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Plant Kingdom
By ASM NAFIS BIOLOGY
for class 11th
Phylogenetic classification systems –
ο‚· It is most acceptable system. It is based on evolutionary relationships between the various organisms.
ο‚· This assumes that organisms belonging to the same taxa have a common ancestor.
Systems of classification –
Artificial System of classification –
ο‚· It used morphological characters such as habit, colour, number and shape of leaves, etc.
ο‚· based mainly on vegetative characters or on the androecium structure (system given by
Linnaeus).
ο‚· Artificial systems gave equal weightage to vegetative and sexual characteristics; this is not
acceptable since we know that often the vegetative characters are more easily affected by
environment.
Cytotaxonomy –It is based on cytological information like chromosome number, structure,
behaviour.
New development in taxonomy –
Numerical Taxonomy –It is carried out using computers and is based on all observable
characteristics. In this way each character is given equal importance and at the same time
hundreds of characters can be considered.
Chemotaxonomy –It is based on the chemical constituents of the plant
Plant Classification
ALGAE / THALLOPHYTA
Algae are chlorophyll-bearing, simple, thalloid, autotrophic and largely aquatic
(both fresh water and marine) organisms.
Some Algae also occur in association with fungi (lichen) and animals (e.g., on
sloth bear).
Size and form of algae –
The unicellular motile – Chlamydomonas,
Unicellular non Motile- Chlorela,
Colonial forms – Volvox
Filamentous Unbranched forms – Ulothrix and Spirogyra,
Filamentous Branched- Cladophora,
Marine and massive plant bodies – kelps
Reproduction in Algae –
οƒ˜ Vegetative reproduction – by fragmentation, each fragment develops into a thallus.
οƒ˜ Asexual reproduction – by the production of different types of spores like
zoospores. They are flagellated (motile) and on germination gives rise to new
plants.
οƒ˜ Sexual reproduction – through fusion of two gametes.
β€’ Isogamous – If gametes are flagellated and similar in size – Chlamydomonas.
β€’ If gametes are non-flagellated and similar in size – Spirogyra.
β€’ Anisogamous – If gametes are dissimilar in size. e.g., some species
of Chlamydomonas.
β€’ Oogamous – Fusion between one large, non-motile female gamete and a smaller,
motile male gamete- Volvox.
Economic importance of Algae –
οƒ˜ Half of the total carbon dioxide fixation on earth is carried out by algae through
photosynthesis.
οƒ˜ Primary producers of energy-rich compounds which form the basis of the food cycles of
all aquatic animals. Many species of Porphyra, Laminaria and Sargassum are used as
food.
οƒ˜ Certain marine brown and red algae produce large amounts of hydrocolloids (water
holding substances), e.g., algin (brown algae) and carrageen (red algae).
οƒ˜ Agar, one of the commercial products obtained from Gelidium and Gracilaria are used to
grow microbes and in preparations of ice-creams and jellies.
οƒ˜ Chlorella and Spirulina are unicellular algae, rich in proteins and are used as food
supplements even by space travellers. (SCP) single cell protein.
β€’ The algae are divided into three main classes (on the basis of pigment and stored food):
Chlorophyceae, Phaeophyceae and Rhodophyceae.
Chlorophyceae (Green algae)
οƒ˜ The plant body may be unicellular, colonial or filamentous. Found in fresh and Marine
Habitat.
οƒ˜ Green due to pigments chlorophyll a and b.
οƒ˜ The chloroplasts may be discoid or plate-like (Ulothrix), cup-shaped (Chlamydomonas),
spiral or ribbon-shaped (Spirogyra).
οƒ˜ pyrenoids located in the chloroplast membrane, contain protein besides starch. And other may
store food in the form of oil droplets.
οƒ˜ Rigid cell wall made of an inner layer of cellulose and an outer layer of pectose.
οƒ˜ Vegetative reproduction – by fragmentation or by formation of different types of spores.
οƒ˜ Asexual reproduction – by flagellated zoospores produced in zoosporangia.
οƒ˜ The sexual reproduction – may be isogamous, anisogamous or oogamous.
β€’ e.g., Chlamydomonas, Ulothrix, Spirogyra, Chara, and Volvox,
Phaeophyceae (Brown algae) also called calps or moss
οƒ˜ Found in marine habitats.
οƒ˜ Present in from simple branched, filamentous forms (Ectocarpus) to profusely branched forms as
represented by kelps.
οƒ˜ They possess chlorophyll a, c, carotenoids and xanthophylls (fucoxanthin).
οƒ˜ Food is stored as complex carbohydrates, which may be in the form of laminarin or mannitol.
οƒ˜ Cell wall made-up of cellulose and has outer coating of gelatinous substance algin.
οƒ˜ Body is usually attached to the substratum by a holdfast, and has a stalk – the stipe and leaf like
photosynthetic organ – the frond.
οƒ˜ Vegetative reproduction – by fragmentation.
οƒ˜ Asexual reproduction – by biflagellate zoospores and have two un-equal laterally attached flagella.
