Slides from a talk presented at the 2014 London Evolutionary Research Network (LERN) conference. Abstract: An evolutionary interpretation of cooperative behaviours must ultimately include an increase the inclusive fitness of actors as well as recipients. At the proximate level, mechanisms for encouraging and maintaining cooperation include factors such as kin discrimination, limited dispersal as well as direct and indirect reciprocity. However humans are also known for co-operating with individuals who are not necessarily close relatives, and often in co-operative groups. Here, we aim to quantify the relative importance of kinship and social group membership as mediators of cooperative behaviour. Using a modified, externally valid Dictator Game, we test whether indigenous Saami reindeer herders in Norway preferentially give gifts to genetic relatives or to members of their cooperative herding group (the ‘siida’). Membership of the same siida strongly increased the odds of receiving a gift. Kinship had a small (and not statistically significant) effect, even for close relatives. Gifts were not preferentially given to younger family members. These patterns suggest that social grouping can trump genetic factors in mediating cooperative behaviour in this population. This is likely due to the importance of herding groups in day-to-day subsistence.