23. Relative Payoff Sum
Si (t) = xSi (t
1) + (1
x)ri + Pi (t)
where 0 < x < 1 is a memory factor,
ri > 0 is the residual value associated with alternative i,
Pi (t) is the payo to alternative i at time t, and
Si (t) is the value that the animal places on the behavioural alternative i at
time t.
24. Relative Payoff Sum
Si (t) = xSi (t
1) + (1
x)ri + Pi (t)
where 0 < x < 1 is a memory factor,
ri > 0 is the residual value associated with alternative i,
Pi (t) is the payo to alternative i at time t, and
Si (t) is the value that the animal places on the behavioural alternative i at
time t.
25. Relative Payoff Sum
Si (t) = xSi (t
1) + (1
x)ri + Pi (t)
where 0 < x < 1 is a memory factor,
ri > 0 is the residual value associated with alternative i,
Pi (t) is the payo to alternative i at time t, and
Si (t) is the value that the animal places on the behavioural alternative i at
time t.
26. Relative Payoff Sum
Si (t) = xSi (t
1) + (1
x)ri + Pi (t)
where 0 < x < 1 is a memory factor,
ri > 0 is the residual value associated with alternative i,
Pi (t) is the payo to alternative i at time t, and
Si (t) is the value that the animal places on the behavioural alternative i at
time t.
27. Relative Payoff Sum
Si (t) = xSi (t
1) + (1
x)ri + Pi (t)
where 0 < x < 1 is a memory factor,
ri > 0 is the residual value associated with alternative i,
Pi (t) is the payo to alternative i at time t, and
Si (t) is the value that the animal places on the behavioural alternative i at
time t.
28. Perfect Memory
Si (t) =
+ Ri (t)/(⇥ + Ni (t))
where Ri (t) is the cumulative payo s from alternative i to time t,
Ni (t) is the number of time periods from the beginning in which the option
was selected,
and ⇥ are parameters.
29. Perfect Memory
Si (t) =
+ Ri (t)/(⇥ + Ni (t))
where Ri (t) is the cumulative payo s from alternative i to time t,
Ni (t) is the number of time periods from the beginning in which the option
was selected,
and ⇥ are parameters.
30. Perfect Memory
Si (t) =
+ Ri (t)/(⇥ + Ni (t))
where Ri (t) is the cumulative payo s from alternative i to time t,
Ni (t) is the number of time periods from the beginning in which the option
was selected,
and ⇥ are parameters.
31. Perfect Memory
Si (t) =
+ Ri (t)/(⇥ + Ni (t))
where Ri (t) is the cumulative payo s from alternative i to time t,
Ni (t) is the number of time periods from the beginning in which the option
was selected,
and ⇥ are parameters.
32. Linear Operator
Si (t) = xSi (t
1) + (1
x)Pi (t)
where 0 < x < 1 is a memory factor,
Pi (t) is the payo to alternative i at time t, and
Si (t) is the value that the animal places on the behavioural alternative i at
time t.
34. Bird Start
At a patch
with food?
Yes
Feed
NO
Produce or
scrounge?
Scrounge
Produce
Move
randomly
No
Any
conspecifics
feeding?
Move to
closest
Yes
There yet?
Yes
No
Closest still
feeding?
No
35. • 5 or 10 birds.
Bird Start
At a patch
with food?
Yes
• Foraging grid is
Feed
NO
a regular 10x10
grid, with
movement in
the 4 cardinal
directions.
Produce or
scrounge?
Scrounge
Produce
Move
randomly
No
Any
conspecifics
feeding?
Move to
closest
Yes
There yet?
• 20 patches on
Yes
No
Closest still
feeding?
No
the grid, with 10
or 20 food items
in each.
36.
37. Relative
Payoff Sum?
Si (t) = xSi (t
1) + (1
Perfect
Memory?
Si (t) =
Linear
Operator?
Si (t) = xSi (t
x)ri + Pi (t)
+ Ri (t)/(⇥ + Ni (t))
1) + (1
x)Pi (t)
38. Relative
Payoff Sum?
Si (t) = xSi (t
1) + (1
Perfect
Memory?
Si (t) =
Linear
Operator?
Si (t) = xSi (t
x)ri + Pi (t)
+ Ri (t)/(⇥ + Ni (t))
1) + (1
x)Pi (t)
39. Relative
Payoff Sum?
Si (t) = xSi (t
1) + (1
Perfect
Memory?
Si (t) =
Linear
Operator?
Si (t) = xSi (t
x)ri + Pi (t)
+ Ri (t)/(⇥ + Ni (t))
1) + (1
x)Pi (t)
Multiple stable rules with multiple parameters?
47. Relative
Payoff Sum
Si (t) = xSi (t
1) + (1
x)ri + Pi (t)
rp >> rs for large population sizes.
5
4
Value
assigned
to
3
behaviour
2
Producer residual
1
Scrounger residual
-1
0
1
2
3
4
5
Time without payo! to behaviour
6
7
8
49. • Under the assumptions of this model,
the Relative Payoff Sum rule is optimal.
• Whether RPS is favored depends on
payoff variance:
• low variance = more attractive power.
• Differences in residuals gives a
prediction for empirical tests.