οƒ˜ Sexual reproduction – may be isogamous, anisogamous or oogamous.
o Union of gametes may take place in water or within the oogonium (oogamous species).
o The gametes are pyriform (pear-shaped) and bear two laterally attached flagella.
β€’ e.g., Ectocarpus, Dictyota, Laminaria, Sargassum and Fucus.
Rhodophyceae (Red algae)
οƒ˜ Predominance of the red pigment, r-phycoerythrin, chlorophyll a, d.
οƒ˜ Found close to the surface of water and also at great depths in oceans where
relatively little light penetrates.
οƒ˜ Cell wall is made up of cellulose and covered with agaropectin.
οƒ˜ The food is stored as floridean starch which is very similar to amylopectin and
glycogen in structure.
οƒ˜ Chloroplast lack external endoplasmic reticulum and contain unstaked (stroma)
Thylakoids.
οƒ˜ Vegetative reproduction – by fragmentation.
οƒ˜ Asexual reproduction – by non-motile spores.
οƒ˜ Sexual reproduction – by non-motile gametes. Sexual reproduction is oogamous.
β€’ e.g., Polysiphonia, Porphyra, Gracilaria and Gelidium
BRYOPHYTES (Amphibians of the plant kingdom)
Bryophytes are also called amphibians of the plant kingdom because these
plants can live in soil but are dependent on water for sexual reproduction.
Bryophytes include mosses and liverworts that are found commonly growing in
moist shaded areas in the hills.
Structure / Plant body –
More differentiated than algae, thallus-like, prostrate or erect, and attached to
the rhizoids (root like structure).
They lack true roots, stem or leaves. They may possess root-like, leaf-like or
stem-like structures.
The main plant body of the bryophyte is haploid. It produces
Sex organs –
β€’ The sex organs in bryophytes are multi-cellular.
β€’ The male sex organ is called antheridium. They produce biflagellate antherozoids.
β€’ The female sex organ called archegonium is flask-shaped and produces a single egg.
Fertilization and development –
οƒ˜ The antherozoids are released into water where they come in contact with
archegonium. and fuses with the egg to produce the zygote.
οƒ˜ Zygotes do not undergo reduction division immediately. They produce a multicellular
body called a sporophyte.
οƒ˜ The sporophyte is not free-living but attached to the photosynthetic gametophyte and
derives nourishment from it. (sporophyte is parasite or dependent on gametophyte).
οƒ˜ Some cells of the sporophyte undergo reduction division (meiosis) to produce haploid
spores.
οƒ˜ These spores germinate to produce gametophyte.
Economic importance –
οƒ˜ some mosses provide food for herbaceous mammals, birds and other
animals.
οƒ˜ Species of Sphagnum, a moss, provide peat that have long been used as fuel,
and because of their capacity to hold water as packing material for trans-
shipment of living material.
οƒ˜ Mosses along with lichens are the first organisms to colonise rocks and
hence, are of great ecological importance.
οƒ˜ They decompose rocks making the substrate suitable for the growth of
higher plants.
οƒ˜ Since mosses form dense mats on the soil, they reduce the impact of falling
rain and prevent soil erosion
The bryophytes are divided into liverworts and mosses.
Liverworts
οƒ˜ The plant body of a liverwort is thalloid.
οƒ˜ The thallus is dorsoventral and closely appressed to the substrate.
οƒ˜ The leafy members have tiny leaf-like appendages in two rows on the stem-like structures.
οƒ˜ Asexual reproduction – by fragmentation of thalli, or by the formation of specialised structures
called gemmae.
o Gemmae are green, multicellular, asexual buds, which develop in small receptacles called gemma cups
located on the thalli.
o The gemmae become detached from the parent body and germinate to form new individuals.
οƒ˜ Sexual reproduction – male and female sex organs are produced either on the same or on
different thalli.
οƒ˜ The sporophyte is having capsule.
οƒ˜ After meiosis, spores are produced within the capsule, and disperse with the help of Elators,
these spores germinate to form free-living gametophytes.
β€’ e.g., Riccia and Marchantia.
Mosses
οƒ˜ The predominant stage of the life cycle of a moss is the gametophyte which consists of
two stages.
o The first stage is the protonema stage, which develops directly from a spore. It is a creeping, green,
branched and frequently filamentous stage.
o The second stage is the leafy stage, which develops from the secondary protonema as a lateral bud. It
has upright, slender axes bearing spirally arranged leaves. They are attached to the soil through
multicellular and branched rhizoids. This stage bears the sex organs.
ο‚· Vegetative reproduction – by fragmentation and budding in the secondary protonema.
ο‚· sexual reproduction – by the sex organs antheridia and archegonia, which are produced at
the apex of the leafy shoots.
ο‚· After fertilization – the zygote develops into a sporophyte, consisting of a foot, seta and
capsule. The capsule contains spores, which are formed after meiosis.
ο‚· The mosses have teeth like structure and elaborate mechanism of spore dispersal.
