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Maize Growth in East and South-east
Asia – Challenges and Opportunities
    Zhang Shihuang, Qiu Huanguang, Lu Kaiyv,
      Ci Xiaoke, Li Mingshun, Zhang Degui,
            Liang Xiaoling, Xu Jiashun

      National Maize Industrial Technology RD Center, MOA
     Chinese Academy of Agricultural Sciences, Beijing, China
                  cshzhang2000@yahoo.com.cn

              Chinese Academy of Agricultural
in Asia

We Are Facing the Same
      Challenges
   demand and imports
       production
        breeding
      seed industry


    Chinese Academy of Agricultural
Maize production in East and SE
             Asia




        Chinese Academy of Agricultural
Maize production in East and SE Asia (1961-2009)


                              180
                                                                                           Maize production
                                                                                           in Asia has been
Maize out put (million ton)




                              160                                y = 3.0 x + 6
                                                                   R2 = 0.96
                              140
                                                                                           accelerating
                              120

                              100
                                                                                           during the past 2
                              80
                                                                        y = 3.0 x + 3.9
                                                                           R2 = 0.96
                                                                                           decades due to
                              60
                                                                         y = 0.54x + 1.4
                                                                                           the increasing
                              40                                            R2 = 0.91
                                                                                           and diversifying
                              20

                               0
                                                                                           demands for
                                                                                           maize
                                 61

                                 64

                                 67

                                 70

                                 73

                                 76

                                 79

                                 82

                                 85

                                 88

                                 91

                                 94

                                 97

                                 00

                                 03

                                 06

                                 09
                              19

                              19

                              19

                              19

                              19

                              19

                              19

                              19

                              19

                              19

                              19

                              19

                              19

                              20

                              20

                              20

                              20
                                    Eastern Asia   South-Eastern Asia       China
Maize Production in East and SE Asia


                                other 31.58
                                   13%




                SE Asia 36.83
                    16%

                                                E Asia
                                              165.89 71%
      Asia
      29%




71%
Maize production (million ton) in East and SE Asia (2009)

                  Total maize output in Asia reached 234 million tons in 2009
                  accounted for one-fourth of the world total production

                                 900.0   818.82
                                 800.0                                               East Asia is the
Maize production (million ton)




                                 700.0                                               main producer of
                                 600.0                                               maize in Asia
                                 500.0
                                 400.0                                               166 million tons
                                 300.0            234.30                             in 2009
                                 200.0
                                                           165.89           164.11
                                 100.0                              36.83            20.2% of the
                                   0.0                                               world’s total
                                         World     Asia Eastern South- China         maize production
                                                         Asia Eastern
                                                                 Asia
Maize production (million ton) in different areas


                                                                                               •Maize production in
                             900.00


                             800.00

                                                                                               East Asia comes
Maize output (million ton)




                             700.00


                             600.00                                                            mainly from China
                             500.00


                             400.00
                                                                                               •accounted for 99%
                             300.00                                                            of East Asia’s total
                             200.00
                                                                                               out put and 20% of
                             100.00
                                                                                               the world output in
                               0.00
                                                                         South-
                                      World     Asia    Eastern Asia                  China

                               1961   205.03   31.60       19.40
                                                                       Eastern Asia
                                                                          4.71        18.03
                                                                                               2009
                               1990   483.34   132.47     101.33          16.44       97.21
                               2009   818.82   234.30     165.89          36.83       164.11
Maize Production in East and SE Asia


• Indonesia, the Philippines, Thailand,
  and Vietnam are some other main
  production countries of maize in
  Southeast Asia
Increment of maize production in the world, East and
SE Asia during 1961-2009 and 1990-2009

                800         755
                                        World
                                                              the average
                                  682
                                                              growth rate of
                700                     Eastern Asia
                                        South-Eastern Asia
Increment (%)




                600

                500                                           maize
                400
                      299
                                                              production in
                                                              East and SE
                300

                200
                                                              Asia is faster
                                                        124
                100                             69 64

