The Paleozoic Era lasted from 543 to 248 million years ago. It was divided into seven periods: Cambrian, Ordovician, Silurian, Devonian, Mississippian, Pennsylvanian, and Permian. Major events included the diversification of multi-celled animal life in the Cambrian and a mass extinction at the end of the Paleozoic that wiped out 90% of marine species. Life colonized land during this era, with plants, fungi and insects moving ashore.

1. The Paleozoic Era543 to 248 Million Years Ago

2. Paleozoic Era: Stratigraphy

3. The Paleozoic Era The Paleozoic is bracketed by two of the most important events in the history of animal life. At its beginning, multi-celled animals underwent a dramatic "explosion" in diversity, and almost all living animal phyla appeared within a few millions of years. At the other end of the Paleozoic, the largest mass extinction in history wiped out approximately 90% of all marine animal species. The causes of both these events are still not fully understood and the subject of much research and controversy. Roughly halfway in between, animals, fungi, and plants alike colonized the land, the insects took to the air, and the limestone shown in this picture was deposited near Burlington, Missouri.

4. A favorite exercise in introductory paleontology courses is to invent mnemonics for the periods of the Paleozoic. These mnemonics all stand for the seven time periods into which the Paleozoic is divided: Cambrian, Ordovician, Silurian, Devonian, Mississippian, Pennsylvanian, Permian. Note that European geologists lump the Mississippian and Pennsylvanian into a single time period, the Carboniferous. Most of these names derive from locations where rocks of these ages were first studied. Cambria was the Latin name for Wales, and the Ordovices and Silures were two Welsh Celtic tribes.

5. The Devonian is named for Devonshire, England. The Mississippian is named for the upper Mississippi River valley, NOT the state of Mississippi, which has very few rocks of this age. The Pennsylvanian is named for Pennsylvania, surprisingly enough, while the Permian was described from rocks in the region of Perm, a town in the Ural Mountains of Russia. The exception to this rule is the Carboniferous Period; its name means "coal-bearing," and this period is in fact the time when extensive coal beds were formed around the world.

6. The Permian290 to 248 Million Years Ago

7. The Permian The Permian period lasted from 290 to 248 million years ago and was the last period of the Paleozoic Era. The distinction between the Paleozoic and the Mesozoic is made at the end of the Permian in recognition of the largest mass extinction recorded in the history of life on Earth. It affected many groups of organisms in many different environments, but it affected marine communities the most by far, causing the extinction of most of the marine invertebrates of the time.

8. Some groups survived the Permian mass extinction in greatly diminished numbers, but they never again reached the ecological dominance they once had, clearing the way for another group of sea life. On land, a relatively smaller extinction of diapsids and synapsids cleared the way for other forms to dominate, and led to what has been called the "Age of Dinosaurs".

9. Also, the great forests of fern-like plants shifted to gymnosperms, plants with their offspring enclosed within seeds. Modern conifers, the most familiar gymnosperms of today, first appear in the fossil record of the Permian. In all, the Permian was the last of the time for some organisms and a pivotal point for others, and life on earth was never the same again.

10. The global geography of the Permian included massive areas of land and water. By the beginning of the Permian, the motion of the Earth's crustal plates had brought much of the total land together, fused in a supercontinent known as Pangaea. Many of the continents of today in somewhat intact form met in Pangaea (only Asia was broken up at the time), which stretched from the northern to the southern pole.

11. Most of the rest of the surface area of the Earth was occupied by a corresponding single ocean, known as Panthalassa, with a smaller sea to the east of Pangaea known as Tethys. Models indicate that the interior regions of this vast continent were probably dry, with great seasonal fluctuations, because of the lack of the moderating effect of nearby bodies of water, and that only portions received rainfall throughout the year. The ocean itself still has little known about it. There are indications that the climate of the Earth shifted at this time, and that glaciation decreased, as the interiors of continents became drier.

13. The Carboniferous354 to 290 Million Years Ago

14. The Carboniferous The Carboniferous Period occurred from about 354 to 290 million years ago during the late Paleozoic Era. The term "Carboniferous" comes from England, in reference to the rich deposits of coal that occur there. These deposits of coal occur throughout northern Europe, Asia, and midwestern and eastern North America.

15. The term "Carboniferous" is used throughout the world to describe this period, although this period has been separated into the Mississippian (Lower Carboniferous) and the Pennsylvanian (Upper Carboniferous) in the United States. This system was adopted to distinguish the coal-bearing layers of the Pennsylvanian from the mostly limestone Mississippian, and is a result of differing stratigraphy on the different continents.

16. Carboniferous Forest :The Carboniferous Period is famous for its vast coal swamps, such as the one depicted here. Such swamps produced the coal from which the term "Carboniferous", or "carbon-bearing" comes.

17. In addition to having the ideal conditions for the beginnings of coal, several major biological, geological, and climatic events occurred during this time. One of the greatest evolutionary innovations of the Carboniferous was the amniotes egg, which allowed for the further exploitation of the land by certain tetra pods. The amniote egg allowed the ancestors of birds, mammals, and reptiles to reproduce on land by preventing the desiccation of the embryo inside. There was also a trend towards mild temperatures during the Carboniferous, as evidenced by the decrease in lycopods and large insects and an increase in the number of tree ferns.

