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Top 11 Indiana Invasive Plant Species
A partnership between the Center for Earth and Environmental Science at IUPUI and Indy Parks & Recreation
Throughout this compendium there are numerous references to vocabulary
that may be unfamiliar to those outside the lens of environmental work. Here,
you will find explanations of this information.
Native plant species (referred to as ‘natives’) are those that historically grew
in a location before an external force, namely human colonizers, arrived in
the area.
Non-native plant species are the opposite, they historically grew in areas
other than where they have been recognized as non-native. They were later
brought to the new area and escaped into the wild, either accidentally or on
purpose. These species are not necessarily dangerous to natives, or invasive,
and may even require human aid to propagate.
Naturalized plant species are non-natives that have begun to flourish without
the aid of humans. These plants are not necessarily invasive but invasive
species are a subcategory of these plants.
Invasive plant species (referred to as ‘invasives’) are a subcategory of
naturalize plants that have become a threat to natives. These dangers are
numerous and far reaching, the specifics of which will be explained in more
detail later.
Many terms exist to describe the growth durations, or time to complete the
plant’s biological life cycle, of plants. There are also terms that relate to
herbicides. The terms used in this compendium are as follows:
Annual: Plant completes its life cycle, from germination to the production of seeds, within one year, and then
dies.
Biennial: Flowering plant that takes two years to complete its biological lifecycle. In the first year, the plant
grows leaves, stems, and roots (vegetative structures), then it enters a period of dormancy over the colder
months.
Perennial: Flowering plants, that grow and bloom over the spring and summer, die back every autumn and
winter, and then return in the spring from their rootstock.
Basal spray: Apply herbicide to the base of the main trunk of the plant.
Penetrant: A chemical that increases the ability of a poison to apply its toxic effect to a living organism.
Surfactant: A compound that lowers the surface tension (or interfacial tension) between two liquids, between a
gas and a liquid or between a liquid and a solid.
Invasive plants have many traits that allow them to flourish as natives in their
country or area of origin. These same traits, when used in a new area of the
world, often give them an unreasonable advantage over their native
counterparts.
Many invasives will have a different growing season than the surrounding plants,
most often their growing season allows them to begin presenting leaves, flowers,
and fruit weeks or months before their native counterparts will. If the invasive
plant is able to establish itself so early in the growing season it can easily starve
a native plant of sunlight and space for growth. This creates a vicious cycle of
aggressive growth by the native as the natives surrounding it die off.
Native animals, such as nesting birds, are also affected. If these bird species
have always sought out the branches or leaves of a native plant that has been
out-competed, and subsequently killed off, by an invasive plant it may force the
birds to nest elsewhere or use sub-par nesting materials.
Insects are another cause for concern as they may rely on the leaves of native
plants for their normal diet. If these leaves cannot be found then the insects in
the area will die off. This trend continues up the food chain and may cause a
domino effect of death or exodus by native animal species.
Various herbicides are described throughout this compendium, the specifics of
use will be described on the slides pertaining to particular species. These
chemicals are often effective tools for controlling the spread of dangerous
invasive plants.
Extreme caution should be used when handling these chemicals. They are
chemical agents designed to destroy biological systems that are toxic, or fatal,
to non-target plants, animals, and humans if ingested or absorbed.
The herbicides that are advised for use in this compendium are listed below.
Names with a registered trademark symbol (®) are brand-name, others are
generic chemicals. All should be made into solutions, as directed, using water.
Arsenal AC ®
Clopyralid
Escort ®
Garlon 3A ®
Garlon 4 ®
Glyphosate
MCPA
Phenoxy
Roundup ®
Tordon 101 ®
Tordon K ®
Triclopyr
Vanquish ®
Some herbicides (such as Tordon 101 and Tordon K) are safer than others in
regards to aquatic life. Herbicides should be used as a last resort to remove
invasive plants but consideration should be given first to herbicides that have the
least impact on the surrounding environment.
Shrubs & Trees
Autumn Olive (Elaeagnus umbellata)
Burning Bush (Euonymus alatus)
Bush Honeysuckle (Lonicera maackii)
Callery Pear (Pyrus calleryana)
Japanese Barberry (Berberis thunbergii)
Multiflora Rose (Rosa multiflora)
Other names include: Japanese Silverberry
Duration: Perennial
Family: Elaegnaceae
Height: 1 – 6 m (3 – 20 ft) tall
Foliage
Leaves: 5 – 8 cm (2 – 3 in) long and 2 – 3 cm
(0.8 – 1.2 in) wide
• Alternate pattern
• Elliptic
• Bright green to gray green
• Silvery scaly midvein
• Silver scaly beneath
• Petioles short and silvery
Thorns: Formed from twigs
Flowers: 8-10 mm (0.3-0.4 in) broad
• Fragrant
• Clusters of white to yellow
• 1-7 per cluster
• 4 lobed
Fruit (not pictured) : Red and finely doted with
silvery to silvery-brown scales, 7 – 10 mm (0.3 –
0.4 in) diameter
• August - November
• Pulpy, juicy, sweet
• Thin skin covering
• Drupe
• Has a pit with fleshy coating
Native to: China & Japan
Introduction (1830): Wildlife
habitat, strip mine reclamation,
shelterbelts (a barrier of plant
life that protects against wind
and reduces erosion)
Threat
• Invades old fields, woodland
edges, and other disturbed
areas
• Can form dense shrub layer
• Displaces native species and
closes open areas
Mechanical Controls
Pull sprouts when possible.
Cutting or burning of the
branches is heavily discouraged
as it can further the spread of
the plant.
Herbicidal Controls
Foliar Spray Method: Wet leaves
with Arsenal AC or Vanquish as a
1% solution in water (4 ounces
of herbicide per 3 gallon mix),
use with a surfactant. Use
between the months of April to
October.