                 0                                            than that of the
                      1961-2009                 1990-2009
                                                              world average
Maize Production in East and SE Asia


• 1961~2009
• maize production and area increased by
  2.99 times and 50% in the world
• but by 7.55 times and 101% in East Asia
• and by 6.82 times and 88% in SE Asia
Maize Production in East and SE Asia


• The growth rates of both yield and area have
  been slowing down in the last 2 decades
• maize production increased by 69% in the
  world, but increased by 64% and 124% in East
  and SE Asia
• Maize area increased 21% in the world, but
  43% and 7% in East Asia and SE Asia
Maize yield in East and SE Asia (1961-2009)


                        6.0
                                                             y = 0.03x + 4.58
                        5.0                                                               The trend of
Maize yield ( Ton/ha)




                        4.0
                                                                                          maize yield
                                                                  y = 0.04x + 4.59
                              y=0.11x+1.0                                                 change in
                        3.0
                                                  y=0.11x+0.974                           East Asia was
                        2.0                                             y = 0.1x + 1.64
                                                                                          contrary to
                        1.0
                                             y=0.03x+0.790
                                                                                          that in SE
                        0.0                                                               Asia
                              Eastern Asia         South-Eastern Asia           China
Maize Yield in East and SE Asia


• Maize yield increased faster before 1990,
  and have been slowing down since then
  in East Asia (esp in China)
• However, it has increased faster in SE
  Asia after 1990
• So the gap between East and SE Asia
  has been being narrower after 1990
Maize Yield in East and SE Asia


• Maize yield increment was due to the:
  – adoption of hybrids
  – increase of inputs
Maize Yield in East Asia


• In East Asia (especially in China)
• maize yield increased by 220% with an
  annual gain of 104 kg/ha during 1969-
  1989
• but increased by 15% with an annual
  gain of 35 kg/ha during 1990-2009
Maize Yield in SE Asia


• In SE Asia
• Maize yield increased by 99% with an
  annual gain of 31 kg/ha before 1990s
• but after that, increased by 110% with
  an annual gain of 99 kg/ha
Maize Yield in East and SE Asia


• Both the growth rate and the absolute
 growth of maize yields in decades
 showed a clear difference between East
 and SE Asia
Yield gains of maize decreasing in China

                                                     SE Asia
                                                                    4.4%
                                                            3.8%    2000
    East Asia                                       2.7%
                                                             1990
                                             2.3%
             5.9%                                    1980
                                    1.9%
           1960                              1970
                      4.5%
                                      1960
                     1970
                               3.2%
                               1980
                                           0.2%
                                         1990       1.7%
                                                    2000
(Data from FAO)

                    Chinese Academy of Agricultural Sciences
Maize production in China




     Chinese Academy of Agricultural
kg/ha          Maize yields since 1950s


                  Single-cross hybrids
                                                        b=23
                          b=126                    Stress tolerance

        OPVs
        b=14
                   b=89
           Double-cross hybrids


                                Year
                 Chinese Academy of Agricultural
Maize Production in China


• Maize yield in China increased by 3.4
 times, with an annual gain of 93 kg/ha
 during the past 5 decades
Maize Production in China(略)

• It should be noted that maize yield in China
  increased at 104 kg/ha/year before 1990s,
  which was even higher than that in Argentina
• but after 1990, maize yield increased at 38
  kg/ha/year which was much lower than the
  growth of 152 kg/ha/year in US and 180
  kg/ha/year in Argentina
• it was even lower than the gains in Vietnam,
  Myanmar, Indonesia and Brazil (more than
  100 kg/ha/year)
Maize yields in US, Argentina and China
12



10
                                          y=0.15x+7.0

 8

     y=0.11x+4.0                            y=0.18x+3.8
 6



 4
       y=0.7x+1.6                                  y=0.04x+4.6

 2
                        y=0.11x+1.0

 0



              America         Argentina      China
Maize Production in SE Asia

• On the contrary, maize yields have
  increased sharply since the mid-1990s in
  several SE Asian countries
• the annual gains of maize yield in
  Indonesia, Vietnam and Myanmar are
  4.1%,4.6% and 6.7%
• and annual gains of 107 kg/ha,143
  kg/ha and 110 kg/ha
Maize Trade in the Region
Maize Trade in the Region

