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PHYSICAL
DORMANCY
SEED DORMANCY
Seed dormancy is an evolutionary adaptation that
prevents seeds from germinating during unsuitable
ecological conditions that would typically lead to a
low probability of seedling survival.
Dormant seeds do not germinate in a specified
period of time under a combination of
environmental factors that are normally conducive
to the germination of non-dormant seeds.
 Seed dormancy can be divided into two major
categories based on what part of the seed produces
dormancy: Exogenous and Endogenous.
There are three types of inherent dormancy based
on their mode of action: Physical, Physiological and
Morphological.
EXOGENOUS DORMANCY
Exogenous dormancy is caused by
conditions outside the embryo and is
often broken down into three
subgroups:
Physical dormancy
Mechanical dormancy
Chemical dormancy
PHYSICAL DORMANCY
• Dormancy caused by an impermeable seed coat is known as physical
dormancy.
• Physical dormancy is the result of impermeable layer(s) that develops
during maturation and drying of the seed or fruit.
• This impermeable layer prevents the seed from taking up water or gases.
• As a result, the seed is prevented from germinating until dormancy is
broken.
• Physical dormancy is believed to have developed more than 100 million
years ago.
• Physical dormancy has been identified in the seeds of plants across 16
angiosperm families like Anacardiaceae,Asteraceae, Cucurbitaceae,
Fabaceae,Malvaceae
REASON FOR THE PHYSICAL DORMANCY
• Generally, physical dormancy is the result of one or
more palisade layers in the fruit or seed coat.
• These layers are lignified with malpighian cells tightly
packed together and impregnated with water-
repellent.
• Specialised structures, which function as a "water-
gap", are associated with the impermeable layers of
the seed to prevent the uptake of water.
• The water-gap is closed at seed maturity and is
opened in response to the appropriate environmental
signal.
• Breaking physical dormancy involves the disruption of
these specialised structures within the seed, and acts
as an environmental signal detector for germination.
• For example,In PY-dormant Fabaceae species (Rosids) the
lens (strophiole) is the spezialized structure localized in the
hilum region that controls water-impermeability.
• In PY-dormant Anacardiaceae species (Rosids), e.g. Rhus
spp., the endocarp consists of three water-impermeable
palisade layers (macrosclereids, osteosclereis,
brachysclereids) and the outer crystalliferous layer.
• Different types of chalazal plugs/caps are the specialized
structures that control water-impermeability of other seeds,
e.g. in PY-dormant species of Bixaceae, Cistaceae, and
Malvaceae.
• Therefore, palisade layers in the fruit perform the functional
role of preventing water uptake.
TREATMENT TO BREAK PHYSICAL DORMANCY
• In natural systems, physical dormancy is broken by
several factors including high temperatures, fluctuating
temperatures, fire, freezing/thawing, drying or passage
through the digestive tracts of animals.
• But the commonly followed method is
SCARIFICATION
• Acid
• Mechanical
• Physical treatment – hot water treatment
SCARIFICATION
• Any treatments may be physical or chemical that
weakens or softens the seed coat is known as
scarification.
• This method is more applicable to Malvaceae and
Leguminaceae group of seeds.
ACID SCARIFICATION
• By using concentrated H2SO4 @ 100ml/kg of seed for 2-3
minutes treatments dormancy can be overcome in the
seeds.
• The duration of treatment will vary and it depends on
type and nature of seed coat.
• E.g. Tree crops 1-3 hours, Rose seeds, treat the seed
partially with acid and then given with warm
stratification.
MECHANICAL SCARIFICATION
• Seeds are rubbed on a sand paper or with a help
of mechanical scarifier or by puncturing on seed
coat with the help of needle to enhance / increase
the moisture absorption by seeds.
• E.g. Bitter gourd for sand scarification, sand and
seed 2:1 ratio should be followed. Rub against
hard surface of seed for 5 to 10 minutes.
HOT WATER TREATMENTS
• It is effective in case of leguminous seeds.
