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Selecting Wine Grape Clones to 
Improve Yield and Quality 
Matthew Fidelibus 
Department of Viticulture and Enology 
University of California, Davis
Species, cultivars, and clones 
• California’s wine industry based on a 
single species of grape, Vitis vinifera L. 
• Most wines made from traditional (old) 
cultivated varieties of V. vinifera 
• Old cultivars have accumulated many 
‘clones’ due to mistaken identity, mutation, 
or both 
• Variation among clones may be exploited 
to improve grapevine yield and quality
What is a grapevine clone? 
• A population of vines propagated 
asexually (e.g., by cuttings) from a single 
mother vine. 
• Virtually all propagated grapevines are 
technically ‘clones’ 
• Of particular interest are clones or 
‘selections’ having distinctive, beneficial, 
characteristics
How to make a mammalian clone
How to make a grapevine clone
How can clonal selection 
research benefit growers? 
• Nurseries offer many different clones of a 
given cultivar 
• These may differ dramatically with respect to 
yield, yield components, fruit composition, 
and susceptibility to pests and diseases 
• The performance of a given clone can 
depend on the climatic region where it is 
grown
Materials and methods 
• Chardonnay, Merlot, and 
Zinfandel/Primitivo vines planted in 1997 
• Bilateral cordons, two-wire trellis, spur 
pruned 
• Vine x row spacing = 7.5’ x 10’ 
• Ten single-vine replicates 
• Randomized complete block design 
• Within a year, all clones harvested on the 
same day
Chardonnay characteristics 
• Most widely cultivated wine grape in CA 
• > 70 different registered clones in CA 
• Clone 4, industry standard 
• High yields, good fruit composition 
• 4 has large, heavy clusters—susceptible 
to sour rot
Chardonnay clones 
Clone Source 
4 Martini 5V21, Olmo #66 
6 Martini 3V4, Olmo #68 
15 Prosser, WA, LR2V6 
18 Rauscedo 8, Italy 
20 Cornegliano 7, Italy 
37 France, 95
Chardonnay yield components 
Clone Clusters / vine Cluster wt. 
(kg.) 
Yield / vine 
4 78 d 0.29 a 21.9 a 
6 107 a 0.22 c 22.0 a 
15 107 a 0.20 d 19.9 b 
18 93 c 0.26 b 22.6 a 
20 98 bc 0.25 b 22.1 a 
37 105 ab 0.24 bc 23.2 a
Chardonnay berries 
Clone Berries / cluster Berry wt 
(g) 
4 192 a 1.5 b 
6 147 cd 1.5 b 
15 138 d 1.4 c 
18 172 b 1.5 b 
20 153 c 1.6 a 
37 158 c 1.5 b
Chardonnay fruit composition 
Clone Brix pH TA 
(g/100 ml) 
4 23.7 a 3.60 c 0.584 a 
6 22.9 b 3.70 a 0.555 b 
15 22.8 b 3.69 ab 0.581 a 
18 22.7 bc 3.67 b 0.534 c 
20 22.4 c 3.68 ab 0.555 b 
37 23.0 b 3.68 ab 0.534 c
Chardonnay, sour rot incidence 
Clone Sour rot incidence 
(%) 
4 16 a 
6 10 b 
15 4 c 
18 14 ab 
20 13 ab 
37 15 a
Chardonnay clones summary 
• Clone 4 had good yields, large clusters, 
and good fruit composition but was very 
susceptible to sour rot 
• Clone 15 had 10% lower yields than 4, but 
had much less sour rot than 4
Merlot characteristics 
• Merlot among the most important red wine 
grapes in Calif. 
• 20%, by area, and 45%, by yield, grown in 
Central Valley 
• Clone 3 is standard; consistent set, yield, 
fruit composition 
• Relative performance compared to other 
clones in the valley has not been tested
Merlot clones 
Clone Source 
1 Inglenook, Calif. 
3 Inglenook, Calif. 
9 Rauscedo, Italy #3 
10 Cornegliano, Italy ISV-V-F2 
11 Cornegliano, Italy ISV-V-F5 
14 Chamber d’Agriculture de al 
Gironde, France
Merlot yield components 
Clone Clusters / vine 
(no.) 