• Over the past half-century, the world
  maize trade has increased significantly
• East Asia has been importing maize for
  a long time, and the imports continue to
  increase
• but some of the SE Asian countries
  turned into maize exporters in the mid-
  1980s
East Asia is a Net Importer (2009)

                                      2008




                                                                                 54970-
                                                  other part of the word




                                                                               9021-
                                                                 Korea




                                                                                16460-
                                                                 Japan




                                                                               220
                                                                  China



-60000     -50000   -40000   -30000      -20000         -10000             0              10000

                                      2008
East Asia is a Net Importer (2009)

                      South-east
                        Asia
                         7%

              13%


                                   Japan
                                    52%



        South Korea
            28%
Maize Trade in the Region (略)

• Japan, Korea and China were the main
  importers
• Maize imports in Japan (16-17 million tons
  annually) accounted for more than half of
  Asia’s total imports
• Korea (8-9 million tons) accounted for 28% of
  Asia’s total imports
Maize Trade in the Region

• Imports of Japan and South Korea have
  been relatively stable since 1990s
• China was a net export country in most
  years
• but its export has been steadily
  declining in the last decade
• China has become a net importer of
  maize in 2010 (1.5 million tons)
Conclusion--1

• Asia is a big importer of maize
• Demand for maize will continue to
  increase in Asia
Increase the Maize Production

    to meet tremendous challenges
Soil Improvement

Small tractors for more
than 20 years and made
the soil hardpan




                          top fertility soil is
                          16.5 cm only
                                       Prof. Zhang Dongxing
Improve the Soil

   soil deep loosing tillage
   to use subsoiler to break soil
    hardpan
   to increase top fertility soil
   to return farm residue and crop
    straw to the soil
Population density and machine

   increase the population density in
    maize production
         45,000/ha   60,000/ha
   farming machine
   new hybrids with stress tolerance
    and suitable to harvest machine
   irrigation works
   farmer collectives
Maize Yield and Population Density
        t/ha        bu/ac    plants/ha plants/ac
                1       16      10,000     4,047
                2       32      20,000     8,094
                3       48      30,000    12,141
                4       64      40,000    16,188
                5       80      45,000    18,211
                6       96      50,000    20,235
                7      112      55,000    22,258
                8      128      60,000    24,282
                9      144      65,000    26,305
               10      160      70,000    28,329
               11      176      75,000    30,352
               12      192      80,000    32,376
               13      208      85,000    34,399
               14      224      90,000    36,423
                                 Lamkey, 2007, Beijing   36
Maize Breeding Strategies in
     East and SE Asia

          Stress tolerance
  Suitable to harvest with machine
Maize Breeding in China

• The rate of maize yield gains in China
  has been reducing although the yield
  potential has been increasing since
  1960 when single cross hybrids were
  introduced
What is Yield?


• What is yield? -- questioned by Duvick
• Yield is stress tolerance !
• Stress tolerance of hybrids has been
 ignored for almost three decades due to
 the central planning economic
 constitution of RD management
Risky variety

Breeders tend to
develop hybrids
with tall stature, too
much leafy, late
maturity or full
season, and big
ears
most of this kind of
hybrids are risky
under stress
                   40
Plant and ear height (Beijing and Wulumuqi,
                                                       2007-2008)

                    280                                                      130
                                   y = 3.71x + 249.5
                    270                R2 = 0.50
                                                                             120
Plant height (cm)




                                                            Ear height(cm)
                    260
                                                                             110                    y = 1.15x + 105.4
                    250
                                                                                                        R2 = 0.10
                    240                                                      100
                          1950 1960 1970 1980 1990 2000                            1950 1960 1970 1980 1990 2000
                              Year of release                                             Year of release
                      Linear regression suggested that plant and ear height increased.
Maturity of hybrids delaied
                 72.0
                           Low
                 71.0      Mid
                           High
Maturity (day)