For stems too tall for foliar
sprays, apply Garlon 4 as a 20%
solution in basal oil, diesel fuel,
or kerosene (2.5 quarts per 3-
gallon mix) with a penetrant
(check with herbicide
distributor) to young bark as a
basal spray (January – February
or May - October)
Cut Stump Method: Cut large
stems and immediately treat the
stumps with one of the following
herbicides in water with a
surfactant:
Arsenal AC: 10% solution (1
quart per 3-gallon mix)
Glyphosate: 20% solution (2.5
quarts per 3-gallon mix)
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Other names include: Winged burning bush or winged
spindletree
Duration: Perennial
Family: Celastraceae
Height: 6.1m (20 ft)
Foliage
Leaves: 2.5 – 7.62 cm (1 – 3 in) long and 1 – 4 cm
(0.39 – 1.57 in) wide
• Opposite branching
• Elliptic
• Serrated
Stems/bark: 4 corky ridges (wings)
Flowers: Not very distinctive
• Late May - June
Fruit (not pictured): Red, 15.2 cm (6 in) capsule
• Enclosed in 4-lobed pink/yellow or orange
capsule
• Ripens in September
Native to: China, Japan, Korea,
Russia
Introduction: Ornamental
Threat
• Lots of seeds
• Bird (and other berry-feeding
animals) dispersal
• Self-fertilizing
• Stump sprouting
• No native predators
• Long growth season
• Shade/sun tolerant
Mechanical Controls
Hand Pull: Can be done at any
time during the year. Pull, dig
and cut the plant out of the
ground. Repeated cutting on a
monthly cycle can inhibit
fruit/seed production and stunt
growth.
Complete removal of root mass
is recommended as secured
roots can sprout new plants.
Herbicidal Controls
Herbicide application in July,
August, and up to mid-
September gives maximum
chemical control.
In March - June the stump
should be cut high (six to
twelve inches), allowed to
stump sprout, and cut again
from July – September to one
inch from the ground before
following cut stump method
instructions.
Cut Stump Method: Cut plant
down to one inch from the
ground.
Glyphosate: Immediately apply
herbicide (straight, no
solution) to stump using paint
brush or sponge applicator.
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Other names include: Amur Honeysuckle
Duration: Perennial
Family: Caprifoliaceae
Height: 6m (~19.5 ft)
Foliage
Leaves: 3.5 – 8.5 am (1.3 – 3.3 in) long
• Opposite pattern for leaves
• Ovate to lance-ovate
• Acuminate
• Tapering gradually to a sharp point, as the tips of certain leaves.
• Dark green above, lighter on lower surface
• Veins are pubescent
Flowers: 1.5 – 2 cm (0.6 – 0.75 in) long
• February - June
• White/yellowish
• Bilabiate
• Having two lips, as the corollas of the
snapdragon.
• 5 parted
• Top 4 petals are fused
Fruit: 6 – 12 mm (0.2 – 0.5 in) in diameter, Green
becoming pink, ripen to red (sometimes yellow or
orange)
• August – February
• Glassy
• In leaf axils
• Usually persist into winter
Native to: Asia
Introduction (1700s – 1800s):
Ornamental
Threat
• Early leaf out in spring
• Takes up space for and shades
out competitors
• Berries available for birds to
eat/disperse even into winter
• Shallow roots do not hold soil
in place
• Reduces surrounding water
quality
Mechanical Controls
Hand Pull: Pulling out small
plants can remove entire root
mass with ease. Important as
secured roots can sprout new
plants.
Can be seasonally controlled by
cutting the main stem of the
plant one inch from the ground.
Herbicidal Controls
Foliar Spray Method:
Thoroughly wetting all leaves is
recommended.
Glyphosate: Use as a 2%
solution (8 ounces per 3-
gallon mix) of glyphosate and
water with a surfactant
Garlon 4: Use as a 20%
solution in basal oil, diesel
fuel, or kerosene (2.5 quarts
per 3-gallon mix), where
commercially available, with
a penetrant (when advised by
herbicide distributor) to
young bark at base of young
plant.
Cut Stump Method:
Recommended for stems too tall
for foliar spray. Use following
herbicides in water with a
surfactant.
Arsenal AC: 10% solution (1
quart per 3-gallon mix).
Glyphosate: 20% solution (2.5
quarts per 3-gallon mix).
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Other names include: Bradford Pear
Duration: Perennial
Family: Rosaceae
Height: 18 m (60 ft) tall, 0.6 m (2ft) diameter
Foliage
Leaves: 5.1-7.6 cm (2-3 in) long
• Alternate
• Simple
• Petiolate
• Shiny
• Wavy, slightly toothed margins
Flowers: 2.5 cm (1in) wide
• April – May
• Emerge before leaves
• White
• Insect pollinated
Fruit (not pictured): 1.3 cm (0.5 in) diameter,
green – brown
Native to: China and Vietnam
Introduction (1900s): Ornamental
Threat
• Usually produces sterile fruit
but can hybridize with other
pears and produce fertile fruit
• Can establish dominance in an
area quickly
Mechanical Controls
Do not plant Callery Pears or
any cultivars such as the
Bradford Pear.
Seedlings and shallow rooted
plants can be pulled if soil is
moist.
Herbicidal Controls
Medium to large trees should be
cut down and stumps treated
with glyphosate or triclopyr-
based herbicide.
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Other names include: Red Barberry
Duration: Perennial
Family: Berberidaceae
Height: 0.6 – 2.5 m (2 – 8.2 ft)
Foliage
Leaves: 1 – 3 cm (0.4 – 1.2 in) long
• Green, blue-green
• May be reddish or purple
• Oval shaped
• Semi – evergreen: alternate or grow in
alternate clusters
Branches: Deeply grooved, spiny
Flowers: 8-10 mm (0.3-0.4 in) broad
• Pale yellow
• Clusters of 2 – 5
• Bloom in March – April
Fruit (not pictured) : Bright red to orange-red berries
• Single seeded
Native to: Japan
Introduction: Ornamental
Threat
• Similar to Japanese
honeysuckle
o Shade tolerant
o Forms dense clumps
• Wide range of
environments
o Closed canopy forests
o Woodlands
o Wetlands
o Pastures
o Meadows
o Wasteland
• Easily dispersed by birds
Mechanical Controls
Hand Pull: Effective method for
small populations of the plant.
Separating the roots from the
plant is also effective as the
plant cannot regrow from its
root mass alone. However,
pulled plants with fruit present
should be removed from the
area to minimize seed dispersal.
Mowing/Cutting: This method
should be used where herbicides
cannot and should be repeated
every growing season as this is
only a temporary solution.