Cluster wt. 
(kg) 
Yield / vine 
(kg) 
1 90 b 0.24 19.83 c 
3 91 b 0.25 20.75 bc 
9 90 b 0.25 21.50 ab 
10 103 a 0.24 22.21 a 
11 92 b 0.26 21.08 abc 
14 85 b 0.23 18.11 d
Merlot berries 
Clone Berries / cluster 
(no.) 
Berry wt 
(g) 
1 150 1.57 c 
3 148 1.66 b 
9 149 1.65 b 
10 145 1.57 c 
11 145 1.73 a 
14 146 1.54 c
Merlot fruit composition 
Clone Brix pH TA 
(g/100 ml) 
1 23.6 c 3.91 b 0.377 cd 
3 23.4 c 3.90 bc 0.374 d 
9 23.7 c 3.90 bc 0.373 d 
10 23.65 bc 3.87 c 0.398 a 
11 24.0 b 3.94 a 0.391 ab 
14 24.4 a 3.94 a 0.385 bc
Merlot clones summary 
• Selection 10 performed best; consistently 
higher yields of fruit with low pH and high 
TA 
• Selection 11 was undesirable; the largest 
berries, was the most susceptible to rot, 
and had high pH 
• Selection 3 performed similar to selections 
1 and 9 and was not superior to 10
‘Crljenak kastelanski’ 
• “The red grape of Kastela” 
• Zinfandel and Primitivo are probably 
selections of the “Red grape” 
• Origin of Zinfandel is obscure, 1850s? 
• Primitivo means “early” in Italian 
• Primitivo has been grown in Italy for about 
200 years
Zinfandel characteristics 
• Big berries, large clusters, high 
fruitfulness, high yields 
• Very susceptible to “sour rot” in warm 
climates 
• Uneven ripening 
• Yield declines as the season progresses 
• Color can be poor, especially in warm 
climates
Zinfandel & Primitivo Clones 
Clone Source 
Zinfandel 1A Lodi, CA, Handel 
Zinfandel 2 Lodi, CA, Handel 
Zinfandel 3 Livermore, CA, Ruetz 
Primitivo 3 Bari, Italy 
Primitivo 5 Calo, Italy 
Primitivo 6 Calo, Italy
Zinfandel, cluster characteristics 
Clone Clusters / vine 
(no.) 
Cluster wt. 
(kg) 
Yield / vine 
(kg) 
1A 73 b 0.30 20.4 c 
2 75 b 0.31 21.3 bc 
3 75 b 0.35 23.4 b 
P3 91 a 0.32 26.0 a 
P5 79 b 0.30 22.2 bc 
P6 80 b 0.31 22.1 bc
Zinfandel, berry characteristics 
Clone Berries / 
cluster 
Berry wt 
(g) 
1A 104 2.77 a 
2 107 2.75 a 
3 126 2.76 a 
P3 116 2.64 b 
P5 117 2.57 b 
P6 110 2.78 a
Zinfandel, fruit composition 
Clone Brix pH TA 
(g/100 ml) 
1A 20.05 c 3.47 b 0.60 
2 20.16 c 3.45 b 0.60 
3 19.56 d 3.45 b 0.59 
P3 21.10 b 3.46 b 0.61 
P5 20.82 b 3.48 b 0.62 
P6 22.00 a 3.52 a 0.60
Zinfandel, sour rot susceptibility 
Clone Sour rot incidence 
(%) 
1A 50 ab 
2 55 a 
3 52 ab 
P3 40 cd 
P5 44 bc 
P6 34 d
Zinfandel clones summary 
• Few differences between the Zinfandel 
selections tested 
• Primitivo selections generally were better 
• Earlier maturing 
• Similar or higher yield 
• Similar or less sour rot
Acknowledgements 
• Thanks to L. Peter Christensen, Kimberley 
Cathline, Jorge Aguilar Osorio, Don 
Katayama, & Karin Kawagoe 
• Viticulture Consortium & American 
Vineyard Foundation
For more details… 
• Email mwf@uckac.edu

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Winegrape cultivar and clone selection for warm climates

  • 1. Selecting Wine Grape Clones to Improve Yield and Quality Matthew Fidelibus Department of Viticulture and Enology University of California, Davis
  • 2. Species, cultivars, and clones • California’s wine industry based on a single species of grape, Vitis vinifera L. • Most wines made from traditional (old) cultivated varieties of V. vinifera • Old cultivars have accumulated many ‘clones’ due to mistaken identity, mutation, or both • Variation among clones may be exploited to improve grapevine yield and quality
  • 3. What is a grapevine clone? • A population of vines propagated asexually (e.g., by cuttings) from a single mother vine. • Virtually all propagated grapevines are technically ‘clones’ • Of particular interest are clones or ‘selections’ having distinctive, beneficial, characteristics
  • 4. How to make a mammalian clone
  • 5. How to make a grapevine clone
  • 6.