                 70.0

                 69.0

                 68.0

                 67.0

                 66.0
                    1960   1970   1980   1990   2000   2010
                                  Era (Year)
Shelling percentage(2008)
                85.0
 Shelling (%)




                80.0

                                                  y = -0.19x + 82.2
                                                       R2 = 0.03

                75.0
                       1950   1960     1970     1980        1990      2000
                              Year of release
Shelling percentage showed a trend to reduction during
1950s-2000s, especially reduced greatly during 1980s-2000s.
1.40
                    1.20           y = 0.0889x + 0.6097
 Barrent tip (cm)


                    1.00
                    0.80
                    0.60
                    0.40
                    0.20
                    0.00
                           1950s   1970s   1980s   1990s   2000s
秃尖长度                                       Year                    44
45
Pioneer hybrids 2006
Pioneer hybrids 2006 46
Stine Seeds, Iowa


150,000/ha

Sept. 2011


have to change
row spacing
from 76 cm to 50
cm, to increase
the population
density
redesigning the
farming machine


             47
Risky variety

• Breeders also tend to develop hybrids
  with excessive stay green and late
  maturity during the past 4 decades
• caused application of chemical
  fertilizers in a non-judicious manner
• reducing the efficiency of N utilization
Heterosis

• The contribution of heterosis to yield
  gains is higher in China than that
  reported in American hybrids
• the contribution of heterosis to yield gains showed
  weak increase with the year of hybrids release
Heterosis incerased (1970-2000)
                             90
                                     Het(%)
                             80
                             70
    Relative heterosis (%)




                             60
                                    66%                                69%
                             50
                             40           y = 0.09x - 107
                                             R2 = 0.02
                             30
                             20
                             10
                             0
                             1960     1970     1980     1990    2000         2010

                                              Year of release
(11 environments, 2007-2009)

                                     Chinese Academy of Agricultural
Schnell, F.W. 1974. Trends and problems in breeding methods for hybrid corn. p. 86–98.
In Proc. of the British Poultry Breeders Roundtable, 16th. Birmingham, England.          51
Duvick, D.N. 1999. Heterosis: Feeding people and protecting natural resources. p. 19–29. 52 J.G.
                                                                                         In
Coors and S. Pandey (ed.) The genetics and exploitation of heterosis in crops. ASA, Madison, WI.
Campbell, B.T., D.T. Bowman, and D.B. Weaver. 2008. Heterotic eff ects in topcrosses
of modern and obsolete cotton cultivars. Crop Sci. 48:593–605.              53
Reorienting the Strategy


• Maize breeders both in China and SE
 Asian countries are now actively
 reorienting the strategy of maize
 breeding, and develop stress resilient
 and input responsive hybrids
Reorienting the Strategy

• increase yield potential
• enhance stress tolerance
• increase population density in both
  crop production and breeding nursery
Strategies for breeding in Asia--1

• continue to introduce, improve and
  utilize exotic germplasm to broaden the
  genetic base in maize breeding
  programs
• to enhance genetic variance in breeding
  populations
• strengthen the pre-breeding efforts
Strategy --2

• germplasm should be divided into 2
  heterotic groups or 2 alignments
• based on the current genetic
  backgrounds, which will keep the SCA
  effects in two groups
Duvick, 2005



NSS




      SS




               58
Strategy --3

• improve the germplasm within each
  group and push them to the opposite
  directions in terms of gene frequency
• which will enhance and accumulate
  GCA effects in breeding populations of
  germplasm
Strategy -- 4

• improve germplasm and select inbred
  lines under high population density and
  other stress conditions based on G×E
  effects to enhance stress tolerance
• double density for inbred line
  development
• plus 15,000/ha for evaluation of new
  hybrids (Troyer,1981)
Strategy -- 5

• multi-location testing and information
  management
• science and art
• data and experience
• modern machine and IP technology
• pipeline and profolio
Conclusion--2