Herbicidal Controls
Foliar Spray Method: Should be
used for large thickets of the
plant when risk to non-target
plants is minimal.
Glyphosate: Use as a 2%
solution of glyphosate and
water plus a 0.5% non-ionic
surfactant to thoroughly wet all
leaves.
Triclopyr: Use as a 2% solution
of triclopyr and water plus a
0.5% non-ionic surfactant to
thoroughly wet all leaves.
Cut Stump Method: This
method should be considered
when treating individual bushes
or where the presence of
desirable species precludes
foliar application. Stump
treatments can be used as long
as the ground is not frozen.
Glyphosate: Horizontally cut
barberry stems at or near
ground level. Immediately
apply a 25% solution of
glyphosate and water to the
cut stump, covering the outer
20% of the stump.
Triclopyr: Horizontally cut
barberry stems at or near
ground level. Immediately
apply a 25% solution of
triclopyr and water to the cut
stump, covering the outer 20%
of the stump.
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Other names include: Japanese rose, baby rose,
seven-sisters rose, rambler rose
Duration: Perennial
Family: Rosaceae
Height: 3-5 m (10 – 16.5 ft)
Foliage
Leaves: 2.5 – 3.8 cm (1 – 1.5 in) long
• Serrated
• Fringed petioles
• 7 – 9 leaflets
Prickles: Not always present
Flowers: Large corymbs
• Small
• White to pinkish
• 5 petaled
• Clustered
Fruit (not pictured): 6-8 mm diameter, reddish
to purple
Native to: Japan, Korea
Introduction (1800s):
Ornamental, livestock
containment, wildlife habitat
Threat
• Displaces native vegetation
• Creates natural barrier to
wildlife
• Bird dispersed
• Tolerant of many conditions
Mechanical Controls
Hand Pull: Pull sprout to
remove root mass.
Cut stems, repeat to stunt
growth.
Herbicidal Controls
Cut Stump Method: Apply to cut
stumps if necessary
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Woody Vines
Japanese Honeysuckle (Lonicera japonica)
Oriental Bittersweet (Celastrus orbiculatus)
Winter Creeper (Euonymus fortunei)
Other names include: Chinese honeysuckle
Duration: Perennial
Family: Caprifoliaceae
Height: 24 m (80 ft)
Foliage
Leaves: 4 – 6.5 am (1.6 – 2.6 in) long, 2-4
cm (0.8-1.5 in) wide
• Opposite branching
• Ovate to elliptic to oblong
• Both surfaces smooth – rough hairy with
white underside
Branches: Deeply grooved, spiny
Flowers: 2 – 3 cm (0.8 – 1.2 in) long, White-pink
and pale yellow
• April – August
• Axillary pairs
• 5 lobes, 2 lips
• 5 stamens, 1 pistil
Fruit (not pictured): Green ripening to black, 5 –
6 mm (0.2 in) diameter, Stalks: 1 – 3 cm (0.4 –
1.2 in)
• June – March
• 2-3 seeds
Native to: Japan
Introduction (1800): Ornamental,
deer browse
Threat
• Forms dense mats in tree canopies
• Useful for erosion control which
leading to more planting
Mechanical Controls
Controlled burning in spring can
reduce ground mats.
Herbicidal Controls
Apply following herbicides to
foliage with a surfactant from
July – October.
Glyphosate: Use as a 2%
solution (8 ounces per 3-
gallon mix) of glyphosate
and water.
Garlon 3A or 4: Use as a
3% - 5% solution (12 – 20
ounces per 3-gallon mix.
Apply Escort to foliage with a
surfactant from June – August 2
ounces per acre in water (0.6
dry ounces per 3-gallon mix) or
spot spraying 2 – 4 ounces per
acre in water (0.6 – 1.2 dry
ounces per 3-gallon mix).
Cut Stump Method: This
method should be used for
severed vines.
Glyphosate or Garlon 3A: Use
as a 20% solution (2.5 quarts
per 3-gallon mix) in water with
surfactant from July - October.
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Other names include: Chinese bittersweet
Duration: Perennial
Family: Celastraceae
Height: 18.3 m (60 ft) tall, 10 cm (4 in) diameter
Foliage
Leaves: 5-13 cm (2-5 in) long
• Alternate
• Elliptical-circular
Flowers: Axillary
• Greenish-white
• Bloom from May – early June
Fruit (not pictured): Green when young, ripen to
yellow then split to reveal scarlet berries which
persist into winter
Native to: Asia
Introduction (1700s – 1800s):
Ornamental
Threat
• Fast growing: covers, shades,
and outcompetes natives
• Can kill large trees
• Seed distributed
Mechanical Controls
Hand Pull: Removal of small or
young plants is advised.
Herbicidal Controls
Foliar Spray Method: Apply one
of the following herbicides with
a surfactant from July –
October.
Garlon 3A or 4 or glyphosate:
Use as a 2% solution (8 ounces
per 3-gallon mix).
Cut Stump Method: For stems
too tall for foliar sprays, apply
Garlon 4 as a 20% solution in
basal oil, diesel fuel, or
kerosene (2.5 quarts per 3-
gallon mix) with a penetrant
(check with herbicide
distributor).
Or, cut large stems and
immediately treat with one of
the following with a surfactant:
Garlon 4 or glyphosate:
25% solution (32 ounces
per 1-gallon mix)
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Other names include: Climbing euonymus, spindle,
fortune’s spindle
Duration: Perennial
Family: Celastraceae
Height: 20 m (66 ft)
Foliage
Leaves: 1-2.5 in (2.5-6.4 cm) long
• Opposite
• Glossy
• Dark green
• Oval
• Slightly toothed
• Light colored veins
Flowers: 5 petals, long branched stalks
Fruit (not pictured) : Pink - red
• Split open to expose seeds with red-orange
arils
Native to: China, Japan, Korea
Introduction: Ornamental
groundcover
Threat
• Takes over open areas
• Grows across ground, displaces
natives and kills seedlings
• Tolerates wide range of soil
types
Mechanical Controls
Cutting: Sever all vertical
climbing stems to prevent
fruiting and spread by birds.
