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Introduction to Scientific Nomenclature Hendrik Segers Royal Belgian Institute of Natural Sciences Hue University, Vietnam  January 2009
What nomenclature? A set of  mandatory   rules  and  voluntary   recommendations  that determine the structure and formation of names of organisms, for use in scientific communication.
Why nomenclature? “ The objects of the Code are  to promote  stability  and  universability  in the scientific names of animals and to ensure that the name of each taxon is unique and distinct.  All its provisions and recommendations are subservient to those ends and none restricts the freedom of taxonomic thought or actions”  (ICZN 1999:2) Fundamental aim of nomenclature: avoid a Tower of Babel Need for universal codes!
Nomenclature codes ,[object Object],[object Object],[object Object],[object Object]
The ICZN
History of zoological nomenclature ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Why do names change? Correct application of nomenclatural codes (e.g., correction of errors; homonyms;…) Increased scientific understanding   (e.g., discoveries; changes in species concept ( s.l. ) and phylogenetic understanding)
Why do names change? Correct application of nomenclatural codes (e.g., correction of errors; homonyms;…) Increased scientific understanding   (e.g., discoveries; changes in species concept ( s.l. ) and phylogenetic understanding) ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Ruling principles ,[object Object],[object Object],Tool  Science
Ruling principles ,[object Object],[object Object],[object Object]
Principle of Typification ,[object Object],[object Object],[object Object],[object Object],[object Object],=>  Identity of a name relies only on its  type , not on its description or diagnosis
Principle of Typification Types of name-bearing types  (in the species group) Original designation (Fixed in the original publication) Subsequent designation (Not fixed in the original publication) Holotype :  the single specimen upon which a new species-group taxon is based in the original publication (see also  isotype ) Paratypes :  remaining specimens of the original type series  (see also allotype) Syntypes :  specimens of a type series that collectively constitute the name-bearing type Hapantotype : (special case) Lectotype :  a syntype designated as the single-name bearing type specimen Paralectotypes :  each specimen of the former syntype series remaining after lectotype designation  (see also isolectotype) Neotype :  the single specimen designated as the name-bearing type when no name-bearing type specimen is believed to exist (anymore)
How does it work? Syntypes Type localities New Species A Species B
How does it work? Holotype + Paratypes Type locality New Species A Species B
How does it work? Lectotype + paralectotype Type localities Species A Species B
How does it work? Lectotype + paralectotype Type localities Species A Species B
Principle of Typification Terms not regulated by the Codes Allotype :  a designated specimen of opposite sex of the holotype Cotype :  a term formerly used for either syntype or paratype Genotype :  a term formerly used to designate the type species of a genus (generotype) Topotype : a specimen originating from the type locality or localities of the species or subspecies to which it is thought to belong, whether or not the specimen is part of the type series (see also  topotypical  specimen) Some peculiar cases:  Cleptotype ,  Iconotype , … Type of a family name = genus  Type of a genus = species
Ruling principles ,[object Object],[object Object],[object Object],[object Object]
Synonyms … ,[object Object],[object Object],[object Object]
Synonyms … ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Ruling principles ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
…  and   Homonyms ,[object Object],[object Object],[object Object],Problem: name for  L. ornata  (Harring & Myers, 1926) non (Daday, 1897)?