β€’ e.g., Funaria and Sphagnum.
Fig. Bryophytes: A liverworts- Marchantia a) male Thallus b) Female thallus; B Mosses- c)
Funaria, Gamatophyte and Sporophyte; d) Sphagum Gamatophyte
PTERIDOPHYTES
ο‚· The Pteridophytes include horsetails and ferns.
ο‚· First terrestrial plants to possess vascular tissues – xylem and phloem.
ο‚· The pteridophytes are found in cool, damp, shady places though some may nourish well in
sandy-soil conditions.
β€’ Structure / Plant body –
ο‚· The main plant body is a sporophyte which is differentiated into true root, stem and leaves.
ο‚· These organs possess well-differentiated vascular tissues.
ο‚· The leaves are small (microphylls) – Selaginella or large (macrophylls) – ferns.
ο‚· The sporophytes bear sporangia that are subtended by leaf-like appendages called
sporophylls.
ο‚· In some case sporophylls may form distinct compact structures called strobili or cones
(Selaginella, Equisetum).
Life cycle –
οƒ˜ The sporangia produce spores by meiosis in spore mother cells.
οƒ˜ The spores germinate to give rise to inconspicuous, small but
multicellular, free-living, mostly photosynthetic thalloid gametophytes
called prothallus.
οƒ˜ These gametophytes require cool, damp, shady places to grow. Because of
this specific restricted requirement and the need for water for fertilisation,
the spread of living pteridophytes is limited and restricted to narrow
geographical regions.
Sexual Reproduction –
ο‚· The gametophytes bear male and female sex organs called Antheridia and
Archegonia, respectively.
ο‚· Water is required for transfer of Antherozoids (the male gametes) to the
mouth of archegonium.
ο‚· Fusion of male gamete with the egg present in the archegonium result in
the formation of zygote.
Development of zygote –Zygote thereafter produces a multicellular well-
differentiated sporophyte which is the dominant phase of the pteridophytes.
There are two types of sporophytes in pteridophytes –
οƒ˜ Homosporous – all spores are of similar kinds, e.g., In majority of the
pteridophytes.
οƒ˜ Heterosporous – 2 types of spores are produced, (a) small, male
microspores, and (b) large, female megaspores. e.g., Selaginella and the
megaspores and microspores germinate and give rise to female and male
gametophytes, respectively.
β€’ The female gametophytes in these plants are retained on the parent
sporophytes for variable periods.
β€’ The development of the zygotes into young embryos takes place within
the female gametophytes. This event is a precursor to the seed habit
considered an important step in evolution
Economic uses –
οƒ˜ Pteridophytes are used for medicinal purposes and as soil-binders.
οƒ˜ They are also frequently grown as ornamentals.
β€’ The pteridophytes are further classified into four classes:
β€’ 1. Psilopsida – e.g., Psilotum. 2. Lycopsida – e.g., Selaginella, Lycopodium. 3.
Sphenopsida – e.g., Equisetum. 4. Pteropsida – e.g., Dryopteris, Pteris, Adiantum.
GYMNOSPERMS
οƒ˜ Plants in which the ovules are not enclosed by any ovary wall and remain
exposed, both before and after fertilisation.
οƒ˜ The seeds that develop post-fertilisation, are not covered (naked).
Plant body / structure –
οƒ˜ Gymnosperms include medium-sized trees or tall trees and shrubs. One of the
gymnosperms, the giant redwood tree Sequoia is one of the tallest tree species.
οƒ˜ Roots –
o The roots are generally tap roots.
o Roots in Pinus have fungal association, in Cycas small specialized roots called coralloid roots are
associated with N2- fixing cyanobacteria.
οƒ˜ Stem – The stems are unbranched (Cycas) or branched (Pinus, Cedrus).
οƒ˜ Leaves –
o The leaves may be simple or compound.
o In Cycas the pinnate leaves persist for a few years.
o The leaves in gymnosperms are well-adapted to withstand extremes of temperature, humidity and
wind.
o In conifers, the needle-like leaves reduce the surface area.
β€’ Their thick cuticle and sunken stomata also help to reduce water loss
Development of spores and gametophyte –
οƒ˜ The gymnosperms are They produce haploid microspores and megaspores.
οƒ˜ Spores are produced within sporangia that are borne on sporophylls, which are
arranged spirally along an axis to form lax or compact strobili or cones.
Male –
οƒ˜ The strobili bearing micro sporophylls also called male strobili.
οƒ˜ The microspores develop into a male gametophytic generation which is highly
reduced and is conned to only a limited number of cells.
οƒ˜ This reduced male gametophyte is called a pollen grain.
Female –
οƒ˜ The cones bearing mega sporophylls with ovules or megasporangia are called
macrosporangiate or female strobili.
οƒ˜ The megaspore mother cell is differentiated from one of the cells of the nucellus.
οƒ˜ The nucellus is protected by envelopes and the composite structure is called an ovule.