• public
• private
• pre-breeding
• pipeline
• profolio
Conclusion--3

• improve the soil and infrastruture
• improve hybrids to meet challenges
• stress tolerance is more important than
  heterosis for hybrid yields
• GCA is more important than SCA
• reorient the strategy of maize breeding
  programs
Chinese Academy of Agricultural
Demand for maize in China - 2020

• 230 million tons by 2020

• Processing 58 mt (26%)

• Feeding stuff 151 mt (70%)




         Chinese Academy of Agricultural
Three concepts of yield


• Genetic yield (maximum) —The highest yield
  record
• Attainable yield —average yield in Maize
  Variety Regional Evaluation Trials
• Actual yield — statistic data issued by MOA
The differences between yields and yield potential
                              great




         Genetic yieldyield
            Genetic yield
            Genetic yield
             Genetic           Attainable yield
                                Attainable yield
                                 Attainable yield      Actual yield
                                                     Actual yield
                                                     Actual yield
                                 Attainable yield
                                 Attainable yield
                                  Attainable yield   Actual yield
              227/120/127                860/95/30              Dr. RF Hu

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S(3) Maize Growth in East and South-east Asia – Challenges and Opportunities

  • 1. Maize Growth in East and South-east Asia – Challenges and Opportunities Zhang Shihuang, Qiu Huanguang, Lu Kaiyv, Ci Xiaoke, Li Mingshun, Zhang Degui, Liang Xiaoling, Xu Jiashun National Maize Industrial Technology RD Center, MOA Chinese Academy of Agricultural Sciences, Beijing, China cshzhang2000@yahoo.com.cn Chinese Academy of Agricultural
  • 2. in Asia We Are Facing the Same Challenges demand and imports production breeding seed industry Chinese Academy of Agricultural
  • 3. Maize production in East and SE Asia Chinese Academy of Agricultural
  • 4. Maize production in East and SE Asia (1961-2009) 180 Maize production in Asia has been Maize out put (million ton) 160 y = 3.0 x + 6 R2 = 0.96 140 accelerating 120 100 during the past 2 80 y = 3.0 x + 3.9 R2 = 0.96 decades due to 60 y = 0.54x + 1.4 the increasing 40 R2 = 0.91 and diversifying 20 0 demands for maize 61 64 67 70 73 76 79 82 85 88 91 94 97 00 03 06 09 19 19 19 19 19 19 19 19 19 19 19 19 19 20 20 20 20 Eastern Asia South-Eastern Asia China
  • 5. Maize Production in East and SE Asia other 31.58 13% SE Asia 36.83 16% E Asia 165.89 71% Asia 29% 71%
  • 6. Maize production (million ton) in East and SE Asia (2009) Total maize output in Asia reached 234 million tons in 2009 accounted for one-fourth of the world total production 900.0 818.82 800.0 East Asia is the Maize production (million ton) 700.0 main producer of 600.0 maize in Asia 500.0 400.0 166 million tons 300.0 234.30 in 2009 200.0 165.89 164.11 100.0 36.83 20.2% of the 0.0 world’s total World Asia Eastern South- China maize production Asia Eastern Asia
  • 7. Maize production (million ton) in different areas •Maize production in 900.00 800.00 East Asia comes Maize output (million ton) 700.00 600.00 mainly from China 500.00 400.00 •accounted for 99% 300.00 of East Asia’s total 200.00 out put and 20% of 100.00 the world output in 0.00 South- World Asia Eastern Asia China 1961 205.03 31.60 19.40 Eastern Asia 4.71 18.03 2009 1990 483.34 132.47 101.33 16.44 97.21 2009 818.82 234.30 165.89 36.83 164.11
  • 8. Maize Production in East and SE Asia • Indonesia, the Philippines, Thailand, and Vietnam are some other main production countries of maize in Southeast Asia