18. Geologically, the Late Carboniferous collision of Laurussia (present-day Europe and North America) into Gondwanaland (present-day Africa and South America) produced the Appalachian mountain belt of eastern North America and the Hercynian Mountains in the United Kingdom. A further collision of Siberia and eastern Europe created the Ural Mountains.The stratigraphy of the Lower Carboniferous can be easily distinguished from that of the Upper Carboniferous.

19. The environment of the Lower Carboniferous in North America was heavily marine, when seas covered parts of the continents. As a result, most of the mineral found in Lower Carboniferous is limestone, which are composed of the remains of crinoids, lime-encrusted green algae, or calcium carbonate shaped by waves. The North American Upper Carboniferous environment was alternately terrestrial and marine, with the transgression and regression of the seas caused by glaciations.

20. These environmental conditions, with the vast amount of plant material provided by the extensive coal forests, allowed for the production of coal. Plant material did not decay when the seas covered them and pressure and heat eventually built up over the millions of years to transform the plant material to coal.

21. Fossils

24. The Devonian417 to 354 Million Years Ago

25. The DevonianThe RhynieChert in Scotland is a Devonian age deposit containing fossils of both Zosterophyllophytes and Trimerophytes, the two major lines of vascular plants. This indicates that prior to the start of the Devonian, the first major radiations of the plants had already happened. The oldest known vascular plants in the Northern Hemisphere are Devonian. The vegetation of the early Devonian consisted primarily of small plants, the tallest being only a meter tall. By the end of the Devonian, ferns, horsetails and seed plants had also appeared, producing the first trees and the first forests.

26. Archaeopterisone of the first trees.

27. The DevonianAlso during the Devonian, two major animal groups colonized the land. The first tetra pods, or land-living vertebrates, appeared during the Devonian, as did the first terrestrial arthropods, including wingless insects and the earliest arachnids. In the oceans, brachiopods flourished, like the beautifully pyritized brachiopod Paraspiriferbownockeri from Ohio, pictured above and to the right. Crinoids and other echinoderms, tabulate and rugose corals, and ammonites were also common. Many new kinds of fish appeared. During the Devonian, there were three major continental masses: North America and Europe sat together near the equator, much of their current land underneath seas. To the north lay a portion of modern Siberia. A composite continent of South America, Africa, Antarctica, India, and Australia dominated the southern hemisphere.

28. Paraspiriferbownockeri

30. The Silurian443 to 417 Million Years Ago

31. The Silurian The Silurian (443 to 417 million years ago) was a time when the Earth underwent considerable changes that had important repercussions for the environment and life within it. The Silurian witnessed a relative stabilization of the earth's general climate, ending the previous pattern of erratic climatic fluctuations. One result of these changes was the melting of large glacial formations. This contributed to a substantial rise in the levels of the major seas.

32. Coral reefs made their first appearance during this time, and the Silurian was also a remarkable time in the evolution of fishes. Not only does this time period mark the wide and rapid spread of jawless fish, but also the highly significant appearances of both the first known freshwater fish as well as the first fish with jaws. It is also at this time that our first good evidence of life on land is preserved, including relatives of spiders and centipedes, and also the earliest fossils of vascular plants.

35. The Ordovician490 to 443 Million Years Ago

36. The Ordovician The Ordovician period began approximately 490 million years ago, with the end of the Cambrian, and ended around 443 million years ago, with the beginning of the Silurian. At this time, the area north of the tropics was almost entirely ocean, and most of the world's land was collected into the southern super-continent Gondwana. Throughout the Ordovician, Gondwana shifted towards the South Pole and much of it was submerged underwater.

37. The Ordovician is best known for the presence of its diverse marine invertebrates, including graptolites, trilobites, brachiopods, and the conodonts (early vertebrates). A typical marine community consisted of these animals, plus red and green algae, primitive fish, cephalopods, corals, crinoids, and gastropods. More recently, there has been found evidence of tetrahedral spores that are similar to those of primitive land plants, suggesting that plants invaded the land at this time.

38. From the Early to Middle Ordovician, the earth experienced a milder climate in which the weather was warm and the atmosphere contained a lot of moisture. However, when Gondwana finally settled on the South Pole during the Late Ordovician, massive glaciers formed causing shallow seas to drain and sea levels to drop. This likely caused the mass extinctions that characterize the end of the Ordovician, in which 60% of all marine invertebrate genera and 25% of all families went extinct.

41. The Cambrian Period543 to 490 Million Years Ago

42. The Cambrian Period The Cambrian Period marks an important point in the history of life on earth; it is the time when most of the major groups of animals first appear in the fossil record. This event is sometimes called the "Cambrian Explosion", because of the relatively short time over which this diversity of forms appears. It was once thought that the Cambrian rocks contained the first and oldest fossil animals, but these are now to be found in the earlier Vendian strata.