  • 7. How can clonal selection research benefit growers? • Nurseries offer many different clones of a given cultivar • These may differ dramatically with respect to yield, yield components, fruit composition, and susceptibility to pests and diseases • The performance of a given clone can depend on the climatic region where it is grown
  • 8. Materials and methods • Chardonnay, Merlot, and Zinfandel/Primitivo vines planted in 1997 • Bilateral cordons, two-wire trellis, spur pruned • Vine x row spacing = 7.5’ x 10’ • Ten single-vine replicates • Randomized complete block design • Within a year, all clones harvested on the same day
  • 9. Chardonnay characteristics • Most widely cultivated wine grape in CA • > 70 different registered clones in CA • Clone 4, industry standard • High yields, good fruit composition • 4 has large, heavy clusters—susceptible to sour rot
  • 10. Chardonnay clones Clone Source 4 Martini 5V21, Olmo #66 6 Martini 3V4, Olmo #68 15 Prosser, WA, LR2V6 18 Rauscedo 8, Italy 20 Cornegliano 7, Italy 37 France, 95
  • 11. Chardonnay yield components Clone Clusters / vine Cluster wt. (kg.) Yield / vine 4 78 d 0.29 a 21.9 a 6 107 a 0.22 c 22.0 a 15 107 a 0.20 d 19.9 b 18 93 c 0.26 b 22.6 a 20 98 bc 0.25 b 22.1 a 37 105 ab 0.24 bc 23.2 a
  • 12. Chardonnay berries Clone Berries / cluster Berry wt (g) 4 192 a 1.5 b 6 147 cd 1.5 b 15 138 d 1.4 c 18 172 b 1.5 b 20 153 c 1.6 a 37 158 c 1.5 b
  • 13. Chardonnay fruit composition Clone Brix pH TA (g/100 ml) 4 23.7 a 3.60 c 0.584 a 6 22.9 b 3.70 a 0.555 b 15 22.8 b 3.69 ab 0.581 a 18 22.7 bc 3.67 b 0.534 c 20 22.4 c 3.68 ab 0.555 b 37 23.0 b 3.68 ab 0.534 c
  • 14. Chardonnay, sour rot incidence Clone Sour rot incidence (%) 4 16 a 6 10 b 15 4 c 18 14 ab 20 13 ab 37 15 a
  • 15. Chardonnay clones summary • Clone 4 had good yields, large clusters, and good fruit composition but was very susceptible to sour rot • Clone 15 had 10% lower yields than 4, but had much less sour rot than 4
  • 16. Merlot characteristics • Merlot among the most important red wine grapes in Calif. • 20%, by area, and 45%, by yield, grown in Central Valley • Clone 3 is standard; consistent set, yield, fruit composition • Relative performance compared to other clones in the valley has not been tested
  • 17. Merlot clones Clone Source 1 Inglenook, Calif. 3 Inglenook, Calif. 9 Rauscedo, Italy #3 10 Cornegliano, Italy ISV-V-F2 11 Cornegliano, Italy ISV-V-F5 14 Chamber d’Agriculture de al Gironde, France
  • 18. Merlot yield components Clone Clusters / vine (no.) Cluster wt. (kg) Yield / vine (kg) 1 90 b 0.24 19.83 c 3 91 b 0.25 20.75 bc 9 90 b 0.25 21.50 ab 10 103 a 0.24 22.21 a 11 92 b 0.26 21.08 abc 14 85 b 0.23 18.11 d