Ruling principles ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Synonyms … « Oldest fool »: Lecane ludwigii  (Eckstein, 1883) ,[object Object],[object Object],[object Object]
…  and   Homonyms Argus  Bohadsch, 1761(gastropod);  Argus  Scopoli, 1763 (butterfly);  Argus  Scopoli, 1777 (butterfly);  Argus  Poli, 1791 (mollusk);  Argus  Temminck, 1807 (bird);   Argus  Lamarck, 1817 (hesperid);  Argus  Boisduval, 1832 (lycaenid);  Argus  Walckenaer, 1836 (arachnid);  Argus  Gray, 1847(mollusk);  Argus  Gerhard, 1850 (lycaenid) ) Principle of Priority: Argus  Bohadsch, 1761  is the  only valid name
…  and   Homonyms ,[object Object],[object Object],[object Object],Problem: name for  L. ornata  (Harring & Myers, 1926) non (Daday, 1897)? No “Oldest fool” available!  Nomen novum  required:  Lecane myersi
Principle of Priority ,[object Object],The Commission can intervene ( nomina rejicienda, conservanda )
Ruling principles ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Monostyla closterocerca  Schmarda, 1853 What’s in a name? With subgenus name: Lecane (Monostyla) closterocerca  (Schmarda, 1853) Edmondson, 1935 Short :  Lecane (M.) closterocerca  (Shmarda, 1853)  Or:   L. closterocerca   new combination:   Lecane closterocerca  (Schmarda, 1853) Edmondson, 1935 With  subspecies  name: trinomen Lecane (Monostyla) closterocerca   amazonica  Koste, 1972  Or:   L. closterocerca amazonica
Scientific Names are Latin ,[object Object],[object Object],[object Object],[object Object],[object Object]
Formation of Scientific Names ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Formation of Scientific Names ,[object Object],[object Object],[object Object],[object Object],[object Object]
Formation of Scientific Names ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Formation of Scientific Names ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Formation of Scientific Names ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Formal requirements in ICZN (1) ,[object Object]
Formal requirements in ICZN (1) ,[object Object],[object Object]
Formal requirements in ICZN (1) ,[object Object],[object Object],[object Object],[object Object]
Some examples: One-letter difference… Eucosma  fandana Eucosma  gandana  Eucosma  handana  Eucosma  landana Eucosma  mandana  Eucosm a  nandana  Eucosma  pandana Eucosma  sandana   Eucosma wandana But also: Cydi a  candana  Epiblema  tandana Epinotia  xandana  Epinotia  zandana Pelochrista  randana Pelochrista  vandana  Tortricidae  (Moths, Northern Mexico : ( Kearfo t t, 1907)
[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],More examples….
[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],Even more examples….
Formal requirements in ICZN (2) ,[object Object],[object Object],[object Object],[object Object],[object Object]
The different codes: a comparison ,[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],The different codes: a comparison ,[object Object],[object Object]
Inter-Code Generic homonyms     Botanical genus names homonyms in Zool. record     Total   64,419 8,784  (13.6%)     in common use 28,041 3,554  (12.7%)       Bacteriological genus names Homonyms in Zool.  Record Homonyms in ING (Botanical) Homonyms in both Total 739 50 (6.8%) 29 (3.9%) 15 in common use   701 48 (6.9%) 27 (3.9%) 14
[object Object],[object Object],[object Object],Examples of genus + species homonoms:
[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],The different codes: a comparison
[object Object],[object Object],[object Object],[object Object],The different codes: a comparison
[object Object],[object Object],[object Object],[object Object],The different codes: a comparison
[object Object],[object Object],[object Object],[object Object],The different codes: a comparison
Table 2: Standardized endings for names of taxa 1 : for fungi; ²: for algae   Rank :   Zoological Botanical Bacteriological phylum/divisio * - phyta/-mycota 1   classis * - opsida/-mycetes 1 / -phyceae²   subclassis * - idea/-mycetidae 1 /-phycidae 2   superordo * - anae   ordo * - ales - ales subordo * - ineae - ineae superfamily - oidea  (not used) (not used) family - ida e - aceae - aceae subfamily - inae - oideae - oideae tribus - ini - eae - eae subtribus - ina - inae - inae
Interesting websites ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],Did I mention these ones?
Thank you for your attention

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Segers Introduction To Scientific Nomenclature

  • 1. Introduction to Scientific Nomenclature Hendrik Segers Royal Belgian Institute of Natural Sciences Hue University, Vietnam January 2009
  • 2. What nomenclature? A set of mandatory rules and voluntary recommendations that determine the structure and formation of names of organisms, for use in scientific communication.
  • 3. Why nomenclature? “ The objects of the Code are to promote stability and universability in the scientific names of animals and to ensure that the name of each taxon is unique and distinct. All its provisions and recommendations are subservient to those ends and none restricts the freedom of taxonomic thought or actions” (ICZN 1999:2) Fundamental aim of nomenclature: avoid a Tower of Babel Need for universal codes!
  • 4.
  • 6.
  • 7. Why do names change? Correct application of nomenclatural codes (e.g., correction of errors; homonyms;…) Increased scientific understanding (e.g., discoveries; changes in species concept ( s.l. ) and phylogenetic understanding)
  • 8.
  • 9.
  • 10.
  • 11.