οƒ˜ The megaspore mother cell divides meiotically to form four megaspores.
οƒ˜ One of the megaspores enclosed within the megasporangium (nucellus) develops into a
multicellular female gametophyte that bears two or more archegonia or female sex
organs.
οƒ˜ The multicellular female gametophyte is also retained within megasporangium.
οƒ˜ The male or female cones or strobili may be borne on the same tree –
bisexual/monoecious – Pinus or on different trees – unisexual/dioecious – Cycas.
οƒ˜ In gymnosperms, the male and the female gametophytes do not have an independent
free-living existence. They remain within the sporangia retained on the sporophytes
Pollination and fertilisation –
ο‚· Pollination by air.
ο‚· The pollen grain developed pollen tube on opening of ovule to carry
male gametes towards archegonia in ovules.
ο‚· Following fertilisation, zygote develops into an embryo and the
ovules into seeds. These seeds are not covered.
ANGIOSPERMS (Flowering plants)
οƒ˜ In the angiosperms pollen grains and ovules are developed in specialized
structures called flower.
οƒ˜ In angiosperms, the seeds are enclosed by fruits.
οƒ˜ The angiosperms are present in wide range of habitat.
οƒ˜ Smallest angiosperm – Wolfia.
οƒ˜ Tallest angiosperm – Eucalyptus.
οƒ˜ They provide us with food, fodder, fuel, medicines and several other
commercially important products.
οƒ˜ They are divided into two classes: the dicotyledons and the monocotyledons.
οƒ˜ The dicotyledons are characterised by having two cotyledons in their seeds
while the monocotyledons have only one
οƒ˜ Some Plant having only one gamate either of male or female called monoecious and other
have both called dioecious.
οƒ˜ The male sex organs in a flower is the stamen. Each stamen consists of a slender filament
with an anther at the tip. The anthers, following meiosis, produce pollen grains.
οƒ˜ The female sex organs in a flower is the pistil or the carpel. Pistil consists of an ovary
enclosing one to many ovules. Within ovules highly reduced female gametophytes
(embryosacs) are present.
οƒ˜ The embryo-sac formation is preceded by meiosis. Hence, each of the cells of an embryo-sac
is haploid.
o Each embryo-sac has a three-celled egg apparatus – one egg cell, two synergids, three antipodal cells and
two polar nuclei.
o The two polar nuclei fuse to produce a diploid secondary nucleus.
οƒ˜ Pollen grain, after dispersal from the anthers, are carried by wind or various other agencies to
the stigma of a pistil. This is termed as pollination.
οƒ˜ The pollen grains germinate on the stigma and the resulting pollen tubes grow through the
tissues of stigma and style and reach the ovule. The pollen tubes enter the embryo-sac where
two male gametes are discharged.
Double fertilisation –
οƒ˜ One of the male gametes fuses with the egg cell to form a zygote (syngamy). The
other male gamete fuses with the diploid secondary nucleus to produce the
triploid primary endosperm nucleus (PEN). This process is known as Triple
Fusion and dauble fertilization.
οƒ˜ Because of the involvement of two fusions, this event is termed as double
fertilisation.
Development of zygote –
οƒ˜ The zygote develops into an embryo (with one or two cotyledons) and the PEN
develops into endosperm which provides nourishment to the developing embryo.
οƒ˜ The synergids and antipodal degenerate after fertilisation.
οƒ˜ During these events the ovules develop into seeds and the ovaries develop into
fruit.
PLANT LIFE CYCLES AND ALTERNATION OF GENERATIONS
β€’ In plants, both haploid and diploid cells can divide by mitosis. This
ability leads to the formation of different plant bodies – haploid and
diploid and alternation of generation.
β€’ This plant body represents a gametophyte. Following fertilisation the
zygote also divides by mitosis to produce a diploid sporophytic plant
body.
β€’ Haploid spores are produced by this plant body by meiosis. These in
turn, divide by mitosis to form a haploid plant body once again.
Different types of life cycle in plants –
1. Haplontic life cycle –
β€’ In this, Sporophytic generation is represented only by the one-celled zygote and there are no free-
living sporophytes. Meiosis in the zygote results in the formation of haploid spores. The haploid
spores divide mitotically and form the gametophyte. The dominant, photosynthetic phase in such
plants is the free-living gametophyte.
β€’ e.g., Volvox, Spirogyra and some species of Chlamydomonas. (most of the algae)
2. Diplontic life cycle –
β€’ In this, diploid sporophyte is the dominant, photosynthetic, independent phase of the plant. The
gametophytic phase is represented by the single to few-celled haploid gametophyte.
β€’ e.g., All seed-bearing plants i.e. gymnosperms and angiosperms, some algae like Fucus.
3. Haplo-diplontic / Intermediate life cycle –
β€’ In this, both gametophytic and sporophytic phases are multicellular and often free living.
β€’ e.g., Bryophytes (dominant phase – gametophyte) and pteridophytes (dominanat phase –
sporophyte), some algae like Ectocarpus, Polysiphonia.