  • 19. Merlot berries Clone Berries / cluster (no.) Berry wt (g) 1 150 1.57 c 3 148 1.66 b 9 149 1.65 b 10 145 1.57 c 11 145 1.73 a 14 146 1.54 c
  • 20. Merlot fruit composition Clone Brix pH TA (g/100 ml) 1 23.6 c 3.91 b 0.377 cd 3 23.4 c 3.90 bc 0.374 d 9 23.7 c 3.90 bc 0.373 d 10 23.65 bc 3.87 c 0.398 a 11 24.0 b 3.94 a 0.391 ab 14 24.4 a 3.94 a 0.385 bc
  • 21. Merlot clones summary • Selection 10 performed best; consistently higher yields of fruit with low pH and high TA • Selection 11 was undesirable; the largest berries, was the most susceptible to rot, and had high pH • Selection 3 performed similar to selections 1 and 9 and was not superior to 10
  • 22. ‘Crljenak kastelanski’ • “The red grape of Kastela” • Zinfandel and Primitivo are probably selections of the “Red grape” • Origin of Zinfandel is obscure, 1850s? • Primitivo means “early” in Italian • Primitivo has been grown in Italy for about 200 years
  • 23. Zinfandel characteristics • Big berries, large clusters, high fruitfulness, high yields • Very susceptible to “sour rot” in warm climates • Uneven ripening • Yield declines as the season progresses • Color can be poor, especially in warm climates
  • 24.
  • 25. Zinfandel & Primitivo Clones Clone Source Zinfandel 1A Lodi, CA, Handel Zinfandel 2 Lodi, CA, Handel Zinfandel 3 Livermore, CA, Ruetz Primitivo 3 Bari, Italy Primitivo 5 Calo, Italy Primitivo 6 Calo, Italy
  • 26. Zinfandel, cluster characteristics Clone Clusters / vine (no.) Cluster wt. (kg) Yield / vine (kg) 1A 73 b 0.30 20.4 c 2 75 b 0.31 21.3 bc 3 75 b 0.35 23.4 b P3 91 a 0.32 26.0 a P5 79 b 0.30 22.2 bc P6 80 b 0.31 22.1 bc
  • 27. Zinfandel, berry characteristics Clone Berries / cluster Berry wt (g) 1A 104 2.77 a 2 107 2.75 a 3 126 2.76 a P3 116 2.64 b P5 117 2.57 b P6 110 2.78 a
  • 28. Zinfandel, fruit composition Clone Brix pH TA (g/100 ml) 1A 20.05 c 3.47 b 0.60 2 20.16 c 3.45 b 0.60 3 19.56 d 3.45 b 0.59 P3 21.10 b 3.46 b 0.61 P5 20.82 b 3.48 b 0.62 P6 22.00 a 3.52 a 0.60
  • 29. Zinfandel, sour rot susceptibility Clone Sour rot incidence (%) 1A 50 ab 2 55 a 3 52 ab P3 40 cd P5 44 bc P6 34 d
  • 30. Zinfandel clones summary • Few differences between the Zinfandel selections tested • Primitivo selections generally were better • Earlier maturing • Similar or higher yield • Similar or less sour rot
  • 31. Acknowledgements • Thanks to L. Peter Christensen, Kimberley Cathline, Jorge Aguilar Osorio, Don Katayama, & Karin Kawagoe • Viticulture Consortium & American Vineyard Foundation
  • 32. For more details… • Email mwf@uckac.edu