  • 12. Principle of Typification Types of name-bearing types (in the species group) Original designation (Fixed in the original publication) Subsequent designation (Not fixed in the original publication) Holotype : the single specimen upon which a new species-group taxon is based in the original publication (see also isotype ) Paratypes : remaining specimens of the original type series (see also allotype) Syntypes : specimens of a type series that collectively constitute the name-bearing type Hapantotype : (special case) Lectotype : a syntype designated as the single-name bearing type specimen Paralectotypes : each specimen of the former syntype series remaining after lectotype designation (see also isolectotype) Neotype : the single specimen designated as the name-bearing type when no name-bearing type specimen is believed to exist (anymore)
  • 13. How does it work? Syntypes Type localities New Species A Species B
  • 14. How does it work? Holotype + Paratypes Type locality New Species A Species B
  • 15. How does it work? Lectotype + paralectotype Type localities Species A Species B
  • 16. How does it work? Lectotype + paralectotype Type localities Species A Species B
  • 17. Principle of Typification Terms not regulated by the Codes Allotype : a designated specimen of opposite sex of the holotype Cotype : a term formerly used for either syntype or paratype Genotype : a term formerly used to designate the type species of a genus (generotype) Topotype : a specimen originating from the type locality or localities of the species or subspecies to which it is thought to belong, whether or not the specimen is part of the type series (see also topotypical specimen) Some peculiar cases: Cleptotype , Iconotype , … Type of a family name = genus Type of a genus = species
  • 18.
  • 19.
  • 20.
  • 21.
  • 22.
  • 23.
  • 24.
  • 25. … and Homonyms Argus Bohadsch, 1761(gastropod); Argus Scopoli, 1763 (butterfly); Argus Scopoli, 1777 (butterfly); Argus Poli, 1791 (mollusk); Argus Temminck, 1807 (bird); Argus Lamarck, 1817 (hesperid); Argus Boisduval, 1832 (lycaenid); Argus Walckenaer, 1836 (arachnid); Argus Gray, 1847(mollusk); Argus Gerhard, 1850 (lycaenid) ) Principle of Priority: Argus Bohadsch, 1761 is the only valid name
  • 26.
  • 27.
  • 28.
  • 29. Monostyla closterocerca Schmarda, 1853 What’s in a name? With subgenus name: Lecane (Monostyla) closterocerca (Schmarda, 1853) Edmondson, 1935 Short : Lecane (M.) closterocerca (Shmarda, 1853) Or: L. closterocerca new combination: Lecane closterocerca (Schmarda, 1853) Edmondson, 1935 With subspecies name: trinomen Lecane (Monostyla) closterocerca amazonica Koste, 1972 Or: L. closterocerca amazonica
  • 30.
  • 31.
  • 32.
  • 33.
  • 34.
  • 35.
  • 36.
  • 37.
  • 38.
  • 39. Some examples: One-letter difference… Eucosma fandana Eucosma gandana Eucosma handana Eucosma landana Eucosma mandana Eucosm a nandana Eucosma pandana Eucosma sandana Eucosma wandana But also: Cydi a candana Epiblema tandana Epinotia xandana Epinotia zandana Pelochrista randana Pelochrista vandana Tortricidae (Moths, Northern Mexico : ( Kearfo t t, 1907)
  • 40.
  • 41.
  • 42.
  • 43.
  • 44.
  • 45. Inter-Code Generic homonyms   Botanical genus names homonyms in Zool. record     Total 64,419 8,784 (13.6%)     in common use 28,041 3,554 (12.7%)       Bacteriological genus names Homonyms in Zool. Record Homonyms in ING (Botanical) Homonyms in both Total 739 50 (6.8%) 29 (3.9%) 15 in common use 701 48 (6.9%) 27 (3.9%) 14
  • 46.
  • 47.
  • 48.
  • 49.
  • 50.
  • 51. Table 2: Standardized endings for names of taxa 1 : for fungi; ²: for algae Rank :   Zoological Botanical Bacteriological phylum/divisio * - phyta/-mycota 1   classis * - opsida/-mycetes 1 / -phyceae²   subclassis * - idea/-mycetidae 1 /-phycidae 2   superordo * - anae   ordo * - ales - ales subordo * - ineae - ineae superfamily - oidea (not used) (not used) family - ida e - aceae - aceae subfamily - inae - oideae - oideae tribus - ini - eae - eae subtribus - ina - inae - inae
  • 52.
  • 53.
  • 54. Thank you for your attention