END OF THIS CHAPTER

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Plant kingdom (11th Biology) for complete 11th and 12th notes 1999 and for PPT(@4999) call/what up on 8076540093

  • 1. Plant Kingdom By ASM NAFIS BIOLOGY for class 11th
  • 2. Phylogenetic classification systems – ο‚· It is most acceptable system. It is based on evolutionary relationships between the various organisms. ο‚· This assumes that organisms belonging to the same taxa have a common ancestor. Systems of classification – Artificial System of classification – ο‚· It used morphological characters such as habit, colour, number and shape of leaves, etc. ο‚· based mainly on vegetative characters or on the androecium structure (system given by Linnaeus). ο‚· Artificial systems gave equal weightage to vegetative and sexual characteristics; this is not acceptable since we know that often the vegetative characters are more easily affected by environment.
  • 3. Cytotaxonomy –It is based on cytological information like chromosome number, structure, behaviour. New development in taxonomy – Numerical Taxonomy –It is carried out using computers and is based on all observable characteristics. In this way each character is given equal importance and at the same time hundreds of characters can be considered. Chemotaxonomy –It is based on the chemical constituents of the plant
  • 5. ALGAE / THALLOPHYTA Algae are chlorophyll-bearing, simple, thalloid, autotrophic and largely aquatic (both fresh water and marine) organisms. Some Algae also occur in association with fungi (lichen) and animals (e.g., on sloth bear). Size and form of algae – The unicellular motile – Chlamydomonas, Unicellular non Motile- Chlorela, Colonial forms – Volvox Filamentous Unbranched forms – Ulothrix and Spirogyra, Filamentous Branched- Cladophora, Marine and massive plant bodies – kelps
  • 6. Reproduction in Algae – οƒ˜ Vegetative reproduction – by fragmentation, each fragment develops into a thallus. οƒ˜ Asexual reproduction – by the production of different types of spores like zoospores. They are flagellated (motile) and on germination gives rise to new plants. οƒ˜ Sexual reproduction – through fusion of two gametes. β€’ Isogamous – If gametes are flagellated and similar in size – Chlamydomonas. β€’ If gametes are non-flagellated and similar in size – Spirogyra. β€’ Anisogamous – If gametes are dissimilar in size. e.g., some species of Chlamydomonas. β€’ Oogamous – Fusion between one large, non-motile female gamete and a smaller, motile male gamete- Volvox.
  • 7. Economic importance of Algae – οƒ˜ Half of the total carbon dioxide fixation on earth is carried out by algae through photosynthesis. οƒ˜ Primary producers of energy-rich compounds which form the basis of the food cycles of all aquatic animals. Many species of Porphyra, Laminaria and Sargassum are used as food. οƒ˜ Certain marine brown and red algae produce large amounts of hydrocolloids (water holding substances), e.g., algin (brown algae) and carrageen (red algae). οƒ˜ Agar, one of the commercial products obtained from Gelidium and Gracilaria are used to grow microbes and in preparations of ice-creams and jellies. οƒ˜ Chlorella and Spirulina are unicellular algae, rich in proteins and are used as food supplements even by space travellers. (SCP) single cell protein. β€’ The algae are divided into three main classes (on the basis of pigment and stored food): Chlorophyceae, Phaeophyceae and Rhodophyceae.
  • 8. Chlorophyceae (Green algae) οƒ˜ The plant body may be unicellular, colonial or filamentous. Found in fresh and Marine Habitat. οƒ˜ Green due to pigments chlorophyll a and b. οƒ˜ The chloroplasts may be discoid or plate-like (Ulothrix), cup-shaped (Chlamydomonas), spiral or ribbon-shaped (Spirogyra). οƒ˜ pyrenoids located in the chloroplast membrane, contain protein besides starch. And other may store food in the form of oil droplets. οƒ˜ Rigid cell wall made of an inner layer of cellulose and an outer layer of pectose. οƒ˜ Vegetative reproduction – by fragmentation or by formation of different types of spores. οƒ˜ Asexual reproduction – by flagellated zoospores produced in zoosporangia. οƒ˜ The sexual reproduction – may be isogamous, anisogamous or oogamous. β€’ e.g., Chlamydomonas, Ulothrix, Spirogyra, Chara, and Volvox,
  • 9.
  • 10. Phaeophyceae (Brown algae) also called calps or moss οƒ˜ Found in marine habitats. οƒ˜ Present in from simple branched, filamentous forms (Ectocarpus) to profusely branched forms as represented by kelps. οƒ˜ They possess chlorophyll a, c, carotenoids and xanthophylls (fucoxanthin). οƒ˜ Food is stored as complex carbohydrates, which may be in the form of laminarin or mannitol. οƒ˜ Cell wall made-up of cellulose and has outer coating of gelatinous substance algin. οƒ˜ Body is usually attached to the substratum by a holdfast, and has a stalk – the stipe and leaf like photosynthetic organ – the frond. οƒ˜ Vegetative reproduction – by fragmentation. οƒ˜ Asexual reproduction – by biflagellate zoospores and have two un-equal laterally attached flagella. οƒ˜ Sexual reproduction – may be isogamous, anisogamous or oogamous. o Union of gametes may take place in water or within the oogonium (oogamous species). o The gametes are pyriform (pear-shaped) and bear two laterally attached flagella. β€’ e.g., Ectocarpus, Dictyota, Laminaria, Sargassum and Fucus.