Herbicidal Controls
Foliar Spray Method: Wet leaves
until runoff with one of the
following:
Tordon 101: Use as a 3%
solution (12 ounces per 3-
gallon mix).
Tordon K: Use as a 2% solution
(8 ounces per 3-gallon mix).
Or repeatedly apply one of the
following (less effective but
reduces harm to non-target
plants):
Garlon 4 or glyphosate: Use as
a 2% solution (8 ounces per 3-
gallon mix) in water with a
surfactant.
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
Herbaceous plants
Canada Thistle (Cirsium arvense)
Garlic Mustard (Alliaria petiolata)
Other names include: Creeping thistle, bull thistle,
spear thistle, lettuce from hell thistle, cursed thistle
Duration: Perennial
Family: Asteraceae
Height: 0.3 – 1.5 m (1 – 5 ft)
Foliage
Leaves: 12 – 20 cm (5 – 8 in) long (basal)
• Lanceolate to oblong-lanceolate
• Glaborous above
• Undersides have short, white hairs
Flowers:
• Female: 1 – 1.5 cm (0.4 – 0.6 in) diameter, 1 – 2 cm
(0.4 – 0.75 in) tall
• Flask-shaped
• Fragrant
• Male
• Smaller
• More globose
Fruit (not pictured) : 2- 3 mm (0.1 in) long, 1 mm
(0.04 in) diameter
• White – light brown pappus
Native to: Europe, Northern Asia
Introduction (17th century):
Escaped weed
Threat
• Hard to remove
• Reduces species diversity
• Decreases crop yield
• Problem in prairie and riparian
habitats
Mechanical Controls
Cutting: At flower stem
extension before bud opens.
Predatory insects: Orellia
ruficauda and Aceria
anthocoptes
Competing plants: Growing
forages such as alfalfa which
must be cut frequently)
Herbicidal Controls
Phenoxy compounds
MCPA and clopyralid where
approved
Other names include: Hedge garlic, jack-by-the-hedge
garlic, poor man’s mustard, garlic root, mustard root,
garlicwort
Duration: Annual/biennial
Family: Brassicaceae
Height: 60 – 120 cm (2 – 4 ft) flower stalks
Foliage
Leaves: 3 – 9 cm (1.2 – 3.6 in) long, 2.5 – 10 cm (1
– 4 in) wide
• Rosette of kidney-shaped
• Later alternate heart-shaped – triangular
Petioles: 1 – 8 cm (0.4 – 3 in) long & reduced
upwards
Flowers: 5 – 7 mm (0.2 – 0.3 in) long, 10 – 14 mm (0.4 –
0.6 in) wide
• Terminal
• Tight clusters of small white, four-petaled flowers
• Bloom April – May
Fruit (not pictured): 2.5 – 12 cm (1 – 5 in) long, 1.5 mm
(0.06 in) wide, green ripening to tan & papery
• Four sided, thin pod
• Alternately whorled along stalk
• Explode to expel tiny black seeds arranged in rows
• Present May – June
Native to: Antarctic, Asia, China,
Europe, Morocco, Pakistan,
Scandinavia
Introduction (1800s): Medical and
culinary use, escaped weed
Threat
• High shade tolerance
• Can invade high quality, mature
woodlands
• Forms dense strands
Mechanical Controls
Hand Pull: Both young and
mature plants can be pulled
with ease. Should be done
before seed formation.
Repeated, annual burning in
early spring or fall can control
plant. Individual burning of
plants with propane torches has
shown some success.
Herbicidal Controls
Foliar Spray Method: Used to
control multiple generations of
the plant:
Glyphosate: Use as a 2%
solution in water (8 ounces
per 3-gallon mix) during
flowering (April – June) and
include a surfactant unless
plants are near surface
waters.
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southern Research Station. 93 p.
References
All images are courtesy of the Center for Earth and Environmental Science at
Indiana University – Purdue University Indianapolis unless otherwise noted.
Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide
for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S.
Department of Agriculture, Forest Service, Southern Research Station. 93 p.
https://en.wikipedia.org/wiki/Surfactant?oldformat=true
https://en.wikipedia.org/wiki/Penetrant_(biochemical)?oldformat=true
Plant references
Bush Honeysuckle (Lonicera maackii)
https://plants.usda.gov/core/profile?symbol=loma6
https://www.invasiveplantatlas.org/subject.html?sub=3040
https://www.invasive.org/eastern/srs/BH.html
Japanese Barberry (Berberis thunbergii)
https://plants.usda.gov/core/profile?symbol=beth
https://www.invasiveplantatlas.org/subject.html?sub=3010
https://wiki.bugwood.org/Archive:SEEPPC/Berberis_thunbergii
Burning Bush (Euonymus alatus)
https://plants.usda.gov/core/profile?symbol=EUAL13
http://www.hort.uconn.edu/plants/detail.php?pid=167
https://www.ecolandscaping.org/04/invasive-plants/winged-euonymus-an-exotic-invasive-plant-
fact-sheet/
Plant references continued
Autumn Olive (Elaeagnus umbellata)
https://plants.usda.gov/core/profile?symbol=ELUM
https://www.invasiveplantatlas.org/subject.html?sub=3021
https://www.invasive.org/eastern/srs/AO.html
Multiflora Rose (Rosa multiflora)
https://plants.usda.gov/core/profile?symbol=romu
https://www.invasiveplantatlas.org/subject.html?sub=3071
https://www.invasive.org/eastern/srs/ER.html
Winter Creeper (Euonymus fortunei)
https://www.invasiveplantatlas.org/subject.html?sub=3024
https://plants.usda.gov/core/profile?symbol=EUFO5
Plant references continued
Oriental Bittersweet (Celastrus orbiculatus)
https://www.invasiveplantatlas.org/subject.html?sub=3012
https://plants.usda.gov/core/profile?symbol=CEOR7
https://plants.usda.gov/java/charProfile?symbol=CEOR7
Japanese Honeysuckle (Lonicera japonica)
https://www.invasiveplantatlas.org/subject.html?sub=3039
https://plants.usda.gov/core/profile?symbol=LOJA
https://www.invasive.org/eastern/srs/JH.html
Callery Pear = Bradford Pear (Pyrus calleryana)
https://plants.usda.gov/core/profile?symbol=PYCA80
https://www.invasiveplantatlas.org/subject.html?sub=10957
https://www.invasive.org/alien/pubs/midatlantic/pyca.htm
Plant references continued
A Garlic Mustard (Alliaria petiolata formerly A. officinalis)
https://www.invasiveplantatlas.org/subject.html?sub=3005
https://www.invasive.org/eastern/srs/GM.html
https://plants.usda.gov/core/profile?symbol=alpe4

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Top 11 indiana invasive plant species

  • 1. Top 11 Indiana Invasive Plant Species A partnership between the Center for Earth and Environmental Science at IUPUI and Indy Parks & Recreation
  • 2. Throughout this compendium there are numerous references to vocabulary that may be unfamiliar to those outside the lens of environmental work. Here, you will find explanations of this information. Native plant species (referred to as ‘natives’) are those that historically grew in a location before an external force, namely human colonizers, arrived in the area. Non-native plant species are the opposite, they historically grew in areas other than where they have been recognized as non-native. They were later brought to the new area and escaped into the wild, either accidentally or on purpose. These species are not necessarily dangerous to natives, or invasive, and may even require human aid to propagate. Naturalized plant species are non-natives that have begun to flourish without the aid of humans. These plants are not necessarily invasive but invasive species are a subcategory of these plants. Invasive plant species (referred to as ‘invasives’) are a subcategory of naturalize plants that have become a threat to natives. These dangers are numerous and far reaching, the specifics of which will be explained in more detail later.