  • 9. Increment of maize production in the world, East and SE Asia during 1961-2009 and 1990-2009 800 755 World the average 682 growth rate of 700 Eastern Asia South-Eastern Asia Increment (%) 600 500 maize 400 299 production in East and SE 300 200 Asia is faster 124 100 69 64 0 than that of the 1961-2009 1990-2009 world average
  • 10. Maize Production in East and SE Asia • 1961~2009 • maize production and area increased by 2.99 times and 50% in the world • but by 7.55 times and 101% in East Asia • and by 6.82 times and 88% in SE Asia
  • 11. Maize Production in East and SE Asia • The growth rates of both yield and area have been slowing down in the last 2 decades • maize production increased by 69% in the world, but increased by 64% and 124% in East and SE Asia • Maize area increased 21% in the world, but 43% and 7% in East Asia and SE Asia
  • 12. Maize yield in East and SE Asia (1961-2009) 6.0 y = 0.03x + 4.58 5.0 The trend of Maize yield ( Ton/ha) 4.0 maize yield y = 0.04x + 4.59 y=0.11x+1.0 change in 3.0 y=0.11x+0.974 East Asia was 2.0 y = 0.1x + 1.64 contrary to 1.0 y=0.03x+0.790 that in SE 0.0 Asia Eastern Asia South-Eastern Asia China
  • 13. Maize Yield in East and SE Asia • Maize yield increased faster before 1990, and have been slowing down since then in East Asia (esp in China) • However, it has increased faster in SE Asia after 1990 • So the gap between East and SE Asia has been being narrower after 1990
  • 14. Maize Yield in East and SE Asia • Maize yield increment was due to the: – adoption of hybrids – increase of inputs
  • 15. Maize Yield in East Asia • In East Asia (especially in China) • maize yield increased by 220% with an annual gain of 104 kg/ha during 1969- 1989 • but increased by 15% with an annual gain of 35 kg/ha during 1990-2009
  • 16. Maize Yield in SE Asia • In SE Asia • Maize yield increased by 99% with an annual gain of 31 kg/ha before 1990s • but after that, increased by 110% with an annual gain of 99 kg/ha
  • 17. Maize Yield in East and SE Asia • Both the growth rate and the absolute growth of maize yields in decades showed a clear difference between East and SE Asia
  • 18. Yield gains of maize decreasing in China SE Asia 4.4% 3.8% 2000 East Asia 2.7% 1990 2.3% 5.9% 1980 1.9% 1960 1970 4.5% 1960 1970 3.2% 1980 0.2% 1990 1.7% 2000 (Data from FAO) Chinese Academy of Agricultural Sciences
  • 19. Maize production in China Chinese Academy of Agricultural
  • 20. kg/ha Maize yields since 1950s Single-cross hybrids b=23 b=126 Stress tolerance OPVs b=14 b=89 Double-cross hybrids Year Chinese Academy of Agricultural
  • 21. Maize Production in China • Maize yield in China increased by 3.4 times, with an annual gain of 93 kg/ha during the past 5 decades
  • 22. Maize Production in China(略) • It should be noted that maize yield in China increased at 104 kg/ha/year before 1990s, which was even higher than that in Argentina • but after 1990, maize yield increased at 38 kg/ha/year which was much lower than the growth of 152 kg/ha/year in US and 180 kg/ha/year in Argentina • it was even lower than the gains in Vietnam, Myanmar, Indonesia and Brazil (more than 100 kg/ha/year)
  • 23. Maize yields in US, Argentina and China 12 10 y=0.15x+7.0 8 y=0.11x+4.0 y=0.18x+3.8 6 4 y=0.7x+1.6 y=0.04x+4.6 2 y=0.11x+1.0 0 America Argentina China
  • 24. Maize Production in SE Asia • On the contrary, maize yields have increased sharply since the mid-1990s in several SE Asian countries • the annual gains of maize yield in Indonesia, Vietnam and Myanmar are 4.1%,4.6% and 6.7% • and annual gains of 107 kg/ha,143 kg/ha and 110 kg/ha
  • 25. Maize Trade in the Region
  • 26. Maize Trade in the Region • Over the past half-century, the world maize trade has increased significantly • East Asia has been importing maize for a long time, and the imports continue to increase • but some of the SE Asian countries turned into maize exporters in the mid- 1980s