  • 11.
  • 12. Rhodophyceae (Red algae) οƒ˜ Predominance of the red pigment, r-phycoerythrin, chlorophyll a, d. οƒ˜ Found close to the surface of water and also at great depths in oceans where relatively little light penetrates. οƒ˜ Cell wall is made up of cellulose and covered with agaropectin. οƒ˜ The food is stored as floridean starch which is very similar to amylopectin and glycogen in structure. οƒ˜ Chloroplast lack external endoplasmic reticulum and contain unstaked (stroma) Thylakoids. οƒ˜ Vegetative reproduction – by fragmentation. οƒ˜ Asexual reproduction – by non-motile spores. οƒ˜ Sexual reproduction – by non-motile gametes. Sexual reproduction is oogamous. β€’ e.g., Polysiphonia, Porphyra, Gracilaria and Gelidium
  • 13.
  • 14. BRYOPHYTES (Amphibians of the plant kingdom) Bryophytes are also called amphibians of the plant kingdom because these plants can live in soil but are dependent on water for sexual reproduction. Bryophytes include mosses and liverworts that are found commonly growing in moist shaded areas in the hills. Structure / Plant body – More differentiated than algae, thallus-like, prostrate or erect, and attached to the rhizoids (root like structure). They lack true roots, stem or leaves. They may possess root-like, leaf-like or stem-like structures. The main plant body of the bryophyte is haploid. It produces
  • 15. Sex organs – β€’ The sex organs in bryophytes are multi-cellular. β€’ The male sex organ is called antheridium. They produce biflagellate antherozoids. β€’ The female sex organ called archegonium is flask-shaped and produces a single egg. Fertilization and development – οƒ˜ The antherozoids are released into water where they come in contact with archegonium. and fuses with the egg to produce the zygote. οƒ˜ Zygotes do not undergo reduction division immediately. They produce a multicellular body called a sporophyte. οƒ˜ The sporophyte is not free-living but attached to the photosynthetic gametophyte and derives nourishment from it. (sporophyte is parasite or dependent on gametophyte). οƒ˜ Some cells of the sporophyte undergo reduction division (meiosis) to produce haploid spores. οƒ˜ These spores germinate to produce gametophyte.
  • 16. Economic importance – οƒ˜ some mosses provide food for herbaceous mammals, birds and other animals. οƒ˜ Species of Sphagnum, a moss, provide peat that have long been used as fuel, and because of their capacity to hold water as packing material for trans- shipment of living material. οƒ˜ Mosses along with lichens are the first organisms to colonise rocks and hence, are of great ecological importance. οƒ˜ They decompose rocks making the substrate suitable for the growth of higher plants. οƒ˜ Since mosses form dense mats on the soil, they reduce the impact of falling rain and prevent soil erosion
  • 17. The bryophytes are divided into liverworts and mosses. Liverworts οƒ˜ The plant body of a liverwort is thalloid. οƒ˜ The thallus is dorsoventral and closely appressed to the substrate. οƒ˜ The leafy members have tiny leaf-like appendages in two rows on the stem-like structures. οƒ˜ Asexual reproduction – by fragmentation of thalli, or by the formation of specialised structures called gemmae. o Gemmae are green, multicellular, asexual buds, which develop in small receptacles called gemma cups located on the thalli. o The gemmae become detached from the parent body and germinate to form new individuals. οƒ˜ Sexual reproduction – male and female sex organs are produced either on the same or on different thalli. οƒ˜ The sporophyte is having capsule. οƒ˜ After meiosis, spores are produced within the capsule, and disperse with the help of Elators, these spores germinate to form free-living gametophytes. β€’ e.g., Riccia and Marchantia.
  • 18. Mosses οƒ˜ The predominant stage of the life cycle of a moss is the gametophyte which consists of two stages. o The first stage is the protonema stage, which develops directly from a spore. It is a creeping, green, branched and frequently filamentous stage. o The second stage is the leafy stage, which develops from the secondary protonema as a lateral bud. It has upright, slender axes bearing spirally arranged leaves. They are attached to the soil through multicellular and branched rhizoids. This stage bears the sex organs. ο‚· Vegetative reproduction – by fragmentation and budding in the secondary protonema. ο‚· sexual reproduction – by the sex organs antheridia and archegonia, which are produced at the apex of the leafy shoots. ο‚· After fertilization – the zygote develops into a sporophyte, consisting of a foot, seta and capsule. The capsule contains spores, which are formed after meiosis. ο‚· The mosses have teeth like structure and elaborate mechanism of spore dispersal. β€’ e.g., Funaria and Sphagnum.
