In Cyperaceae plants like sedges, only one of the four nuclei formed after meiosis functions during pollen development, while the other three degenerate. The functional nucleus remains in the center and divides to form a vegetative cell and generative cell, while the non-functional nuclei are pushed to the side and may begin to divide abortively before degenerating. As a result, within each pollen tetrad of Cyperaceae, only one pollen grain becomes functional, similar to how one megaspore becomes functional during embryo sac development in these plants.