  • 3. Many terms exist to describe the growth durations, or time to complete the plant’s biological life cycle, of plants. There are also terms that relate to herbicides. The terms used in this compendium are as follows: Annual: Plant completes its life cycle, from germination to the production of seeds, within one year, and then dies. Biennial: Flowering plant that takes two years to complete its biological lifecycle. In the first year, the plant grows leaves, stems, and roots (vegetative structures), then it enters a period of dormancy over the colder months. Perennial: Flowering plants, that grow and bloom over the spring and summer, die back every autumn and winter, and then return in the spring from their rootstock. Basal spray: Apply herbicide to the base of the main trunk of the plant. Penetrant: A chemical that increases the ability of a poison to apply its toxic effect to a living organism. Surfactant: A compound that lowers the surface tension (or interfacial tension) between two liquids, between a gas and a liquid or between a liquid and a solid.
  • 4. Invasive plants have many traits that allow them to flourish as natives in their country or area of origin. These same traits, when used in a new area of the world, often give them an unreasonable advantage over their native counterparts. Many invasives will have a different growing season than the surrounding plants, most often their growing season allows them to begin presenting leaves, flowers, and fruit weeks or months before their native counterparts will. If the invasive plant is able to establish itself so early in the growing season it can easily starve a native plant of sunlight and space for growth. This creates a vicious cycle of aggressive growth by the native as the natives surrounding it die off. Native animals, such as nesting birds, are also affected. If these bird species have always sought out the branches or leaves of a native plant that has been out-competed, and subsequently killed off, by an invasive plant it may force the birds to nest elsewhere or use sub-par nesting materials. Insects are another cause for concern as they may rely on the leaves of native plants for their normal diet. If these leaves cannot be found then the insects in the area will die off. This trend continues up the food chain and may cause a domino effect of death or exodus by native animal species.
  • 5. Various herbicides are described throughout this compendium, the specifics of use will be described on the slides pertaining to particular species. These chemicals are often effective tools for controlling the spread of dangerous invasive plants. Extreme caution should be used when handling these chemicals. They are chemical agents designed to destroy biological systems that are toxic, or fatal, to non-target plants, animals, and humans if ingested or absorbed. The herbicides that are advised for use in this compendium are listed below. Names with a registered trademark symbol (®) are brand-name, others are generic chemicals. All should be made into solutions, as directed, using water. Arsenal AC ® Clopyralid Escort ® Garlon 3A ® Garlon 4 ® Glyphosate MCPA Phenoxy Roundup ® Tordon 101 ® Tordon K ® Triclopyr Vanquish ®
  • 6. Some herbicides (such as Tordon 101 and Tordon K) are safer than others in regards to aquatic life. Herbicides should be used as a last resort to remove invasive plants but consideration should be given first to herbicides that have the least impact on the surrounding environment.
  • 7. Shrubs & Trees Autumn Olive (Elaeagnus umbellata) Burning Bush (Euonymus alatus) Bush Honeysuckle (Lonicera maackii) Callery Pear (Pyrus calleryana) Japanese Barberry (Berberis thunbergii) Multiflora Rose (Rosa multiflora)
  • 8. Other names include: Japanese Silverberry Duration: Perennial Family: Elaegnaceae Height: 1 – 6 m (3 – 20 ft) tall Foliage Leaves: 5 – 8 cm (2 – 3 in) long and 2 – 3 cm (0.8 – 1.2 in) wide • Alternate pattern • Elliptic • Bright green to gray green • Silvery scaly midvein • Silver scaly beneath • Petioles short and silvery Thorns: Formed from twigs
  • 9. Flowers: 8-10 mm (0.3-0.4 in) broad • Fragrant • Clusters of white to yellow • 1-7 per cluster • 4 lobed Fruit (not pictured) : Red and finely doted with silvery to silvery-brown scales, 7 – 10 mm (0.3 – 0.4 in) diameter • August - November • Pulpy, juicy, sweet • Thin skin covering • Drupe • Has a pit with fleshy coating
  • 10. Native to: China & Japan Introduction (1830): Wildlife habitat, strip mine reclamation, shelterbelts (a barrier of plant life that protects against wind and reduces erosion) Threat • Invades old fields, woodland edges, and other disturbed areas • Can form dense shrub layer • Displaces native species and closes open areas
  • 11. Mechanical Controls Pull sprouts when possible. Cutting or burning of the branches is heavily discouraged as it can further the spread of the plant. Herbicidal Controls Foliar Spray Method: Wet leaves with Arsenal AC or Vanquish as a 1% solution in water (4 ounces of herbicide per 3 gallon mix), use with a surfactant. Use between the months of April to October. For stems too tall for foliar sprays, apply Garlon 4 as a 20% solution in basal oil, diesel fuel, or kerosene (2.5 quarts per 3- gallon mix) with a penetrant (check with herbicide distributor) to young bark as a basal spray (January – February or May - October) Cut Stump Method: Cut large stems and immediately treat the stumps with one of the following herbicides in water with a surfactant: Arsenal AC: 10% solution (1 quart per 3-gallon mix) Glyphosate: 20% solution (2.5 quarts per 3-gallon mix) Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 12. Other names include: Winged burning bush or winged spindletree Duration: Perennial Family: Celastraceae Height: 6.1m (20 ft) Foliage Leaves: 2.5 – 7.62 cm (1 – 3 in) long and 1 – 4 cm (0.39 – 1.57 in) wide • Opposite branching • Elliptic • Serrated Stems/bark: 4 corky ridges (wings)