  • 19. Fig. Bryophytes: A liverworts- Marchantia a) male Thallus b) Female thallus; B Mosses- c) Funaria, Gamatophyte and Sporophyte; d) Sphagum Gamatophyte
  • 20. PTERIDOPHYTES ο‚· The Pteridophytes include horsetails and ferns. ο‚· First terrestrial plants to possess vascular tissues – xylem and phloem. ο‚· The pteridophytes are found in cool, damp, shady places though some may nourish well in sandy-soil conditions. β€’ Structure / Plant body – ο‚· The main plant body is a sporophyte which is differentiated into true root, stem and leaves. ο‚· These organs possess well-differentiated vascular tissues. ο‚· The leaves are small (microphylls) – Selaginella or large (macrophylls) – ferns. ο‚· The sporophytes bear sporangia that are subtended by leaf-like appendages called sporophylls. ο‚· In some case sporophylls may form distinct compact structures called strobili or cones (Selaginella, Equisetum).
  • 21. Life cycle – οƒ˜ The sporangia produce spores by meiosis in spore mother cells. οƒ˜ The spores germinate to give rise to inconspicuous, small but multicellular, free-living, mostly photosynthetic thalloid gametophytes called prothallus. οƒ˜ These gametophytes require cool, damp, shady places to grow. Because of this specific restricted requirement and the need for water for fertilisation, the spread of living pteridophytes is limited and restricted to narrow geographical regions.
  • 22. Sexual Reproduction – ο‚· The gametophytes bear male and female sex organs called Antheridia and Archegonia, respectively. ο‚· Water is required for transfer of Antherozoids (the male gametes) to the mouth of archegonium. ο‚· Fusion of male gamete with the egg present in the archegonium result in the formation of zygote. Development of zygote –Zygote thereafter produces a multicellular well- differentiated sporophyte which is the dominant phase of the pteridophytes.
  • 23. There are two types of sporophytes in pteridophytes – οƒ˜ Homosporous – all spores are of similar kinds, e.g., In majority of the pteridophytes. οƒ˜ Heterosporous – 2 types of spores are produced, (a) small, male microspores, and (b) large, female megaspores. e.g., Selaginella and the megaspores and microspores germinate and give rise to female and male gametophytes, respectively. β€’ The female gametophytes in these plants are retained on the parent sporophytes for variable periods. β€’ The development of the zygotes into young embryos takes place within the female gametophytes. This event is a precursor to the seed habit considered an important step in evolution
  • 24. Economic uses – οƒ˜ Pteridophytes are used for medicinal purposes and as soil-binders. οƒ˜ They are also frequently grown as ornamentals. β€’ The pteridophytes are further classified into four classes: β€’ 1. Psilopsida – e.g., Psilotum. 2. Lycopsida – e.g., Selaginella, Lycopodium. 3. Sphenopsida – e.g., Equisetum. 4. Pteropsida – e.g., Dryopteris, Pteris, Adiantum.
  • 25.
  • 26. GYMNOSPERMS οƒ˜ Plants in which the ovules are not enclosed by any ovary wall and remain exposed, both before and after fertilisation. οƒ˜ The seeds that develop post-fertilisation, are not covered (naked).
  • 27. Plant body / structure – οƒ˜ Gymnosperms include medium-sized trees or tall trees and shrubs. One of the gymnosperms, the giant redwood tree Sequoia is one of the tallest tree species. οƒ˜ Roots – o The roots are generally tap roots. o Roots in Pinus have fungal association, in Cycas small specialized roots called coralloid roots are associated with N2- fixing cyanobacteria. οƒ˜ Stem – The stems are unbranched (Cycas) or branched (Pinus, Cedrus). οƒ˜ Leaves – o The leaves may be simple or compound. o In Cycas the pinnate leaves persist for a few years. o The leaves in gymnosperms are well-adapted to withstand extremes of temperature, humidity and wind. o In conifers, the needle-like leaves reduce the surface area. β€’ Their thick cuticle and sunken stomata also help to reduce water loss
  • 28. Development of spores and gametophyte – οƒ˜ The gymnosperms are They produce haploid microspores and megaspores. οƒ˜ Spores are produced within sporangia that are borne on sporophylls, which are arranged spirally along an axis to form lax or compact strobili or cones. Male – οƒ˜ The strobili bearing micro sporophylls also called male strobili. οƒ˜ The microspores develop into a male gametophytic generation which is highly reduced and is conned to only a limited number of cells. οƒ˜ This reduced male gametophyte is called a pollen grain.