  • 27. East Asia is a Net Importer (2009) 2008 54970- other part of the word 9021- Korea 16460- Japan 220 China -60000 -50000 -40000 -30000 -20000 -10000 0 10000 2008
  • 28. East Asia is a Net Importer (2009) South-east Asia 7% 13% Japan 52% South Korea 28%
  • 29. Maize Trade in the Region (略) • Japan, Korea and China were the main importers • Maize imports in Japan (16-17 million tons annually) accounted for more than half of Asia’s total imports • Korea (8-9 million tons) accounted for 28% of Asia’s total imports
  • 30. Maize Trade in the Region • Imports of Japan and South Korea have been relatively stable since 1990s • China was a net export country in most years • but its export has been steadily declining in the last decade • China has become a net importer of maize in 2010 (1.5 million tons)
  • 31. Conclusion--1 • Asia is a big importer of maize • Demand for maize will continue to increase in Asia
  • 32. Increase the Maize Production to meet tremendous challenges
  • 33. Soil Improvement Small tractors for more than 20 years and made the soil hardpan top fertility soil is 16.5 cm only Prof. Zhang Dongxing
  • 34. Improve the Soil  soil deep loosing tillage  to use subsoiler to break soil hardpan  to increase top fertility soil  to return farm residue and crop straw to the soil
  • 35. Population density and machine  increase the population density in maize production  45,000/ha 60,000/ha  farming machine  new hybrids with stress tolerance and suitable to harvest machine  irrigation works  farmer collectives
  • 36. Maize Yield and Population Density t/ha bu/ac plants/ha plants/ac 1 16 10,000 4,047 2 32 20,000 8,094 3 48 30,000 12,141 4 64 40,000 16,188 5 80 45,000 18,211 6 96 50,000 20,235 7 112 55,000 22,258 8 128 60,000 24,282 9 144 65,000 26,305 10 160 70,000 28,329 11 176 75,000 30,352 12 192 80,000 32,376 13 208 85,000 34,399 14 224 90,000 36,423 Lamkey, 2007, Beijing 36
  • 37. Maize Breeding Strategies in East and SE Asia Stress tolerance Suitable to harvest with machine
  • 38. Maize Breeding in China • The rate of maize yield gains in China has been reducing although the yield potential has been increasing since 1960 when single cross hybrids were introduced
  • 39. What is Yield? • What is yield? -- questioned by Duvick • Yield is stress tolerance ! • Stress tolerance of hybrids has been ignored for almost three decades due to the central planning economic constitution of RD management
  • 40. Risky variety Breeders tend to develop hybrids with tall stature, too much leafy, late maturity or full season, and big ears most of this kind of hybrids are risky under stress 40
  • 41. Plant and ear height (Beijing and Wulumuqi, 2007-2008) 280 130 y = 3.71x + 249.5 270 R2 = 0.50 120 Plant height (cm) Ear height(cm) 260 110 y = 1.15x + 105.4 250 R2 = 0.10 240 100 1950 1960 1970 1980 1990 2000 1950 1960 1970 1980 1990 2000 Year of release Year of release Linear regression suggested that plant and ear height increased.
  • 42. Maturity of hybrids delaied 72.0 Low 71.0 Mid High Maturity (day) 70.0 69.0 68.0 67.0 66.0 1960 1970 1980 1990 2000 2010 Era (Year)
  • 43. Shelling percentage(2008) 85.0 Shelling (%) 80.0 y = -0.19x + 82.2 R2 = 0.03 75.0 1950 1960 1970 1980 1990 2000 Year of release Shelling percentage showed a trend to reduction during 1950s-2000s, especially reduced greatly during 1980s-2000s.
  • 44. 1.40 1.20 y = 0.0889x + 0.6097 Barrent tip (cm) 1.00 0.80 0.60 0.40 0.20 0.00 1950s 1970s 1980s 1990s 2000s 秃尖长度 Year 44
  • 47. Stine Seeds, Iowa 150,000/ha Sept. 2011 have to change row spacing from 76 cm to 50 cm, to increase the population density redesigning the farming machine 47