  • 13. Flowers: Not very distinctive • Late May - June Fruit (not pictured): Red, 15.2 cm (6 in) capsule • Enclosed in 4-lobed pink/yellow or orange capsule • Ripens in September
  • 14. Native to: China, Japan, Korea, Russia Introduction: Ornamental Threat • Lots of seeds • Bird (and other berry-feeding animals) dispersal • Self-fertilizing • Stump sprouting • No native predators • Long growth season • Shade/sun tolerant
  • 15. Mechanical Controls Hand Pull: Can be done at any time during the year. Pull, dig and cut the plant out of the ground. Repeated cutting on a monthly cycle can inhibit fruit/seed production and stunt growth. Complete removal of root mass is recommended as secured roots can sprout new plants. Herbicidal Controls Herbicide application in July, August, and up to mid- September gives maximum chemical control. In March - June the stump should be cut high (six to twelve inches), allowed to stump sprout, and cut again from July – September to one inch from the ground before following cut stump method instructions. Cut Stump Method: Cut plant down to one inch from the ground. Glyphosate: Immediately apply herbicide (straight, no solution) to stump using paint brush or sponge applicator. Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 16. Other names include: Amur Honeysuckle Duration: Perennial Family: Caprifoliaceae Height: 6m (~19.5 ft) Foliage Leaves: 3.5 – 8.5 am (1.3 – 3.3 in) long • Opposite pattern for leaves • Ovate to lance-ovate • Acuminate • Tapering gradually to a sharp point, as the tips of certain leaves. • Dark green above, lighter on lower surface • Veins are pubescent
  • 17. Flowers: 1.5 – 2 cm (0.6 – 0.75 in) long • February - June • White/yellowish • Bilabiate • Having two lips, as the corollas of the snapdragon. • 5 parted • Top 4 petals are fused Fruit: 6 – 12 mm (0.2 – 0.5 in) in diameter, Green becoming pink, ripen to red (sometimes yellow or orange) • August – February • Glassy • In leaf axils • Usually persist into winter
  • 18. Native to: Asia Introduction (1700s – 1800s): Ornamental Threat • Early leaf out in spring • Takes up space for and shades out competitors • Berries available for birds to eat/disperse even into winter • Shallow roots do not hold soil in place • Reduces surrounding water quality
  • 19. Mechanical Controls Hand Pull: Pulling out small plants can remove entire root mass with ease. Important as secured roots can sprout new plants. Can be seasonally controlled by cutting the main stem of the plant one inch from the ground. Herbicidal Controls Foliar Spray Method: Thoroughly wetting all leaves is recommended. Glyphosate: Use as a 2% solution (8 ounces per 3- gallon mix) of glyphosate and water with a surfactant Garlon 4: Use as a 20% solution in basal oil, diesel fuel, or kerosene (2.5 quarts per 3-gallon mix), where commercially available, with a penetrant (when advised by herbicide distributor) to young bark at base of young plant. Cut Stump Method: Recommended for stems too tall for foliar spray. Use following herbicides in water with a surfactant. Arsenal AC: 10% solution (1 quart per 3-gallon mix). Glyphosate: 20% solution (2.5 quarts per 3-gallon mix). Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 20. Other names include: Bradford Pear Duration: Perennial Family: Rosaceae Height: 18 m (60 ft) tall, 0.6 m (2ft) diameter Foliage Leaves: 5.1-7.6 cm (2-3 in) long • Alternate • Simple • Petiolate • Shiny • Wavy, slightly toothed margins
  • 21. Flowers: 2.5 cm (1in) wide • April – May • Emerge before leaves • White • Insect pollinated Fruit (not pictured): 1.3 cm (0.5 in) diameter, green – brown
  • 22. Native to: China and Vietnam Introduction (1900s): Ornamental Threat • Usually produces sterile fruit but can hybridize with other pears and produce fertile fruit • Can establish dominance in an area quickly
  • 23. Mechanical Controls Do not plant Callery Pears or any cultivars such as the Bradford Pear. Seedlings and shallow rooted plants can be pulled if soil is moist. Herbicidal Controls Medium to large trees should be cut down and stumps treated with glyphosate or triclopyr- based herbicide. Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 24. Other names include: Red Barberry Duration: Perennial Family: Berberidaceae Height: 0.6 – 2.5 m (2 – 8.2 ft) Foliage Leaves: 1 – 3 cm (0.4 – 1.2 in) long • Green, blue-green • May be reddish or purple • Oval shaped • Semi – evergreen: alternate or grow in alternate clusters Branches: Deeply grooved, spiny
  • 25. Flowers: 8-10 mm (0.3-0.4 in) broad • Pale yellow • Clusters of 2 – 5 • Bloom in March – April Fruit (not pictured) : Bright red to orange-red berries • Single seeded
  • 26. Native to: Japan Introduction: Ornamental Threat • Similar to Japanese honeysuckle o Shade tolerant o Forms dense clumps • Wide range of environments o Closed canopy forests o Woodlands o Wetlands o Pastures o Meadows o Wasteland • Easily dispersed by birds