• The seeds should be soaked in boiled water for
1-5 minutes in 60-80 °C.
SST_21.pptx

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SST_21.pptx

  • 2. SEED DORMANCY Seed dormancy is an evolutionary adaptation that prevents seeds from germinating during unsuitable ecological conditions that would typically lead to a low probability of seedling survival. Dormant seeds do not germinate in a specified period of time under a combination of environmental factors that are normally conducive to the germination of non-dormant seeds.  Seed dormancy can be divided into two major categories based on what part of the seed produces dormancy: Exogenous and Endogenous. There are three types of inherent dormancy based on their mode of action: Physical, Physiological and Morphological.
  • 3. EXOGENOUS DORMANCY Exogenous dormancy is caused by conditions outside the embryo and is often broken down into three subgroups: Physical dormancy Mechanical dormancy Chemical dormancy
  • 4. PHYSICAL DORMANCY • Dormancy caused by an impermeable seed coat is known as physical dormancy. • Physical dormancy is the result of impermeable layer(s) that develops during maturation and drying of the seed or fruit. • This impermeable layer prevents the seed from taking up water or gases. • As a result, the seed is prevented from germinating until dormancy is broken. • Physical dormancy is believed to have developed more than 100 million years ago. • Physical dormancy has been identified in the seeds of plants across 16 angiosperm families like Anacardiaceae,Asteraceae, Cucurbitaceae, Fabaceae,Malvaceae
  • 5. REASON FOR THE PHYSICAL DORMANCY • Generally, physical dormancy is the result of one or more palisade layers in the fruit or seed coat. • These layers are lignified with malpighian cells tightly packed together and impregnated with water- repellent. • Specialised structures, which function as a "water- gap", are associated with the impermeable layers of the seed to prevent the uptake of water. • The water-gap is closed at seed maturity and is opened in response to the appropriate environmental signal. • Breaking physical dormancy involves the disruption of these specialised structures within the seed, and acts as an environmental signal detector for germination.
  • 6. • For example,In PY-dormant Fabaceae species (Rosids) the lens (strophiole) is the spezialized structure localized in the hilum region that controls water-impermeability. • In PY-dormant Anacardiaceae species (Rosids), e.g. Rhus spp., the endocarp consists of three water-impermeable palisade layers (macrosclereids, osteosclereis, brachysclereids) and the outer crystalliferous layer. • Different types of chalazal plugs/caps are the specialized structures that control water-impermeability of other seeds, e.g. in PY-dormant species of Bixaceae, Cistaceae, and Malvaceae. • Therefore, palisade layers in the fruit perform the functional role of preventing water uptake.
  • 7. TREATMENT TO BREAK PHYSICAL DORMANCY • In natural systems, physical dormancy is broken by several factors including high temperatures, fluctuating temperatures, fire, freezing/thawing, drying or passage through the digestive tracts of animals. • But the commonly followed method is SCARIFICATION • Acid • Mechanical • Physical treatment – hot water treatment
  • 8. SCARIFICATION • Any treatments may be physical or chemical that weakens or softens the seed coat is known as scarification. • This method is more applicable to Malvaceae and Leguminaceae group of seeds. ACID SCARIFICATION • By using concentrated H2SO4 @ 100ml/kg of seed for 2-3 minutes treatments dormancy can be overcome in the seeds. • The duration of treatment will vary and it depends on type and nature of seed coat. • E.g. Tree crops 1-3 hours, Rose seeds, treat the seed partially with acid and then given with warm stratification.
  • 9. MECHANICAL SCARIFICATION • Seeds are rubbed on a sand paper or with a help of mechanical scarifier or by puncturing on seed coat with the help of needle to enhance / increase the moisture absorption by seeds. • E.g. Bitter gourd for sand scarification, sand and seed 2:1 ratio should be followed. Rub against hard surface of seed for 5 to 10 minutes. HOT WATER TREATMENTS • It is effective in case of leguminous seeds. • The seeds should be soaked in boiled water for 1-5 minutes in 60-80 °C.