelle et al., eds.), LIPI Press, 1-12. 2008 (with M. Shekelle, S. Gursky, I.Neri-Arboleda & A.Nietsch). A method for multivariate analysis and classification of tarsier tail tuft.In Primates of the Oriental Night (M Shekelle et al., eds.), LIPI Press, 71-84. 2008 (with I.Maryanto). Craniometry of slow lorises (genus Nycticebus) of insular Southeast Asia.In Primates of the Oriental Night (M Shekelle et al., eds.), LIPI Press, 115-122. 2008 (with J.F.Oates & P.D.Jenkins). The type locality of Pan troglodytes vellerosus (Gray, 1862), and implications for the nomenclature of West African chimpanzees. 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