  • 27. Mechanical Controls Hand Pull: Effective method for small populations of the plant. Separating the roots from the plant is also effective as the plant cannot regrow from its root mass alone. However, pulled plants with fruit present should be removed from the area to minimize seed dispersal. Mowing/Cutting: This method should be used where herbicides cannot and should be repeated every growing season as this is only a temporary solution. Herbicidal Controls Foliar Spray Method: Should be used for large thickets of the plant when risk to non-target plants is minimal. Glyphosate: Use as a 2% solution of glyphosate and water plus a 0.5% non-ionic surfactant to thoroughly wet all leaves. Triclopyr: Use as a 2% solution of triclopyr and water plus a 0.5% non-ionic surfactant to thoroughly wet all leaves. Cut Stump Method: This method should be considered when treating individual bushes or where the presence of desirable species precludes foliar application. Stump treatments can be used as long as the ground is not frozen. Glyphosate: Horizontally cut barberry stems at or near ground level. Immediately apply a 25% solution of glyphosate and water to the cut stump, covering the outer 20% of the stump. Triclopyr: Horizontally cut barberry stems at or near ground level. Immediately apply a 25% solution of triclopyr and water to the cut stump, covering the outer 20% of the stump. Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 28. Other names include: Japanese rose, baby rose, seven-sisters rose, rambler rose Duration: Perennial Family: Rosaceae Height: 3-5 m (10 – 16.5 ft) Foliage Leaves: 2.5 – 3.8 cm (1 – 1.5 in) long • Serrated • Fringed petioles • 7 – 9 leaflets Prickles: Not always present
  • 29. Flowers: Large corymbs • Small • White to pinkish • 5 petaled • Clustered Fruit (not pictured): 6-8 mm diameter, reddish to purple
  • 30. Native to: Japan, Korea Introduction (1800s): Ornamental, livestock containment, wildlife habitat Threat • Displaces native vegetation • Creates natural barrier to wildlife • Bird dispersed • Tolerant of many conditions
  • 31. Mechanical Controls Hand Pull: Pull sprout to remove root mass. Cut stems, repeat to stunt growth. Herbicidal Controls Cut Stump Method: Apply to cut stumps if necessary Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 32. Woody Vines Japanese Honeysuckle (Lonicera japonica) Oriental Bittersweet (Celastrus orbiculatus) Winter Creeper (Euonymus fortunei)
  • 33. Other names include: Chinese honeysuckle Duration: Perennial Family: Caprifoliaceae Height: 24 m (80 ft) Foliage Leaves: 4 – 6.5 am (1.6 – 2.6 in) long, 2-4 cm (0.8-1.5 in) wide • Opposite branching • Ovate to elliptic to oblong • Both surfaces smooth – rough hairy with white underside Branches: Deeply grooved, spiny
  • 34. Flowers: 2 – 3 cm (0.8 – 1.2 in) long, White-pink and pale yellow • April – August • Axillary pairs • 5 lobes, 2 lips • 5 stamens, 1 pistil Fruit (not pictured): Green ripening to black, 5 – 6 mm (0.2 in) diameter, Stalks: 1 – 3 cm (0.4 – 1.2 in) • June – March • 2-3 seeds
  • 35. Native to: Japan Introduction (1800): Ornamental, deer browse Threat • Forms dense mats in tree canopies • Useful for erosion control which leading to more planting
  • 36. Mechanical Controls Controlled burning in spring can reduce ground mats. Herbicidal Controls Apply following herbicides to foliage with a surfactant from July – October. Glyphosate: Use as a 2% solution (8 ounces per 3- gallon mix) of glyphosate and water. Garlon 3A or 4: Use as a 3% - 5% solution (12 – 20 ounces per 3-gallon mix. Apply Escort to foliage with a surfactant from June – August 2 ounces per acre in water (0.6 dry ounces per 3-gallon mix) or spot spraying 2 – 4 ounces per acre in water (0.6 – 1.2 dry ounces per 3-gallon mix). Cut Stump Method: This method should be used for severed vines. Glyphosate or Garlon 3A: Use as a 20% solution (2.5 quarts per 3-gallon mix) in water with surfactant from July - October. Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 37. Other names include: Chinese bittersweet Duration: Perennial Family: Celastraceae Height: 18.3 m (60 ft) tall, 10 cm (4 in) diameter Foliage Leaves: 5-13 cm (2-5 in) long • Alternate • Elliptical-circular
  • 38. Flowers: Axillary • Greenish-white • Bloom from May – early June Fruit (not pictured): Green when young, ripen to yellow then split to reveal scarlet berries which persist into winter
  • 39. Native to: Asia Introduction (1700s – 1800s): Ornamental Threat • Fast growing: covers, shades, and outcompetes natives • Can kill large trees • Seed distributed
  • 40. Mechanical Controls Hand Pull: Removal of small or young plants is advised. Herbicidal Controls Foliar Spray Method: Apply one of the following herbicides with a surfactant from July – October. Garlon 3A or 4 or glyphosate: Use as a 2% solution (8 ounces per 3-gallon mix). Cut Stump Method: For stems too tall for foliar sprays, apply Garlon 4 as a 20% solution in basal oil, diesel fuel, or kerosene (2.5 quarts per 3- gallon mix) with a penetrant (check with herbicide distributor). Or, cut large stems and immediately treat with one of the following with a surfactant: Garlon 4 or glyphosate: 25% solution (32 ounces per 1-gallon mix) Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 41. Other names include: Climbing euonymus, spindle, fortune’s spindle Duration: Perennial Family: Celastraceae Height: 20 m (66 ft) Foliage Leaves: 1-2.5 in (2.5-6.4 cm) long • Opposite • Glossy • Dark green • Oval • Slightly toothed • Light colored veins
  • 42. Flowers: 5 petals, long branched stalks Fruit (not pictured) : Pink - red • Split open to expose seeds with red-orange arils
  • 43. Native to: China, Japan, Korea Introduction: Ornamental groundcover Threat • Takes over open areas • Grows across ground, displaces natives and kills seedlings • Tolerates wide range of soil types