  • 48. Risky variety • Breeders also tend to develop hybrids with excessive stay green and late maturity during the past 4 decades • caused application of chemical fertilizers in a non-judicious manner • reducing the efficiency of N utilization
  • 49. Heterosis • The contribution of heterosis to yield gains is higher in China than that reported in American hybrids • the contribution of heterosis to yield gains showed weak increase with the year of hybrids release
  • 50. Heterosis incerased (1970-2000) 90 Het(%) 80 70 Relative heterosis (%) 60 66% 69% 50 40 y = 0.09x - 107 R2 = 0.02 30 20 10 0 1960 1970 1980 1990 2000 2010 Year of release (11 environments, 2007-2009) Chinese Academy of Agricultural
  • 51. Schnell, F.W. 1974. Trends and problems in breeding methods for hybrid corn. p. 86–98. In Proc. of the British Poultry Breeders Roundtable, 16th. Birmingham, England. 51
  • 52. Duvick, D.N. 1999. Heterosis: Feeding people and protecting natural resources. p. 19–29. 52 J.G. In Coors and S. Pandey (ed.) The genetics and exploitation of heterosis in crops. ASA, Madison, WI.
  • 53. Campbell, B.T., D.T. Bowman, and D.B. Weaver. 2008. Heterotic eff ects in topcrosses of modern and obsolete cotton cultivars. Crop Sci. 48:593–605. 53
  • 54. Reorienting the Strategy • Maize breeders both in China and SE Asian countries are now actively reorienting the strategy of maize breeding, and develop stress resilient and input responsive hybrids
  • 55. Reorienting the Strategy • increase yield potential • enhance stress tolerance • increase population density in both crop production and breeding nursery
  • 56. Strategies for breeding in Asia--1 • continue to introduce, improve and utilize exotic germplasm to broaden the genetic base in maize breeding programs • to enhance genetic variance in breeding populations • strengthen the pre-breeding efforts
  • 57. Strategy --2 • germplasm should be divided into 2 heterotic groups or 2 alignments • based on the current genetic backgrounds, which will keep the SCA effects in two groups
  • 59. Strategy --3 • improve the germplasm within each group and push them to the opposite directions in terms of gene frequency • which will enhance and accumulate GCA effects in breeding populations of germplasm
  • 60. Strategy -- 4 • improve germplasm and select inbred lines under high population density and other stress conditions based on G×E effects to enhance stress tolerance • double density for inbred line development • plus 15,000/ha for evaluation of new hybrids (Troyer,1981)
  • 61. Strategy -- 5 • multi-location testing and information management • science and art • data and experience • modern machine and IP technology • pipeline and profolio
  • 62. Conclusion--2 • public • private • pre-breeding • pipeline • profolio
  • 63. Conclusion--3 • improve the soil and infrastruture • improve hybrids to meet challenges • stress tolerance is more important than heterosis for hybrid yields • GCA is more important than SCA • reorient the strategy of maize breeding programs
  • 64. Chinese Academy of Agricultural
  • 65. Demand for maize in China - 2020 • 230 million tons by 2020 • Processing 58 mt (26%) • Feeding stuff 151 mt (70%) Chinese Academy of Agricultural
  • 66. Three concepts of yield • Genetic yield (maximum) —The highest yield record • Attainable yield —average yield in Maize Variety Regional Evaluation Trials • Actual yield — statistic data issued by MOA
  • 67. The differences between yields and yield potential great Genetic yieldyield Genetic yield Genetic yield Genetic Attainable yield Attainable yield Attainable yield Actual yield Actual yield Actual yield Attainable yield Attainable yield Attainable yield Actual yield 227/120/127 860/95/30 Dr. RF Hu