  • 29. Female – οƒ˜ The cones bearing mega sporophylls with ovules or megasporangia are called macrosporangiate or female strobili. οƒ˜ The megaspore mother cell is differentiated from one of the cells of the nucellus. οƒ˜ The nucellus is protected by envelopes and the composite structure is called an ovule. οƒ˜ The megaspore mother cell divides meiotically to form four megaspores. οƒ˜ One of the megaspores enclosed within the megasporangium (nucellus) develops into a multicellular female gametophyte that bears two or more archegonia or female sex organs. οƒ˜ The multicellular female gametophyte is also retained within megasporangium. οƒ˜ The male or female cones or strobili may be borne on the same tree – bisexual/monoecious – Pinus or on different trees – unisexual/dioecious – Cycas. οƒ˜ In gymnosperms, the male and the female gametophytes do not have an independent free-living existence. They remain within the sporangia retained on the sporophytes
  • 30. Pollination and fertilisation – ο‚· Pollination by air. ο‚· The pollen grain developed pollen tube on opening of ovule to carry male gametes towards archegonia in ovules. ο‚· Following fertilisation, zygote develops into an embryo and the ovules into seeds. These seeds are not covered.
  • 31.
  • 32. ANGIOSPERMS (Flowering plants) οƒ˜ In the angiosperms pollen grains and ovules are developed in specialized structures called flower. οƒ˜ In angiosperms, the seeds are enclosed by fruits. οƒ˜ The angiosperms are present in wide range of habitat. οƒ˜ Smallest angiosperm – Wolfia. οƒ˜ Tallest angiosperm – Eucalyptus. οƒ˜ They provide us with food, fodder, fuel, medicines and several other commercially important products. οƒ˜ They are divided into two classes: the dicotyledons and the monocotyledons. οƒ˜ The dicotyledons are characterised by having two cotyledons in their seeds while the monocotyledons have only one
  • 33. οƒ˜ Some Plant having only one gamate either of male or female called monoecious and other have both called dioecious. οƒ˜ The male sex organs in a flower is the stamen. Each stamen consists of a slender filament with an anther at the tip. The anthers, following meiosis, produce pollen grains. οƒ˜ The female sex organs in a flower is the pistil or the carpel. Pistil consists of an ovary enclosing one to many ovules. Within ovules highly reduced female gametophytes (embryosacs) are present. οƒ˜ The embryo-sac formation is preceded by meiosis. Hence, each of the cells of an embryo-sac is haploid. o Each embryo-sac has a three-celled egg apparatus – one egg cell, two synergids, three antipodal cells and two polar nuclei. o The two polar nuclei fuse to produce a diploid secondary nucleus. οƒ˜ Pollen grain, after dispersal from the anthers, are carried by wind or various other agencies to the stigma of a pistil. This is termed as pollination. οƒ˜ The pollen grains germinate on the stigma and the resulting pollen tubes grow through the tissues of stigma and style and reach the ovule. The pollen tubes enter the embryo-sac where two male gametes are discharged.
  • 34. Double fertilisation – οƒ˜ One of the male gametes fuses with the egg cell to form a zygote (syngamy). The other male gamete fuses with the diploid secondary nucleus to produce the triploid primary endosperm nucleus (PEN). This process is known as Triple Fusion and dauble fertilization. οƒ˜ Because of the involvement of two fusions, this event is termed as double fertilisation. Development of zygote – οƒ˜ The zygote develops into an embryo (with one or two cotyledons) and the PEN develops into endosperm which provides nourishment to the developing embryo. οƒ˜ The synergids and antipodal degenerate after fertilisation. οƒ˜ During these events the ovules develop into seeds and the ovaries develop into fruit.
  • 35.
  • 36. PLANT LIFE CYCLES AND ALTERNATION OF GENERATIONS β€’ In plants, both haploid and diploid cells can divide by mitosis. This ability leads to the formation of different plant bodies – haploid and diploid and alternation of generation. β€’ This plant body represents a gametophyte. Following fertilisation the zygote also divides by mitosis to produce a diploid sporophytic plant body. β€’ Haploid spores are produced by this plant body by meiosis. These in turn, divide by mitosis to form a haploid plant body once again.
  • 37. Different types of life cycle in plants – 1. Haplontic life cycle – β€’ In this, Sporophytic generation is represented only by the one-celled zygote and there are no free- living sporophytes. Meiosis in the zygote results in the formation of haploid spores. The haploid spores divide mitotically and form the gametophyte. The dominant, photosynthetic phase in such plants is the free-living gametophyte. β€’ e.g., Volvox, Spirogyra and some species of Chlamydomonas. (most of the algae) 2. Diplontic life cycle – β€’ In this, diploid sporophyte is the dominant, photosynthetic, independent phase of the plant. The gametophytic phase is represented by the single to few-celled haploid gametophyte. β€’ e.g., All seed-bearing plants i.e. gymnosperms and angiosperms, some algae like Fucus. 3. Haplo-diplontic / Intermediate life cycle – β€’ In this, both gametophytic and sporophytic phases are multicellular and often free living. β€’ e.g., Bryophytes (dominant phase – gametophyte) and pteridophytes (dominanat phase – sporophyte), some algae like Ectocarpus, Polysiphonia.
  • 38. END OF THIS CHAPTER