  • 44. Mechanical Controls Cutting: Sever all vertical climbing stems to prevent fruiting and spread by birds. Herbicidal Controls Foliar Spray Method: Wet leaves until runoff with one of the following: Tordon 101: Use as a 3% solution (12 ounces per 3- gallon mix). Tordon K: Use as a 2% solution (8 ounces per 3-gallon mix). Or repeatedly apply one of the following (less effective but reduces harm to non-target plants): Garlon 4 or glyphosate: Use as a 2% solution (8 ounces per 3- gallon mix) in water with a surfactant. Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 45. Herbaceous plants Canada Thistle (Cirsium arvense) Garlic Mustard (Alliaria petiolata)
  • 46. Other names include: Creeping thistle, bull thistle, spear thistle, lettuce from hell thistle, cursed thistle Duration: Perennial Family: Asteraceae Height: 0.3 – 1.5 m (1 – 5 ft) Foliage Leaves: 12 – 20 cm (5 – 8 in) long (basal) • Lanceolate to oblong-lanceolate • Glaborous above • Undersides have short, white hairs
  • 47. Flowers: • Female: 1 – 1.5 cm (0.4 – 0.6 in) diameter, 1 – 2 cm (0.4 – 0.75 in) tall • Flask-shaped • Fragrant • Male • Smaller • More globose Fruit (not pictured) : 2- 3 mm (0.1 in) long, 1 mm (0.04 in) diameter • White – light brown pappus
  • 48. Native to: Europe, Northern Asia Introduction (17th century): Escaped weed Threat • Hard to remove • Reduces species diversity • Decreases crop yield • Problem in prairie and riparian habitats
  • 49. Mechanical Controls Cutting: At flower stem extension before bud opens. Predatory insects: Orellia ruficauda and Aceria anthocoptes Competing plants: Growing forages such as alfalfa which must be cut frequently) Herbicidal Controls Phenoxy compounds MCPA and clopyralid where approved
  • 50. Other names include: Hedge garlic, jack-by-the-hedge garlic, poor man’s mustard, garlic root, mustard root, garlicwort Duration: Annual/biennial Family: Brassicaceae Height: 60 – 120 cm (2 – 4 ft) flower stalks Foliage Leaves: 3 – 9 cm (1.2 – 3.6 in) long, 2.5 – 10 cm (1 – 4 in) wide • Rosette of kidney-shaped • Later alternate heart-shaped – triangular Petioles: 1 – 8 cm (0.4 – 3 in) long & reduced upwards
  • 51. Flowers: 5 – 7 mm (0.2 – 0.3 in) long, 10 – 14 mm (0.4 – 0.6 in) wide • Terminal • Tight clusters of small white, four-petaled flowers • Bloom April – May Fruit (not pictured): 2.5 – 12 cm (1 – 5 in) long, 1.5 mm (0.06 in) wide, green ripening to tan & papery • Four sided, thin pod • Alternately whorled along stalk • Explode to expel tiny black seeds arranged in rows • Present May – June
  • 52. Native to: Antarctic, Asia, China, Europe, Morocco, Pakistan, Scandinavia Introduction (1800s): Medical and culinary use, escaped weed Threat • High shade tolerance • Can invade high quality, mature woodlands • Forms dense strands
  • 53. Mechanical Controls Hand Pull: Both young and mature plants can be pulled with ease. Should be done before seed formation. Repeated, annual burning in early spring or fall can control plant. Individual burning of plants with propane torches has shown some success. Herbicidal Controls Foliar Spray Method: Used to control multiple generations of the plant: Glyphosate: Use as a 2% solution in water (8 ounces per 3-gallon mix) during flowering (April – June) and include a surfactant unless plants are near surface waters. Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p.
  • 54. References All images are courtesy of the Center for Earth and Environmental Science at Indiana University – Purdue University Indianapolis unless otherwise noted. Miller, James H. 2003. Nonnative invasive plants of southern forests: a field guide for identification and control. Gen. Tech. Rep. SRS–62. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southern Research Station. 93 p. https://en.wikipedia.org/wiki/Surfactant?oldformat=true https://en.wikipedia.org/wiki/Penetrant_(biochemical)?oldformat=true
  • 55. Plant references Bush Honeysuckle (Lonicera maackii) https://plants.usda.gov/core/profile?symbol=loma6 https://www.invasiveplantatlas.org/subject.html?sub=3040 https://www.invasive.org/eastern/srs/BH.html Japanese Barberry (Berberis thunbergii) https://plants.usda.gov/core/profile?symbol=beth https://www.invasiveplantatlas.org/subject.html?sub=3010 https://wiki.bugwood.org/Archive:SEEPPC/Berberis_thunbergii Burning Bush (Euonymus alatus) https://plants.usda.gov/core/profile?symbol=EUAL13 http://www.hort.uconn.edu/plants/detail.php?pid=167 https://www.ecolandscaping.org/04/invasive-plants/winged-euonymus-an-exotic-invasive-plant- fact-sheet/
  • 56. Plant references continued Autumn Olive (Elaeagnus umbellata) https://plants.usda.gov/core/profile?symbol=ELUM https://www.invasiveplantatlas.org/subject.html?sub=3021 https://www.invasive.org/eastern/srs/AO.html Multiflora Rose (Rosa multiflora) https://plants.usda.gov/core/profile?symbol=romu https://www.invasiveplantatlas.org/subject.html?sub=3071 https://www.invasive.org/eastern/srs/ER.html Winter Creeper (Euonymus fortunei) https://www.invasiveplantatlas.org/subject.html?sub=3024 https://plants.usda.gov/core/profile?symbol=EUFO5
  • 57. Plant references continued Oriental Bittersweet (Celastrus orbiculatus) https://www.invasiveplantatlas.org/subject.html?sub=3012 https://plants.usda.gov/core/profile?symbol=CEOR7 https://plants.usda.gov/java/charProfile?symbol=CEOR7 Japanese Honeysuckle (Lonicera japonica) https://www.invasiveplantatlas.org/subject.html?sub=3039 https://plants.usda.gov/core/profile?symbol=LOJA https://www.invasive.org/eastern/srs/JH.html Callery Pear = Bradford Pear (Pyrus calleryana) https://plants.usda.gov/core/profile?symbol=PYCA80 https://www.invasiveplantatlas.org/subject.html?sub=10957 https://www.invasive.org/alien/pubs/midatlantic/pyca.htm
  • 58. Plant references continued A Garlic Mustard (Alliaria petiolata formerly A. officinalis) https://www.invasiveplantatlas.org/subject.html?sub=3005 https://www.invasive.org/eastern/srs/GM.html https://plants.usda.gov/core/profile?symbol=alpe4